<document id="D14AC31CE99A5B54F9B2A0C25C6B41AF" ID-DOI="10.5852/ejt.2018.395" ID-GBIF-Dataset="10dbb315-5662-4069-a1ae-9692fdf99499" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829724" ID-ZooBank="A244F97F-8CEB-40CE-A182-7E5440337CAA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="juliana" checkinTime="1589571932603" checkinUser="valdenar" docAuthor="Stekolnikov, Alexandr A." docDate="2018" docId="486DBB53FFD5FFCB88F4FB2ADB5DF832" docLanguage="en" docName="ejt-395_stekolnikov.pdf.imf" docOrigin="European Journal of Taxonomy 395" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Schoengastia hyracis Vercammen-Grandjean &amp; Brennan 1957" docType="treatment" docVersion="19" lastPageNumber="80" masterDocId="B454C32BFF9AFF848951FFD6DF63FFD9" masterDocTitle="Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)" masterLastPageNumber="233" masterPageNumber="1" pageNumber="80" updateTime="1740593697770" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="749C7CA0C737CB3B80E9D46D0FB9F97D">Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)</mods:title>
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<mods:namePart id="CD839F9A77BEF1A24C288E1891E75758">Stekolnikov, Alexandr A.</mods:namePart>
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<mods:affiliation id="1B9F71B1D2CAC567A9FC660812D0D2A2">Zoological Institute, Russian Academy of Sciences, Universitetskaya embankment 1, St. Petersburg 199034, Russia.</mods:affiliation>
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<mods:title id="1DD8D546CAE2C874A3FE72032DEE4258">European Journal of Taxonomy</mods:title>
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<mods:date id="B097AE864AD7729B08F5FB8DB1CBF6EA">2018</mods:date>
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<treatment id="486DBB53FFD5FFCB88F4FB2ADB5DF832" ID-DOI="http://doi.org/10.5281/zenodo.3849151" ID-GBIF-Taxon="164160579" ID-Zenodo-Dep="3849151" LSID="urn:lsid:plazi:treatment:486DBB53FFD5FFCB88F4FB2ADB5DF832" httpUri="http://treatment.plazi.org/id/486DBB53FFD5FFCB88F4FB2ADB5DF832" lastPageNumber="80" pageId="79" pageNumber="80" scope_family="Trombiculidae" scope_subFamily="Trombiculinae" scope_superOrder="Acariformes" scope_tribe="Schoengastiini">
<subSubSection id="88DE59CEFFD5FFCB88F4FB2ADBECFACE" box="[421,1167,1276,1303]" pageId="79" pageNumber="80" type="nomenclature">
<paragraph id="C07B0A45FFD5FFCB88F4FB2ADBECFACE" blockId="79.[421,1167,1276,1303]" box="[421,1167,1276,1303]" pageId="79" pageNumber="80">
<heading id="9B33BD29FFD5FFCB88F4FB2ADBECFACE" box="[421,1167,1276,1303]" centered="true" fontSize="11" level="2" pageId="79" pageNumber="80" reason="2">
<taxonomicName id="07C471C6FFD5FFCB88F4FB2ADBECFACE" ID-CoL="8ZK7X" authority="Vercammen-Grandjean &amp; Brennan, 1957" authorityName="Vercammen-Grandjean &amp; Brennan" authorityYear="1957" box="[421,1167,1276,1303]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="hyracis">
<emphasis id="F2B0D657FFD5FFCB88F4FB2ADDC6FACE" bold="true" box="[421,677,1276,1303]" italics="true" pageId="79" pageNumber="80">Schoengastia hyracis</emphasis>
<bibRefCitation id="A45577B4FFD5FFCB8BFCFB2ADBECFACE" author="Vercammen-Grandjean &amp; Brennan" box="[685,1167,1276,1303]" firstAuthor="Vercammen-Grandjean" pageId="79" pageNumber="80" pagination="484 - 496" refId="ref68553" refString="Vercammen-Grandjean P. H. &amp; Brennan J. M. 1957. Eight new chiggers from East Africa and a new genus, Trombigastia (Acarina: Trombiculidae). Annals of the Entomological Society of America 50 (5): 484 - 496. https: // doi. org / 10.1093 / aesa / 50.5.484" type="journal article" year="1957">Vercammen-Grandjean &amp; Brennan, 1957</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFD5FFCB89ECFA97DB33FA35" pageId="79" pageNumber="80" type="reference_group">
<paragraph id="C07B0A45FFD5FFCB89ECFA97DB4CFA82" blockId="79.[189,1071,1345,1372]" box="[189,1071,1345,1372]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="E0D42D6BFFD5FFCB89ECFA97DB4CFA82" box="[189,1071,1345,1372]" pageId="79" pageNumber="80">
<treatmentCitation id="41652C54FFD5FFCB89ECFA97DCBCFA82" author="Vercammen-Grandjean P. H. &amp; Brennan J. M." box="[189,991,1345,1372]" page="486" pageId="79" pageNumber="80" year="1957">
<taxonomicName id="07C471C6FFD5FFCB89ECFA97DCBCFA82" ID-CoL="8ZK7X" authority="Vercammen-Grandjean &amp; Brennan, 1957: 486" authorityName="Vercammen-Grandjean &amp; Brennan" authorityPageNumber="486" authorityYear="1957" box="[189,991,1345,1372]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="hyracis">
<emphasis id="F2B0D657FFD5FFCB89ECFA97DED4FA82" box="[189,439,1345,1371]" italics="true" pageId="79" pageNumber="80">Schoengastia hyracis</emphasis>
<bibRefCitation id="A45577B4FFD5FFCB88EFFA97DCBCFA82" author="Vercammen-Grandjean &amp; Brennan" box="[446,991,1345,1372]" firstAuthor="Vercammen-Grandjean" page="486" pageId="79" pageNumber="80" pagination="484 - 496" refId="ref68553" refString="Vercammen-Grandjean P. H. &amp; Brennan J. M. 1957. Eight new chiggers from East Africa and a new genus, Trombigastia (Acarina: Trombiculidae). Annals of the Entomological Society of America 50 (5): 484 - 496. https: // doi. org / 10.1093 / aesa / 50.5.484" type="journal article" year="1957">Vercammen-Grandjean &amp; Brennan, 1957: 486</bibRefCitation>
</taxonomicName>
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, fig. 3.
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</paragraph>
<paragraph id="C07B0A45FFD5FFCB89ECFA53DB7DFA46" blockId="79.[189,1104,1413,1517]" box="[189,1054,1413,1440]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="E0D42D6BFFD5FFCB89ECFA53DB7DFA46" box="[189,1054,1413,1440]" pageId="79" pageNumber="80">
<taxonomicName id="07C471C6FFD5FFCB89ECFA53DD0DFA46" ID-CoL="8ZK7X" authorityName="Vercammen-Grandjean &amp; Brennan" authorityYear="1957" box="[189,622,1413,1439]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="hyracis" subGenus="Schoengastia">
<emphasis id="F2B0D657FFD5FFCB89ECFA53DD0DFA46" box="[189,622,1413,1439]" italics="true" pageId="79" pageNumber="80">Schoengastia (Schoengastia) hyracis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="41652C54FFD5FFCB8BDAFA53DB79FA46" author="Vercammen-Grandjean P. H." box="[651,1050,1413,1440]" page="81" pageId="79" pageNumber="80" year="1965">
<bibRefCitation id="A45577B4FFD5FFCB8BDAFA53DB79FA46" author="Vercammen-Grandjean" box="[651,1050,1413,1440]" firstAuthor="Vercammen-Grandjean" page="81" pageId="79" pageNumber="80" refId="ref68052" refString="Vercammen-Grandjean P. H. 1965 c. Trombiculinae of the World. Synopsis with Generic, Subgeneric, and Group Diagnoses (Acarina, Trombiculidae). George Williams Hooper Foundation, San Francisco." type="book" year="1965" yearSuffix="c">Vercammen-Grandjean 1965c: 81</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FFD5FFCB89ECFA7ADC87FA1F" blockId="79.[189,1104,1413,1517]" box="[189,996,1451,1478]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="E0D42D6BFFD5FFCB89ECFA7ADC87FA1F" box="[189,996,1451,1478]" pageId="79" pageNumber="80">
<taxonomicName id="07C471C6FFD5FFCB89ECFA7ADD40FA1F" ID-CoL="8ZK7X" box="[189,547,1452,1478]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="79" pageNumber="80" phylum="Arthropoda" rank="subSpecies" species="haddowi" subSpecies="hyracis">
<emphasis id="F2B0D657FFD5FFCB89ECFA7ADD40FA1F" box="[189,547,1452,1478]" italics="true" pageId="79" pageNumber="80">Schoengastia haddowi hyracis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="41652C54FFD5FFCB8B11FA7DDC83FA1F" author="Vercammen-Grandjean P. H." box="[576,992,1451,1478]" page="652" pageId="79" pageNumber="80" year="1958">
<bibRefCitation id="A45577B4FFD5FFCB8B11FA7DDC83FA1F" author="Vercammen-Grandjean" box="[576,992,1451,1478]" firstAuthor="Vercammen-Grandjean" page="652" pageId="79" pageNumber="80" pagination="619 - 670" refId="ref67603" refString="Vercammen-Grandjean P. H. 1958 b. Revue des Schoengastia centrafricains et remaniements divers de cet important genre (Acarina - Trombiculidae). Annales de parasitologie humaine et comparee 33 (5 - 6): 619 - 670." type="journal article" year="1958" yearSuffix="b">Vercammen-Grandjean 1958b: 652</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FFD5FFCB89ECFA04DB33FA35" blockId="79.[189,1104,1413,1517]" box="[189,1104,1490,1517]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="E0D42D6BFFD5FFCB89ECFA04DB33FA35" box="[189,1104,1490,1517]" pageId="79" pageNumber="80">
<taxonomicName id="07C471C6FFD5FFCB89ECFA04DDB9FA35" ID-CoL="8ZK7X" box="[189,730,1490,1516]" class="Arachnida" family="Trombiculidae" genus="Schoengastia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="79" pageNumber="80" phylum="Arthropoda" rank="subSpecies" species="haddowi" subGenus="Schoengastia" subSpecies="hyracis">
<emphasis id="F2B0D657FFD5FFCB89ECFA04DDB9FA35" box="[189,730,1490,1516]" italics="true" pageId="79" pageNumber="80">Schoengastia (Schoengastia) haddowi hyracis</emphasis>
</taxonomicName>
– 
<treatmentCitation id="41652C54FFD5FFCB8BA9FA04DCA5FA34" author="Zumpt F." box="[760,966,1490,1517]" page="155" pageId="79" pageNumber="80" year="1961">
<bibRefCitation id="A45577B4FFD5FFCB8BA9FA04DCA5FA34" author="Zumpt" box="[760,966,1490,1517]" firstAuthor="Zumpt" page="155" pageId="79" pageNumber="80" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">Zumpt 1961: 155</bibRefCitation>
</treatmentCitation>
, fig. 92a–d.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFD5FFCB89ECF9C1DEF9F926" pageId="79" pageNumber="80" type="materials_examined">
<materialsCitation id="70AC0018FFD5FFCB89ECF9C1DEC5F9BA" box="[189,422,1559,1635]" collectionCode="RMCA" pageId="79" pageNumber="80" specimenCount="1" typeStatus="holotype">
<paragraph id="C07B0A45FFD5FFCB89ECF9C1DE4CF9E8" blockId="79.[189,303,1559,1585]" box="[189,303,1559,1585]" pageId="79" pageNumber="80">
<heading id="9B33BD29FFD5FFCB89ECF9C1DE4CF9E8" bold="true" box="[189,303,1559,1585]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="F2B0D657FFD5FFCB89ECF9C1DE4CF9E8" bold="true" box="[189,303,1559,1585]" pageId="79" pageNumber="80">
<typeStatus id="1F7FB4E7FFD5FFCB89ECF9C1DE4CF9E8" box="[189,303,1559,1585]" pageId="79" pageNumber="80" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFD5FFCB89ECF99EDEC5F9BA" blockId="79.[189,422,1608,1635]" box="[189,422,1608,1635]" pageId="79" pageNumber="80">
<collectionCode id="A6D59280FFD5FFCB89ECF99EDE71F9BB" box="[189,274,1608,1634]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="79" pageNumber="80" type="Museum">RMCA</collectionCode>
(not found).
</paragraph>
</materialsCitation>
<paragraph id="C07B0A45FFD5FFCB89ECF95ADEE9F97F" blockId="79.[189,394,1676,1702]" box="[189,394,1676,1702]" pageId="79" pageNumber="80">
<heading id="9B33BD29FFD5FFCB89ECF95ADEE9F97F" bold="true" box="[189,394,1676,1702]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="F2B0D657FFD5FFCB89ECF95ADEE9F97F" bold="true" box="[189,394,1676,1702]" pageId="79" pageNumber="80">Material revised</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFD5FFCB89ECF968DEF9F926" blockId="79.[189,1399,1726,1791]" pageId="79" pageNumber="80">
<materialsCitation id="70AC0018FFD5FFCB89ECF968DE94F900" box="[189,503,1726,1753]" pageId="79" pageNumber="80" specimenCode="113906" specimenCount="1" typeStatus="paratype">
<specimenCount id="D6C2C1CCFFD5FFCB89ECF968DF8DF901" box="[189,238,1726,1752]" pageId="79" pageNumber="80" type="generic">One</specimenCount>
<typeStatus id="1F7FB4E7FFD5FFCB89A2F969DE3BF900" box="[243,344,1727,1753]" pageId="79" pageNumber="80">paratype</typeStatus>
(No. 
<specimenCode id="9062A23EFFD5FFCB88C7F968DE8CF900" box="[406,495,1726,1753]" pageId="79" pageNumber="80">113906</specimenCode>
)
</materialsCitation>
<materialsCitation id="70AC0018FFD5FFCB88ADF968DCC9F901" box="[508,938,1726,1753]" collectionCode="RMCA" pageId="79" pageNumber="80" specimenCount="21">
and 
<specimenCount id="D6C2C1CCFFD5FFCB8B7AF968DD2BF901" box="[555,584,1726,1752]" pageId="79" pageNumber="80">21</specimenCount>
more specimens from 
<collectionCode id="A6D59280FFD5FFCB8A1DF968DCC5F901" box="[844,934,1726,1752]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="79" pageNumber="80" type="Museum">RMCA</collectionCode>
;
</materialsCitation>
<materialsCitation id="70AC0018FFD5FFCB8AFEF968DEECF926" collectionCode="BMNH" pageId="79" pageNumber="80" specimenCode="1956.9.15.21, 1956.9.15.22" specimenCount="2" typeStatus="paratype">
<specimenCount id="D6C2C1CCFFD5FFCB8AFEF968DCDDF901" box="[943,958,1726,1752]" pageId="79" pageNumber="80" type="generic">2</specimenCount>
<typeStatus id="1F7FB4E7FFD5FFCB8A92F969DB50F900" box="[963,1075,1727,1753]" pageId="79" pageNumber="80">paratypes</typeStatus>
from 
<collectionCode id="A6D59280FFD5FFCB8D27F969DBADF900" box="[1142,1230,1727,1753]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="79" pageNumber="80">BMNH</collectionCode>
(
<specimenCode id="9062A23EFFD5FFCB8D8AF968DA14F900" box="[1243,1399,1726,1753]" pageId="79" pageNumber="80">1956.9.15.21</specimenCode>
and 
<specimenCode id="9062A23EFFD5FFCB89BEF932DEECF926" box="[239,399,1764,1791]" pageId="79" pageNumber="80">1956.9.15.22</specimenCode>
</materialsCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFD5FFCB89ECF8FFDD79F8AD" box="[189,538,1833,1909]" pageId="79" pageNumber="80" type="distribution">
<paragraph id="C07B0A45FFD5FFCB89ECF8FFDE35F89A" blockId="79.[189,342,1833,1859]" box="[189,342,1833,1859]" pageId="79" pageNumber="80">
<heading id="9B33BD29FFD5FFCB89ECF8FFDE35F89A" bold="true" box="[189,342,1833,1859]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="F2B0D657FFD5FFCB89ECF8FFDE35F89A" bold="true" box="[189,342,1833,1859]" pageId="79" pageNumber="80">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFD5FFCB89ECF88CDD79F8AD" blockId="79.[189,538,1882,1909]" box="[189,538,1882,1909]" pageId="79" pageNumber="80">
<collectingCountry id="B8D34AD5FFD5FFCB89ECF88CDE7BF8AD" box="[189,280,1882,1908]" name="Uganda" pageId="79" pageNumber="80">Uganda</collectingCountry>
(
<collectingRegion id="0200C4A7FFD5FFCB8879F88CDEFBF8AC" box="[296,408,1882,1909]" country="Uganda" name="Kaabong" pageId="79" pageNumber="80">Kaabong</collectingRegion>
), 
<collectingCountry id="B8D34AD5FFD5FFCB88FDF88CDD74F8AD" box="[428,535,1882,1908]" name="Tanzania" pageId="79" pageNumber="80">Zanzibar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFFD5FFCB89ECF849DB5DF832" box="[189,1086,1951,2027]" pageId="79" pageNumber="80" type="biology_ecology">
<paragraph id="C07B0A45FFD5FFCB89ECF849DE60F860" blockId="79.[189,259,1951,1977]" box="[189,259,1951,1977]" pageId="79" pageNumber="80">
<heading id="9B33BD29FFD5FFCB89ECF849DE60F860" bold="true" box="[189,259,1951,1977]" fontSize="11" level="3" pageId="79" pageNumber="80" reason="3">
<emphasis id="F2B0D657FFD5FFCB89ECF849DE60F860" bold="true" box="[189,259,1951,1977]" pageId="79" pageNumber="80">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FFD5FFCB89ECF807DB5DF832" blockId="79.[189,1086,2001,2027]" box="[189,1086,2001,2027]" pageId="79" pageNumber="80">
<taxonomicName id="07C471C6FFD5FFCB89ECF807DEC5F832" authorityName="Wolf" authorityYear="1822" box="[189,422,2001,2027]" class="Mammalia" family="Cercopithecidae" genus="Cercopithecus" kingdom="Animalia" order="Primates" pageId="79" pageNumber="80" phylum="Chordata" rank="species" species="mitis">
<emphasis id="F2B0D657FFD5FFCB89ECF807DEC5F832" box="[189,422,2001,2027]" italics="true" pageId="79" pageNumber="80">Cercopithecus mitis</emphasis>
</taxonomicName>
, 
<taxonomicName id="07C471C6FFD5FFCB88E5F807DC20F832" authorityName="Thomas" authorityYear="1897" box="[436,835,2001,2027]" class="Mammalia" family="Macroscelididae" genus="Petrodromus" kingdom="Animalia" order="Macroscelidea" pageId="79" pageNumber="80" phylum="Chordata" rank="subSpecies" species="tetradactylus" subSpecies="sultani">
<emphasis id="F2B0D657FFD5FFCB88E5F807DC20F832" box="[436,835,2001,2027]" italics="true" pageId="79" pageNumber="80">Petrodromus tetradactylus sultani</emphasis>
</taxonomicName>
, 
<taxonomicName id="07C471C6FFD5FFCB8A03F807DCBFF832" authorityName="Thomas" authorityYear="1892" box="[850,988,2001,2027]" class="Mammalia" family="Procaviidae" kingdom="Animalia" order="Hyracoidea" pageId="79" pageNumber="80" phylum="Chordata" rank="family">Procaviidae</taxonomicName>
gen. sp.
</paragraph>
</subSubSection>
</treatment>
</document>