<document id="022DEC9DE95052578EBBE06A38390CBB" ID-DOI="10.5852/ejt.2018.395" ID-GBIF-Dataset="10dbb315-5662-4069-a1ae-9692fdf99499" ID-ISSN="2118-9773" ID-Zenodo-Dep="3829724" ID-ZooBank="A244F97F-8CEB-40CE-A182-7E5440337CAA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1589571932603" checkinUser="valdenar" docAuthor="Stekolnikov, Alexandr A." docDate="2018" docId="486DBB53FF84FF9A889BFEDDDD5FFC2B" docLanguage="en" docName="ejt-395_stekolnikov.pdf.imf" docOrigin="European Journal of Taxonomy 395" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Schoengastiella nasicola" docType="treatment" docVersion="17" lastPageNumber="31" masterDocId="B454C32BFF9AFF848951FFD6DF63FFD9" masterDocTitle="Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)" masterLastPageNumber="233" masterPageNumber="1" pageNumber="31" updateTime="1739281695645" updateUser="valdenar" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="565F1603EA63FB83078C783D0D30A50D">Taxonomy and distribution of African chiggers (Acariformes, Trombiculidae)</mods:title>
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<mods:namePart id="1241B438ECCCDA21F1F125ABBCE12ED3">Stekolnikov, Alexandr A.</mods:namePart>
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<mods:affiliation id="68A504EC2D63FB7DCBC88EB6B5D35A80">Zoological Institute, Russian Academy of Sciences, Universitetskaya embankment 1, St. Petersburg 199034, Russia.</mods:affiliation>
<mods:nameIdentifier id="AA35029BF6CDD28C75B3CD19F35DD291" type="email">Alexandr.Stekolnikov@zin.ru</mods:nameIdentifier>
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<mods:title id="19FEC1948D8B9CDDEDE6560064C1169B">European Journal of Taxonomy</mods:title>
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<mods:date id="B7547C71F5EA7CA94AF7963C3D5CBAD9">2018</mods:date>
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<treatment id="486DBB53FF84FF9A889BFEDDDD5FFC2B" ID-DOI="http://doi.org/10.5281/zenodo.3849542" ID-GBIF-Taxon="164160490" ID-Zenodo-Dep="3849542" LSID="urn:lsid:plazi:treatment:486DBB53FF84FF9A889BFEDDDD5FFC2B" httpUri="http://treatment.plazi.org/id/486DBB53FF84FF9A889BFEDDDD5FFC2B" lastPageNumber="31" pageId="30" pageNumber="31" scope_family="Trombiculidae" scope_subFamily="Gahrliepiinae" scope_superOrder="Acariformes">
<subSubSection id="88DE59CEFF84FF9A889BFEDDDB0AFEFF" box="[458,1129,267,294]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph id="C07B0A45FF84FF9A889BFEDDDB0AFEFF" blockId="30.[458,1129,267,294]" box="[458,1129,267,294]" pageId="30" pageNumber="31">
<heading id="9B33BD29FF84FF9A889BFEDDDB0AFEFF" box="[458,1129,267,294]" centered="true" fontSize="11" level="2" pageId="30" pageNumber="31" reason="2">
<taxonomicName id="07C471C6FF84FF9A889BFEDDDB0AFEFF" ID-CoL="8ZK3F" authority="(Vercammen-Grandjean, 1956)" baseAuthorityName="Vercammen-Grandjean" baseAuthorityYear="1956" box="[458,1129,267,294]" class="Arachnida" family="Trombiculidae" genus="Schoengastiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nasicola">
<emphasis id="F2B0D657FF84FF9A889BFEDDDD96FEFF" bold="true" box="[458,757,267,294]" italics="true" pageId="30" pageNumber="31">Schoengastiella nasicola</emphasis>
(
<bibRefCitation id="A45577B4FF84FF9A8A55FEDDDB02FEFF" author="Vercammen-Grandjean" box="[772,1121,267,294]" firstAuthor="Vercammen-Grandjean" pageId="30" pageNumber="31" pagination="92" refId="ref67279" refString="Vercammen-Grandjean P. H. 1956 a. Addendum [figure of Audya nasicola]. Revue de zoologie et de botanique africaines 54: 92." type="journal article" year="1956">Vercammen-Grandjean, 1956</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF84FF9A89ECFE99DB34FDC7" pageId="30" pageNumber="31" type="reference_group">
<paragraph id="C07B0A45FF84FF9A89ECFE99DC46FEB3" blockId="30.[189,805,335,362]" box="[189,805,335,362]" pageId="30" pageNumber="31">
<taxonomicName id="07C471C6FF84FF9A89ECFE99DC42FEB3" authority="Vercammen-Grandjean, 1956 b: 350" authorityName="Vercammen-Grandjean" authorityPageNumber="350" authorityYear="1956" box="[189,801,335,362]" class="Arachnida" family="Trombiculidae" genus="Audya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nasicola">
<emphasis id="F2B0D657FF84FF9A89ECFE99DE11FEB0" box="[189,370,335,361]" italics="true" pageId="30" pageNumber="31">Audya nasicola</emphasis>
<bibRefCitation id="A45577B4FF84FF9A8828FE99DC42FEB3" author="Vercammen-Grandjean" box="[377,801,335,362]" firstAuthor="Vercammen-Grandjean" page="350" pageId="30" pageNumber="31" pagination="349 - 352" refId="ref67306" refString="Vercammen-Grandjean P. H. 1956 b. Audya nasicola: un Trombiculidae d'un genre nouveau dans la sous-famille des Gahrliepiinae. Revue de zoologie et de botanique africaines 53: 349 - 352." type="journal article" year="1956" yearSuffix="b">Vercammen-Grandjean, 1956b: 350</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFE45DCFAFE74" blockId="30.[189,1111,403,543]" box="[189,921,403,430]" pageId="30" pageNumber="31">
<taxonomicName id="07C471C6FF84FF9A89ECFE45DE11FE74" authorityName="Vercammen-Grandjean" authorityYear="1956" box="[189,370,403,429]" class="Arachnida" family="Trombiculidae" genus="Audya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nasicola">
<emphasis id="F2B0D657FF84FF9A89ECFE45DE11FE74" box="[189,370,403,429]" italics="true" pageId="30" pageNumber="31">Audya nasicola</emphasis>
</taxonomicName>
– 
<bibRefCitation id="A45577B4FF84FF9A88DEFE45DC7DFE74" author="Vercammen-Grandjean" box="[399,798,403,430]" firstAuthor="Vercammen-Grandjean" page="92" pageId="30" pageNumber="31" pagination="92" refId="ref67279" refString="Vercammen-Grandjean P. H. 1956 a. Addendum [figure of Audya nasicola]. Revue de zoologie et de botanique africaines 54: 92." type="journal article" year="1956" yearSuffix="a">
Vercammen-Grandjean 
<date id="B47A2C85FF84FF9A8BF6FE45DD81FE77" box="[679,738,403,430]" pageId="30" pageNumber="31" value="1956">1956</date>
a: 92
</bibRefCitation>
, figs A–B.
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFE6FDCE3FE0A" blockId="30.[189,1111,403,543]" box="[189,896,441,467]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="E0D42D6BFF84FF9A89ECFE6FDCE3FE0A" box="[189,896,441,467]" pageId="30" pageNumber="31">
<taxonomicName id="07C471C6FF84FF9A89ECFE6FDD6DFE0A" box="[189,526,441,467]" class="Arachnida" family="Trombiculidae" genus="Gahrliepia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nasicola" subGenus="Audya">
<emphasis id="F2B0D657FF84FF9A89ECFE6FDD6DFE0A" box="[189,526,441,467]" italics="true" pageId="30" pageNumber="31">Gahrliepia (Audya) nasicola</emphasis>
</taxonomicName>
– 
<treatmentCitation id="41652C54FF84FF9A8B7DFE6FDD99FE0A" author="Zumpt F." box="[556,762,441,467]" page="174" pageId="30" pageNumber="31" year="1961">
<bibRefCitation id="A45577B4FF84FF9A8B7DFE6FDD99FE0A" author="Zumpt" box="[556,762,441,467]" firstAuthor="Zumpt" page="174" pageId="30" pageNumber="31" pagination="1 - 457" refId="ref69765" refString="Zumpt F. (ed.) 1961. The arthropod parasites of vertebrates in Africa south of the Sahara (Ethiopian Region). Vol. 1: Chelicerata. Publications of the South African Institute for Medical Research 11 (50): 1 - 457." type="journal article" year="1961">
Zumpt 
<date id="B47A2C85FF84FF9A8BD0FE6FDDDFFE0A" box="[641,700,441,467]" pageId="30" pageNumber="31" value="1961">1961</date>
: 174
</bibRefCitation>
</treatmentCitation>
, fig. 97f–g.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFE08DD1FFE21" blockId="30.[189,1111,403,543]" box="[189,636,478,505]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="E0D42D6BFF84FF9A89ECFE08DD1FFE21" box="[189,636,478,505]" pageId="30" pageNumber="31">
<taxonomicName id="07C471C6FF84FF9A89ECFE08DE11FE21" authorityName="Vercammen-Grandjean" authorityYear="1956" box="[189,370,478,504]" class="Arachnida" family="Trombiculidae" genus="Audya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nasicola">
<emphasis id="F2B0D657FF84FF9A89ECFE08DE11FE21" box="[189,370,478,504]" italics="true" pageId="30" pageNumber="31">Audya nasicola</emphasis>
</taxonomicName>
– 
<treatmentCitation id="41652C54FF84FF9A88DEFE08DD1BFE21" author="Taufflieb R." box="[399,632,478,505]" page="30" pageId="30" pageNumber="31" year="1965">
<bibRefCitation id="A45577B4FF84FF9A88DEFE08DD1BFE21" author="Taufflieb" box="[399,632,478,505]" firstAuthor="Taufflieb" page="30" pageId="30" pageNumber="31" pagination="21 - 36" refId="ref66511" refString="Taufflieb R. 1965 a. Contribution a l'etude des Trombiculidae (Acarina) d'Angola. Publicacoes culturais da Companhia de Diamantes de Angola 4095: 21 - 36." type="journal article" year="1965" yearSuffix="a">
Taufflieb 
<date id="B47A2C85FF84FF9A8B51FE08DD58FE20" box="[512,571,478,505]" pageId="30" pageNumber="31" value="1965">1965</date>
a: 30
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFDD2DB34FDC7" blockId="30.[189,1111,403,543]" box="[189,1111,516,543]" pageId="30" pageNumber="31">
<treatmentCitationGroup id="E0D42D6BFF84FF9A89ECFDD2DB34FDC7" box="[189,1111,516,543]" pageId="30" pageNumber="31">
<taxonomicName id="07C471C6FF84FF9A89ECFDD2DD24FDC7" ID-CoL="8ZK3F" baseAuthorityName="Vercammen-Grandjean" baseAuthorityYear="1956" box="[189,583,516,542]" class="Arachnida" family="Trombiculidae" genus="Schoengastiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="nasicola" subGenus="Audya">
<emphasis id="F2B0D657FF84FF9A89ECFDD2DD24FDC7" box="[189,583,516,542]" italics="true" pageId="30" pageNumber="31">Schoengastiella (Audya) nasicola</emphasis>
</taxonomicName>
– 
<treatmentCitation id="41652C54FF84FF9A8B35FDD2DC96FDC7" author="Vercammen-Grandjean P. H." box="[612,1013,516,543]" page="402" pageId="30" pageNumber="31" year="1975">
<bibRefCitation id="A45577B4FF84FF9A8B35FDD2DC96FDC7" author="Vercammen-Grandjean" box="[612,1013,516,543]" firstAuthor="Vercammen-Grandjean" page="402" pageId="30" pageNumber="31" pagination="397 - 439" refId="ref68361" refString="Vercammen-Grandjean P. H. 1975. Some larval Trombiculidae of the Ethiopian Region (Acari). Revue de zoologie africaine 89 (2): 397 - 439." type="journal article" year="1975">
Vercammen-Grandjean 
<date id="B47A2C85FF84FF9A8A2AFDD2DCD5FDC6" box="[891,950,516,543]" pageId="30" pageNumber="31" value="1975">1975</date>
: 402
</bibRefCitation>
</treatmentCitation>
, fig. B8.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF84FF9A89ECFD9EDDD1FCD1" pageId="30" pageNumber="31" type="materials_examined">
<paragraph id="C07B0A45FF84FF9A89ECFD9EDE4CFDBB" blockId="30.[189,303,584,610]" box="[189,303,584,610]" pageId="30" pageNumber="31">
<heading id="9B33BD29FF84FF9A89ECFD9EDE4CFDBB" bold="true" box="[189,303,584,610]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="F2B0D657FF84FF9A89ECFD9EDE4CFDBB" bold="true" box="[189,303,584,610]" pageId="30" pageNumber="31">
<typeStatus id="1F7FB4E7FF84FF9A89ECFD9EDE4CFDBB" box="[189,303,584,610]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFDAFDE09FD4A" blockId="30.[189,362,633,659]" box="[189,362,633,659]" pageId="30" pageNumber="31">
<materialsCitation id="70AC0018FF84FF9A89ECFDAFDE09FD4A" ID-GBIF-Occurrence="2620304500" box="[189,362,633,659]" collectionCode="RMCA" pageId="30" pageNumber="31" specimenCode="RMCA 82444" specimenCount="1" typeStatus="holotype">
<specimenCode id="9062A23EFF84FF9A89ECFDAFDE05FD4A" box="[189,358,633,659]" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="30" pageNumber="31" type="Museum">RMCA 82444</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFD6ADEE9FD0F" blockId="30.[189,394,700,726]" box="[189,394,700,726]" pageId="30" pageNumber="31">
<heading id="9B33BD29FF84FF9A89ECFD6ADEE9FD0F" bold="true" box="[189,394,700,726]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="F2B0D657FF84FF9A89ECFD6ADEE9FD0F" bold="true" box="[189,394,700,726]" pageId="30" pageNumber="31">Material revised</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFD3BDDD1FCD1" blockId="30.[189,690,749,776]" box="[189,690,749,776]" pageId="30" pageNumber="31">
<materialsCitation id="70AC0018FF84FF9A89ECFD3BDE49FCDE" ID-GBIF-Occurrence="2620304354" box="[189,298,749,775]" collectionCode="RMCA" pageId="30" pageNumber="31" specimenCount="1" typeStatus="holotype">
<typeStatus id="1F7FB4E7FF84FF9A89ECFD3BDE49FCDE" box="[189,298,749,775]" pageId="30" pageNumber="31" type="holotype">Holotype</typeStatus>
</materialsCitation>
<materialsCitation id="70AC0018FF84FF9A8860FD3BDDD1FCD1" box="[305,690,749,776]" collectionCode="RMCA" pageId="30" pageNumber="31" specimenCount="4" typeStatus="paratype">
and four 
<typeStatus id="1F7FB4E7FF84FF9A88CAFD38DD68FCD1" box="[411,523,750,776]" pageId="30" pageNumber="31" type="paratype">paratypes</typeStatus>
from 
<collectionCode id="A6D59280FF84FF9A8B02FD38DDCEFCD1" box="[595,685,750,776]" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="30" pageNumber="31" type="Museum">RMCA</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF84FF9A89ECFCE7DDCBFCA4" box="[189,680,817,893]" pageId="30" pageNumber="31" type="distribution">
<paragraph id="C07B0A45FF84FF9A89ECFCE7DE35FC92" blockId="30.[189,342,817,843]" box="[189,342,817,843]" pageId="30" pageNumber="31">
<heading id="9B33BD29FF84FF9A89ECFCE7DE35FC92" bold="true" box="[189,342,817,843]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="F2B0D657FF84FF9A89ECFCE7DE35FC92" bold="true" box="[189,342,817,843]" pageId="30" pageNumber="31">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFCB5DDCBFCA4" blockId="30.[189,680,866,893]" box="[189,680,866,893]" pageId="30" pageNumber="31">
<collectingCountry id="B8D34AD5FF84FF9A89ECFCB5DE58FCA5" box="[189,315,866,893]" name="Democratic Republic of the Congo" pageId="30" pageNumber="31">DR Congo</collectingCountry>
(Mbandaka), 
<collectingCountry id="B8D34AD5FF84FF9A888EFCB4DD56FCA5" box="[479,565,866,892]" name="Angola" pageId="30" pageNumber="31">Angola</collectingCountry>
(Dundo).
</paragraph>
</subSubSection>
<subSubSection id="88DE59CEFF84FF9A89ECFC70DD5FFC2B" box="[189,572,934,1010]" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="C07B0A45FF84FF9A89ECFC70DE60FC19" blockId="30.[189,259,934,960]" box="[189,259,934,960]" pageId="30" pageNumber="31">
<heading id="9B33BD29FF84FF9A89ECFC70DE60FC19" bold="true" box="[189,259,934,960]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="F2B0D657FF84FF9A89ECFC70DE60FC19" bold="true" box="[189,259,934,960]" pageId="30" pageNumber="31">Hosts</emphasis>
</heading>
</paragraph>
<paragraph id="C07B0A45FF84FF9A89ECFC01DD5FFC2B" blockId="30.[189,572,983,1010]" box="[189,572,983,1010]" pageId="30" pageNumber="31">
<taxonomicName id="07C471C6FF84FF9A89ECFC01DEF3FC28" baseAuthorityName="Du Chaillu" baseAuthorityYear="1860" box="[189,400,983,1009]" class="Mammalia" family="Tenrecidae" genus="Potamogale" kingdom="Animalia" order="Afrosoricida" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="velox">
<emphasis id="F2B0D657FF84FF9A89ECFC01DEF3FC28" box="[189,400,983,1009]" italics="true" pageId="30" pageNumber="31">Potamogale velox</emphasis>
</taxonomicName>
, 
<emphasis id="F2B0D657FF84FF9A88CEFC0EDD56FC28" box="[415,565,984,1009]" italics="true" pageId="30" pageNumber="31">
<taxonomicName id="07C471C6FF84FF9A88CEFC0EDE89FC28" authorityName="Fischer" authorityYear="1803" box="[415,490,984,1009]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">Rattus</taxonomicName>
<taxonomicName id="07C471C6FF84FF9A88A0FC0EDD56FC28" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[497,565,984,1009]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="rattus">rattus</taxonomicName>
</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
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