Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae) Author Pall-Gergely, Barna https://orcid.org/0000-0002-6167-7221 Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary pallgergely2@gmail.com Author Hunyadi, Andras Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary Author Vermeulen, Jaap J. JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands Author Grego, Jozef Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia Author Sutcharit, Chirasak Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand Author Reischuetz, Alexander Puechhaimgasse 52, A- 3580 Horn, Austria Author Dumrongrojwattana, Pongrat Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand Author Botta-Dukat, Zoltan Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary Author Oerstan, Aydin 12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA Author Fekete, Judit University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary & Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary Author Jochum, Adrienne https://orcid.org/0000-0002-6624-6412 Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland & Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland & Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany text ZooKeys 2023 2023-02-13 1147 1 177 http://dx.doi.org/10.3897/zookeys.1147.93824 journal article http://dx.doi.org/10.3897/zookeys.1147.93824 1313-2970-1147-1 9BB9881B0076473D8E53155D37CA1F50 FF2B6B317B505F9EA0E1000BDCD16CE7 Angustopila prolixa Pall-Gergely & Hunyadi sp. nov. Fig. 53 Type material. Holotype : Vietnam • 1 empty shell (H: 0.67 mm, D: 0.73 mm); Lạng Sơn Province, Hữu Lũng District, Minh Tiến, 1400 m northeast Cầu Cheo Minh Tiến (locality code: 2020/55); 21°34.02'N , 106°17.79'E ; 30 m a.s.l.; 20 Feb. 2020; A. Hunyadi leg.; HNHM 105290. Paratypes : Vietnam • 12 shells; same data as for holotype; coll. HA. Additional material. Vietnam • 1 j/b shell; same data as for holotype; coll. HA. Diagnosis. A small, domed Angustopila species with a very high and long parietal tooth. Description. Shell small for the genus, slightly wider than high or as wide as high; transparent, pale grey, dorsal side domed; body whorl widest from standard apertural view; protoconch consists of almost 1.75 whorls, with strong spiral striation; teleoconch ornamented by weak, irregular radial growth lines and irregularly spaced, much stronger spiral striae (ca. 10 or 11 on body whorl from standard apertural view); whorls 3.75, rounded or depressed-rounded; aperture slightly oblique to shell axis from lateral view; umbilicus very wide; parietal callus detached from penultimate whorl; aperture pear-shaped; peristome expanded but not reflected; parietal tooth starts from parietal callus, strongly elongated inside aperture, conspicuously elevated (extends beyond 1/2 aperture), slightly curves in direction of palatal wall. Figure 53. Angustopila prolixa Pall-Gergely & Hunyadi, sp. nov. Holotype (HNHM 105290): A, C-G, I-J ), paratypes (2020/55, specimen 1: B 2020/55, specimen 3: H ). Apertural ( A, B ), lateral ( C ), apical ( D ) and ventral ( E ) sides of the shell; aperture ( F ), sculpture on the protoconch ( G ), parietal tooth ( H ); ventral ( I ) and frontal ( J ) surface of the body whorl. Measurements (in mm). H = 0.67-0.77, D = 0.73-0.78, H/D*100 = 91.8-100 ( n = 5), RUD = 33.8-36.4 ( n = 2). Differential diagnosis. The nearby occurring, A. vomer sp. nov., is similar in possessing a highly elevated, long parietal tooth. However, it has a larger, concave-conical (rather than domed) shell, a narrower umbilicus, a more rounded aperture (wider sinulus), and its parietal tooth does not turn towards the palatal wall, and does not reach the parietal callus. The most similar species in terms of shell and aperture shape is A. quadridens sp. nov., which possesses four teeth. Etymology. The specific epithet ( prolixo = elongated in Latin) refers to the unusually long parietal tooth of this new species. Distribution. This new species is known only from its type locality (Fig. 27 ).