Two new genera and two new species of troglobitic harvestmen of Stygnopsidae (Opiliones, Laniatores, Gonyleptoidea) from Oaxaca, Mexico, with notes on selected morphological characters Author Cruz-López, Jesús A. Colección Nacional de Arácnidos, Departamento de Zoología, Instituto de Biología, Universidad Nacional Autónoma de México, Apartado postal 70 - 153, 04510, Ciudad de México (Mexico) and Instituto Nacional de Investigaciones Forestales, Agrícolas y Pecuarias (INIFAP), Centro Regional de Investigación Pacífico Sur (CIRPAS), Campo Experimental Valles Centrales, Melchor Ocampo No. 7, Santo Domingo Barrio Bajo, Villa de Etla, 68200 Oaxaca (Mexico) thelyphonidito @ gmail. com thelyphonidito@gmail.com text Zoosystema 2021 2021-02-23 43 5 101 112 journal article 7961 10.5252/zoosystema2021v43a5 1d93b583-d19b-4889-a904-7cb366aca615 1638-9387 4569986 urn:lsid:zoobank.org:pub:B7E88CF5-60E5-4B67-9EB9-C4D8FAC2996A Toojah cimutaa n. gen., n. sp. ( Figs 5-8 ) urn:lsid:zoobank.org:act: 4F3A2A0B-3D59-43D4-A89F-C74D8C7983CC TYPE MATERIAL. — Holotype . Mexico • ♂ ; Oaxaca , Valle Nacional , Te Cimutaa cave ; 17°54’10.5”N , 96°22’37.5”W ; 25.IV. 2008 ; Paul Bryant leg.; CNAN-T01344 . DIAGNOSIS. — As for the genus. ETYMOLOGY. — Noun in apposition referring to the name of the cave where it was found. DESCRIPTION Measurement Scutum length = 3.4 mm ; scutum width at mid-bulge level = 1.7 mm , scutum width at posterior margin = 2.8 mm ; cheliceral hand length = 3.0 mm. Body ( Figs 5 ; 6 ) Scutum type zeta (ζ) with the mid-bulge not marked, lateral margins straight, posterior margin of scutum wider than midbulge section giving the appearance of trapeze; ocularium at the frontal margin, wide and rounded, no eyes. Lateral clear areas teardrop-shaped, at level of mesotergal area II. Sulcus I very deep, sulci II-IV shallow. Coxae I and II with similar size, coxae III and IV similar in size and slightly larger than coxae I and II. Coxae I and II ventrally ornated with a row of long spiniform setiferous tubercles; coxae III and IV with a row of widely spaced tubercles. Stigmatic area reverse “T” shaped, spiracles not hidden. FIG. 5. — Habitus of the holotype (CNAN-T01344) of Toojah cimutaa n. gen., n. sp. A , dorsal; B , lateral. Scale bars: 1.5 mm. FIG. 6. — Scutum and venter of the holotype (CNAN-T01344) of Toojah cimutaa n. gen., n. sp. A , scutum; B , venter. Arrow indicates lateral clear area. Scale bars 1.0 mm. Chelicera ( Fig. 7A, B ) Basichelicerite with long bulla, covering almost all segment. Cheliceral hand swollen, fixed finger with seven teeth, the first and the sixth larger, movable finger five teeth, the basalmost larger and blunt. Pedipalp ( Fig. 7C, D ) Trochanter with two large spiniform setiferous tubercles on ventral side, the distalmost larger; femur slightly compressed laterally, with a ventral row of seven spiniform setiferous tubercles, the first three larger and decreasing in size slightly; patella unarmed; tibia with IIi (1=2>3) and iiIi (3>1=2=4), major SST on mesal and ectal sides indicated on the figure, tarsus with II (1>2) on both sides, first setiferous tubercle on mesal margin very large, almost the same length than tarsus. Tarsal claw of the same size that tarsus. Legs ( Table 2 ) Trochanter III rounded,longer than other trochanters.All segments without armature, except apical portion of femur IV with two ventral rows of reduced tubercles. Tarsal count: 8(3):51(8):6:7. Penis ( Fig. 8 ) Base of pars distalis swollen, with the flimsy lamina thick in lateral view and the apical margin ventrally curved. Follis inflated and almost three times larger than wide, with a bilobular dorsal projection, spines only on the ventro-apical portion of follis, stylus inserted in it and with small bristles on the tip. Macrosetal arrangement Stygnopsis - type sensu Cruz-López & Francke (2017), all MS with similar size, spatulate at the tip and with the bases well marked and deep. MS arranged as follow: two pairs of MS E on the middle of flimsy lamina, two pairs of MS C lateral to the base of follis, a pair of MS A dorsally, below the base of follis and a pair of MS B ventrally, near the base of pars distalis. Female Unknown.