Nine new spider species of Belisana Thorell, 1898 (Araneae, Pholcidae) from karst caves, with a list of species of the genus from Guizhou, southwestern China Author Wang, Bing https://orcid.org/0009-0009-7047-4395 College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China Author Li, Jinglin https://orcid.org/0009-0008-3932-309X College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China Author Li, Shuqiang 0000-0002-3290-5416 Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China Author Yao, Zhiyuan 0000-0002-1631-0949 College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China text ZooKeys 2024 2024-10-25 1216 265 302 journal article 10.3897/zookeys.1216.132561 0E173F72-36D3-4B97-BC43-F096067A3D1E Belisana zhangi Tong & Li, 2007 Figs 16 , 17 , 23 C, D , 25 E, F Belisana zhangi Tong and Li 2007: 505 , figs 1–6. Material examined. China • 1 ♂ ; Guizhou , Anshun , Puding County , Huachu Town , Jinqian Cave ; 26 ° 14.835 ' N , 105 ° 37.521 ' E ; 3 May 2005 ; collector unknown; IZCAS - Ar 45177 • 3 ♀ ; same data as for preceding; IZCAS - Ar 45178 –80 . Diagnosis. The species resembles B. yuhaoi Yang & Yao, 2023 ( Yang et al. 2023 a : 178, figs 2 A, B, 3 A – D, 4 A – H) by having similar bulbal apophysis (hooked; Fig. 17 C ) and epigyne (epigynal pockets on posterior part of epigynal plate, epigynal plate posteriorly curved; Figs 17 A , 25 E ), but can be distinguished by procursus with nearly angular ventro-subdistal membranous lamella (arrow 1 in Figs 16 C , 23 C vs nearly round) and curved retrolateral membranous flap (f in Figs 16 D , 23 D vs angular), by male cheliceral distal apophyses pointing inwards ( da in Fig. 17 D vs downwards), and by vulval pore plates long elliptic ( pp in Figs 17 B , 25 F vs nearly triangular). Figure 16. Belisana zhangi Tong & Li, 2007 , male A, B palp ( A prolateral view B retrolateral view, arrow 1 points at ventral apophysis, arrow 2 points at retrolatero-proximal protrusion) C, D distal part of procursus ( C prolateral view, arrow 1 points at ventro-subdistal membranous lamella, arrow 2 points at prolatero-distal membranous lamella D retrolateral view, arrow 1 points at sclerotized dorso-subdistal apophysis, arrow 2 points at retrolatero-distal spine). Abbreviations: b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.10 ( A, B ); 0.02 ( C, D ). Figure 17. Belisana zhangi Tong & Li, 2007 , male ( C – F ) and female ( A, B, G, H ) A epigyne, ventral view B vulva, dorsal view C bulb, prolateral view D chelicerae, frontal view E – H habitus ( E, G dorsal view F lateral view H ventral view). Abbreviations: aa = anterior arch, b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.10 ( A – D ); 0.40 ( E – H ). Habitat. The species was found in the dark zone inside the cave. Distribution. China ( Guangxi , type locality; Guizhou , Fig. 1 ).