The genus Clavariadelphus (Clavariadelphaceae, Gomphales) in China
Author
Huang, Hong-Yan
School of Pharmaceutical Sciences and Yunnan Key Laboratory of Pharmacology for Natural Products, Kunming Medical University, Kunming, 650500, China
Author
Zhao, Jie
School of Pharmaceutical Sciences and Yunnan Key Laboratory of Pharmacology for Natural Products, Kunming Medical University, Kunming, 650500, China
Author
Zhang, Ping
College of Life Science, Hunan Normal University, Changsha, 410081, China
Author
Ge, Zai-Wei
Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, 650201, China
https://orcid.org/0000-0003-3184-4604
Author
Li, Xian
School of Pharmaceutical Sciences and Yunnan Key Laboratory of Pharmacology for Natural Products, Kunming Medical University, Kunming, 650500, China
xianlikm@163.com
Author
Tang, Li-Ping
School of Pharmaceutical Sciences and Yunnan Key Laboratory of Pharmacology for Natural Products, Kunming Medical University, Kunming, 650500, China
lipingtang11@qq.com
text
MycoKeys
2020
70
89
121
http://dx.doi.org/10.3897/mycokeys.70.54149
journal article
http://dx.doi.org/10.3897/mycokeys.70.54149
1314-4049-70-89
65FFE6B3A0F25CEF8D1B84BCEB9F6D24
3.
Clavariadelphus elongatus J. Khan, Sher & Khalid, Phytotaxa 365: 184, 2018
Figs 2d
, 2e
, 3c
, 4c
, 5c
, 8a
, 8b
Note.
The following description is taken from
Sher et al. (2018)
, field notes of the Chinese material including macro-morphology, growth habit, distribution, host plants and our examination of the specimens.
Figure 8.
Microscopic features of
Clavariadelphus elongatus
(HKAS 76589).
a
Leptocystidia
and immature basidia
b
Basidia.
Description.
Basidiomes
up to 28 cm high, 0.5-1.0 cm diam. basally, enlarged upwards to 1.5 cm diam., subcylindrical to fusiform, simple or occasionally branched, laterally compressed in age;
hymenium
longitudinally rugose, plum colour (13C2-4) or light purple to greyish-purple (14C2-3) or dull-lilac (15D2-3);
apex
tapered, subacute to obtuse, initially smooth, rugulose in age, caramel-brown to sandy-brown or sienna (6C5-6);
base
terete, smooth, white;
mycelial hyphae
scant, white;
flesh
initially solid, then soft and spongy in age.
Odour
and
taste
not recorded.
Hymenium
extending over the apex of the basidiomata, composed of basidia and leptocystidia.
Basidia
75-95
x
6-10
μm
, clavate, hyaline, thin-walled, 4-spored, sterigmata 7-10
μm
in length.
Basidiospores
[40/2/2] (8.3-) 9.0-11.0 (-12.0)
x
(5.5-) 5.7-7.4
μm
,
Q
= (1.43-) 1.44-2.04 (-2.31),
Q
m = 1.71
+/-
0.16, narrowly ellipsoid to ellipsoid, ovate or amygdaliform, with a small apiculus, inamyloid, thin-walled, hyaline in KOH, smooth.
Leptocystidia
70-75
x
3.5-4.5
μm
, scattered amongst and scarcely projecting beyond the basidia, cylindrical to narrowly clavate, thin-walled, smooth, hyaline, non-pigmented, clamped, inflated apically at maturity, at times with apical or subapical branches.
Mycelial hyphae
2-3 or 6-8
μm
diam., interwoven or aggregated into rhizomorphic strands, branched, clamped; the hyphal walls echinulate with light microscopy, encrusted with massive triangular or irregular, flaky crystals up 1
μm
high, which are insoluble in KOH.
Chemical reactions.
(dried basidiomes): KOH = positive, light yellow; FeCl3 = positive, green-yellow; NH4OH = positive, orange; ethanol, FeSO4, phenol and
Melzer's
reagent = negative.
Known distribution and ecology.
NW and SW China (in this study), Pakistan (
Sher et al. 2018
). Solitary to scattered on the ground in coniferous woods (
Abies
spp. and
Picea
spp.) or mixed with broad-leaved trees (
Quercus
spp.,
Rhododendron
spp. and
Salix
spp.) at elevations ranging from 3000-4350 m.
Materials examined.
China. Gansu Province: Zhouqu Prefecture, Shatan National Forest Park,
Abies
spp. woods, 16 August 2012,
X.T Zhu 740
(HKAS 76589). Sichuan Province: Litang Prefecture, Gaowa, Kobresia-Bistorta meadows with extensive areas of dwarf
Rhododendron
and
Salix
scrub with
Picea
spp.,
30°10.10'N
,
100°35.12'E
, alt. 4300-4350 m, 8 August 2006,
Z.W. Ge 1221
(HKAS 50801); Yajiang Prefecture, meadows with shrub thickets and
Picea
spp. forests,
30°2.67'N
,
101°18.48'E
, alt. 3850-3870 m, 4 August 2006,
Z.W. Ge 1162
(HKAS 50742). Yunnan Province: Yulong Prefecture, Lizui Village, mixed coniferous and broad-leaved forests of
Picea
spp. and
Quercus
spp., alt. 3000 m, 23 August 2007,
Y. Zhang 36
(HKAS 52425); Shangri-La Prefecture,
27°29.00'N
,
99°25.00'E
, alt. 3600 m, 13 August 2008,
T.Z. Wei 150
(HMAS 260746).
Comments.
Clavariadelphus elongatus
was originally described from Pakistan (
Sher et al. 2018
). In this study, it was found in NW and SW China. This species is unique in its greyish-purple basidiomes with acute to subacute, non-enlarged apex, hyphae of the basal mycelium encrusted with massive, flaky crystals and basidiomes having a light yellow reaction to KOH.
Clavariadelphus himalayensis
, another Asian taxon, might be confused with
C. elongatus
since both have a tinge of grey-purple when young. However,
C. himalayensis
is distinct in having smaller basidiomes, pastel-red colouration at maturation, shorter basidiospores (8.2-9.4
x
5.0-5.5
μm
), hyphae of the basal mycelium covered nipple-shaped protuberances without crystals and basidiomes having a brown-yellow reaction to KOH.
Phylogenetically,
C. elongatus
is related to
C. pistillaris
and the sequence of "
C. occidentalis
" from GenBank with weak support (Fig.
1
).