A revision of the Neogene Conidae and Conorbidae (Gastropoda) of the Paratethys Sea Author Harzhauser, Mathias Author Landau, Bernard text Zootaxa 2016 4210 1 1 178 journal article 37280 10.11646/zootaxa.4210.1.1 e782e07d-76b7-4e9b-ba34-ed3286254ec6 1175-5326 252966 D39416B8-CF85-440B-84C2-D4380BECC4E3 Kalloconus voeslauensis ( Hoernes & Auinger, 1879 ) Figs 3 U, 14C1–C3,14D1–D3 Conus clavatus Lam. —Hörnes 1851: 25 (partim), pl. 2, figs 5a–b. [ Dendroconus ] [ Conus ] Vöslauensis n. f.— Hoernes 1878a : 195 (nomen nudum). Conus ( Dendroconus ) Voeslauensis nov. form.— Hoernes & Auinger 1879 (partim): 22, pl. 1, fig. 8, pl. 3, fig. 4. [ Conus ( Chelyconus ) mediterraneus ] var. ponderoclavata Sacc. — Sacco 1893b : 104 ( nov. nom. pro Conus clavatus in Hörnes 1851 , pl. 2, fig. 5). Conus ( Dendroconus ) voeslauensis Hoernes et Auinger—Csepreghy-Meznerics 1956: 122, pl. 11, figs 3–4. Conus ( Lithoconus ?) voeslauensis Hoernes & Auinger, 1879 — Strausz 1966 : 456 , pl. 68, figs 1–2. Conus ( Dendroconus ) voeslauensis Hoernes & Auinger—Chira & Voia 2001 : 156 , pl. 4, figs 4a–b. non Conus cf. Voeslauensis R. Hoern. et Au.— Bauer 1900 : 21 , pl. 1, fig. 1. non Conus ( Dendroconus ) voeslauensis Hoernes und Auinger 1879 —Kojumdgieva in Kojumdgieva & Strachimirov 1960 : 215 , pl. 51, fig. 4 [= Lautoconus steindachnerii (Hoernes, 1879) ] Type material. 6 syntypes NHMW 1849 /0023/0005, Bad Vöslau ( Austria ) including specimen illustrated in Hoernes & Auinger (1879, pl. 3, fig. 4) , 2 syntypes NHMW 1851 /0010/0002, Mikulov-Kienberk ( Czech Republic ), including specimen illustrated in Hoernes & Auinger (1879, pl. 1, fig. 8) ; all middle Miocene , Badenian ( Langhian ). Studied material. Syntypes and 3 spec. NHMW 1853 /0010/0005, Enzesfeld ( Austria ), 6 spec . NHMW A1633, Enzesfeld ( Austria ), 3 spec . NHMW 1849 /0033/0002, Bad Vöslau ; 1 spec . NHMW 1846 /0037/0049, Gainfarn ( Austria ), illustrated in Hörnes (1851, pl. 2, fig. 5). Illustrated material. Figs 14 C1–C3: syntype , Bad Vöslau ( Austria ): SL: 55.5 mm , MD: 34.4 mm , NHMW 1849/0023/0005a, specimen illustrated in Hoernes & Auinger (1879, pl. 3, fig. 4) ; Figs 14 D1–D3: syntype , Enzesfeld ( Austria ): SL: 52.0 mm, MD: 31.0 mm, NHMW 1853/0010/0005; Fig. 3 U: syntype , Bad Vöslau ( Austria ): SL: 42.8 mm , MD: 27.4 mm , NHMW 1849/0023/0005b. FIGURE 14A1–A3. Kalloconus tschermaki (Hoernes & Auinger, 1879) , Gainfarn (Austria), NHMW 1853/0003/0003a, syntype. 14B1–B4. Kalloconus tschermaki (Hoernes & Auinger, 1879) , Niederkreuzstetten (Austria), NHMW 1864/0001/0499, syntype. 14C1–C3. Kalloconus voeslauensis (Hoernes & Auinger, 1879) , Bad Vöslau (Austria), NHMW 1849/0023/0005a, syntype. 14D1–D3. Kalloconus voeslauensis (Hoernes & Auinger, 1879) , Enzesfeld (Austria), NHMW 1853/0010/0005, syntype. Revised description. Medium-sized solid shells; low conical to weakly coeloconoid spire; spire whorls bulgy and convex, delicate spiral threads on early whorls, later smooth except for prominent growth lines, suture deeply channelled. Subsutural flexure shallow, moderately curved, moderately asymmetrical. Shoulder rounded, rarely subangulate, position of maximum diameter slightly below; last whorl only weakly ventricose or straight-sided, slightly constricted at base, typically with several strongly incised growth lines. Faint spiral cords on base; entire whorl covered with delicate spiral sculpture, visible only in tangential light. Siphonal canal short, slightly recurved; siphonal fasciole weakly swollen, twisted with prominent growth lines. Aperture moderately narrow, widening towards siphonal canal. Colour pattern consisting of numerous spirals (no subquadratic dots can be observed as illustrated by Hoernes & Auinger 1879 ). Shell measurements and ratios. n = 10: largest specimen: SL: 61.8 mm , MD: 35.0 mm, mean SL: 50.2 mm (σ = 5.7), mean MD: 31.2 (σ = 2.8), spire angle: µ = 118.8° (σ = 4.7°), last whorl angle: µ = 38.9° (σ = 1.0°), LW: µ = 1.6 (σ = 0.08), RD: µ = 0.71 (σ = 0.03), PMD: µ = 0.87 (σ = 0.02), RSH: µ = 0.12 (σ = 0.01). Discussion. Most shells which can be reliably identified as Kalloconus voeslauensis are characterised by several deeply incised growth lines on the last whorl. In their original description of this species, Hoernes & Auinger (1879) referred also to specimens from Lăpugiu de Sus, which, however, represent Kalloconus hungaricus . The specimen illustrated by Hörnes (1851, pl. 2, figs 5a–b) as Conus clavatus is an aberrant specimen of Kalloconus voeslauensis . Sacco (1893b) introduced ponderoclavata as varietal name for this aberrant specimen, which thus is a subjective junior synonym of Conus voeslauensis . Paleoenvironment. The specimens from Bad Vöslau are filled with clay and might have dwelled in basinal settings. Mikulov-Kienberk and Enzesfeld, in contrast, represent shallow marine environments (own data). Distribution in Paratethys. Badenian (middle Miocene): Alpine-Carpathian Foredeep: Grund ( Austria ); Vienna Basin: Bad Vöslau, Gainfarn ( Austria ), Mikulov-Kienberk ( Czech Republic ) ( Hoernes & Auinger 1879 ); Eisenstadt-Sopron Basin : Forchtenau ( Austria ) ( Sieber 1956 ); Transylvanian Basin : Lăpugiu de Sus ( Romania ) ( Chira & Voia 2001 ); Pannonian Basin: Szob, Budapest , Mecsek Mountains ( Hungary ) ( Csepreghy-Meznerics 1956 ; Strausz 1966 ).