The artoriine wolf spiders of Australia: the new genus Kochosa and a key to genera (Araneae: Lycosidae) Author Framenau, Volker W. Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150. & Department of Terrestrial Zoology, Western Australian Museum, Welshpool DC, Western Australia 6986, Australia & Zoological Museum Hamburg, Leibnitz Institute for the Analysis of Biodiversity Change (LIB), Centre for Taxonomy & Morphology, Author Castanheira, Pedro De S. 0000-0002-0623-1622 Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150. & https // orcid. org / 0000 - 0002 - 0623 - 1622 Author Yoo, Jung-Sun 0000-0002-3243-2006 Animal Resources Division, National Institute of Biological Resources, Environmental Research Complex, Incheon, 22689, Korea https // orcid. org / 0000 - 0002 - 3243 - 2006 text Zootaxa 2023 2023-02-10 5239 3 301 357 http://dx.doi.org/10.11646/zootaxa.5239.3.1 journal article 10.11646/zootaxa.5239.3.1 1175-5326 7634797 BF1FF837-56D5-4829-8D46-E821D9D31AB3 Kochosa timwintoni sp. nov. ( Figs 27 , 28A–E , 29A–D ) Holotype . Male , West Mt Barren ( 34º13'S 119º26'E , Western Australia , AUSTRALIA ), R. J. McKay , 15 July 1970 , on sand ( WAM T86692 ). Etymology. The specific epithet is a patronym honouring the Western Australian author Tim Winton for his ongoing environmental advocacy (i.e., Winton 2008 ). His novels have given the senior author hours of pleasure and time of reflection. Other material examined ( 4 males , 6 females ). Australia . Western Australia : 1 male , West Mt Barren , 34º13'S 119º26'E ( WAM 71 /499) ; 2 females , same locality ( WAM 71 /856–7) ; 1 female ( WAM 70 /209), same locality ; 2 males , same locality ( WAM T86693 ) ; 3 females , same locality ( WAM70 /246a–c) . Diagnosis. Males of K. timwintoni sp. nov. can be easily separated from other species of Kochosa gen. nov. by the unique light brown to yellow, broad base of the embolus ( Fig. 28C, E ). They are most similar to those of K. westralia sp. nov. , but in addition to the embolus, the first species has a triangular and pointing retrolaterally basoembolic apophysis, whereas it is broad and round in K. westralia sp. nov. ( Figs 28E vs 31E). The epigyne of female K. timwintoni sp. nov. has a distinct median septum similar to K. obelix sp. nov. and K. westralia sp. nov. , but the posterior transverse part is much broader, so that the median septum forms and inverted “T” ( Figs 29C vs 19D, 32C). Description. Male ( based on holotype , WAM T86692 ) . Cephalothorax . Dorsally dark brown; indistinctly lighter medially ( Fig. 28A ); broad lateral bands with white setae ( Fig. 28A ); Sternum dark shiny brown ( Fig. 28B ). Abdomen . Dorsally dark olive-grey; cardiac mark continuous, but almost dissolved into two spots ( Fig. 28A ). Venter uniformly olive-grey ( Fig. 28B ). Pedipalps ( Figs 28C–E ). Cymbium comparatively broad; embolic division almost entirely exposed; tegular apophysis reduced, shark-tooth-shaped; basoembolic apophysis somewhat duckbill-shaped, pointing retrolatrally ( Fig. 28E ); embolus with light brown base, apically stout. Legs. Dark brown, with light annulations; spination of leg I: femur: 3 dorsal (apical one small), 1 apicoprolateral; tibia: 3 ventral pairs, 1 prolateral; metatarsus: 3 ventral pairs, 1 apicoventral, 2 prolateral, 1 apicoprolateral, 1 retrolateral, 1 apicoretrolateral. Measurements . TL 3.91, CL 2.34, CW 1.64. Eyes: AME 0.09, ALE 0.08, PME 0.26, PLE 0.21. Row of eyes: AE 0.47, PME 0.70, PLE 0.86. Sternum (length/width) 0.98/0.86. Labium (length/width) 0.19/0.30. AL 1.72, AW 1.56. Legs: Length of segments (femur + patella/tibia + metatarsus + tarsus = total length): Pedipalp 0.81+0.56+- +0.81=2.19, I 1.53+1.63+1.28+0.86=5.30; II 1.35+1.72+1.40+0.81=5.28, III 1.40+1.63+1.21+0.70=4.93; IV 1.93 +2.12+2.02+0.91=6.98. Variation . Size (range, mean ± s.d.): TL 3.91–4.40, 4.07 ± 0.22; CL 2.30–2.70, 2.44 ± 0.18; CW 1.64–2.00, 1.81 ± 0.15, n = 4. There is little colour variation between the males examined here except for somewhat lighter legs in one specimen . Female (based on WAM 70/246a; epigyne dorsal WAM 70/246b). Cephalothorax and abdomen . Colouration and setae-arrangement generally as male, but light median and lateral carapace bands less distinct ( Fig. 29A, B ). FIGURE 28A–E. Kochosa timwintoni sp. nov. , male holotype, WAM T86692. A, B, male habitus, dorsal and ventral view; C, D, male, left pedipalp, ventral and retrolateral view; E , male, left pedipalp, apical section. Scale bars: A, B , 1.0 mm; C, D, 0.2 mm; E , 0.1 mm. Epigyne ( Fig. 29C, D ). Ventral view: median septum inverted T-shaped ( Fig. 29C ); dorsal view (based on WAM 70/246b): spermathecal heads spherical, comparatively large, spermathecal stalks basally curved ( Fig. 29D ). Legs . Light brown with dark annulations; spination of leg I: femur: 3 dorsal; tibia: 3 ventral pairs, 1 prolateral; metatarsus: 3 ventral pairs and 1 apicoventral, 2 prolateral; 1 apicoprolateral; 1 retrolateral; 1 apicoretrolateral. FIGURE 29A–D. Kochosa timwintoni sp. nov. , female, WAM 70/246a ( A–C ) and WAM 70/246b ( D ). A, B, female habitus, dorsal and ventral view; C, epigyne, epigyne, ventral view; D , epigyne, dorsal view. Scale bars: A, B , 1.0 m; C, D , 0.2 mm. Measurements . TL 5.50, CL 2.55, CW 1.70. Eyes:AME 0.11, ALE 0.09, PME 0.25, PLE 0.23. Row of eyes: AE 0.67, PME 0.85, PLE 1.16. Sternum (length/width) 1.00/0.90. Labium (length/width) 0.40/0.47. AL 2.60, AW 2.30. Pedipalp 0.75+0.77+-+0.75=2.27, I 1.61+1.75+1.10+0.65=5.10, II 1.45+1.66+1.05+0.65=4.81, III 1.25+1.71+1.26 +0.65=4.76, IV 2.05+2.01+2.01+0.75=6.82. Variation . Size (range, mean ± s.d.): TL 5.32–5.62, 5.46 ± 0.13; CL 2.50–2.55, 2.50 ± 0.08; CW 1.70–1.95, 1.81 ± 0.10, n = 4. The females of K. timwintoni sp. nov. are generally poorly preserved and it is therefore difficult to assess any colour variation. Life history and habitat preferences. Males and females of K. timwintoni sp. nov. were found in July and August suggesting reproductive activity in winter. The habitat preferences appear unusual, as specimens were found “on sand” or “on sand plain”, in contrast to most other species of Kochosa gen. nov. which appear to prefer forests or woodland. Distribution. Only know from the type locality, West Mt Barren, Western Australia ( Fig. 27 ).