A review of the genus Aponema Jensen, 1978 (Nematoda: Microlaimidae) with description of three new species
Author
Kovalyev, Shota V.
Author
Miljutina, Maria A.
text
Zootaxa
2009
2077
56
68
journal article
10.5281/zenodo.187255
2ef5a369-3da8-4342-8d3d-b57d32249c1b
1175-5326
187255
Aponema
Jensen 1978
Emended diagnosis.
Microlaimidae
. Males monorchic with single outstretched anterior testis. Females didelphic-amphidelphic with outstretched ovaries. Amphidial fovea monospiral, outline rounded. Somatic setae inserted directly on body cuticle, never on a process. Cuticle with transverse striae or optically smooth, without thorns. Pharyngeal bulb varying in shape: from spherical to pyriform. Gubernaculum with or without apophyses. Tail conical or elongated.
Type
species.
A. torosum
(
Lorenzen, 1973
)
Jensen, 1978
Remarks.
The tendency to reduction of the posterior testis occurs among three other
Microlaimidae
genera. Among
Microlaimus
species the posterior testis in
M. globiceps
De
Man
, 1880
(see
Turpeenniemi, 1997
) and
M. copulatus
Jensen, 1988
is only half the size of the anterior testis. In the only species of genus
Acanthomicrolaimus
Stewart & Nicholas, 1987
—A.
jenseni
Stewart & Nicholas, 1987
—the posterior testis is also much shorter than the anterior one, and gubernacular apophyses are present as in several
Aponema
species (
Stewart & Nicholas, 1987
). This genus differs from
Aponema
not only by the presence of a short posterior testis but also in the structure of the cuticle, which bears numerous thorns.
Bathynox
Bussau & Vopel, 1999
, like
Aponema
possesses only one anterior testis, but its amphids are located very far from the anterior tip of head and somatic setae are situated on short processes, unlike
Aponema
.
In
Microlaimus nanus
Blome, 1982
, the posterior testis is absent, and this allows us to relate this species to
Aponema
Jensen, 1978
and to regard it as
Aponema nanum
(
Blome, 1982
)
comb.n.
Nevertheless,
Muthumbi & Vincx (1999)
described two
Aponema
having well-developed anterior and posterior testes—
A. decramerae
Muthumbi & Vincx 1999
and
A. mnazi
Muthumbi & Vincx 1999
—without emending the diagnosis of the genus
Aponema
.
We suppose that the main difference of
Aponema
from other Microlaimids is the lack of a posterior testis so it seems doubtful that
A. decraemerae
and
A. mnazi
should be considered as belonging to the genus
Aponema
and we propose to transfer these species to
Microlaimus
.