The recent apple snails of Africa and Asia (Mollusca: Gastropoda: Ampullariidae: Afropomus, Forbesopomus, Lanistes, Pila, Saulea): a nomenclatural and type catalogue. The apple snails of the Americas: addenda and corrigenda
Author
Cowie, Robert H.
text
Zootaxa
2015
3940
1
1
92
journal article
10.11646/zootaxa.3940.1.1
5013eaff-bb37-4be3-a1ea-43c11d0cf5d1
1175-5326
288204
B5D45D1B-4B52-4F0B-8AF6-B587F8857475
speciosa
Ampullaria speciosa
Philippi, 1849b
: 18
[1852a: 40, pl. 11, fig. 2].
Type
material
—possible
syntypes
: MNHNCL 15900 (2 opercula, old number 61.042), ZMB 1323 (
Köhler & Glaubrecht 2006: 210
).
Type
locality
—“
India
orientalis
” [error; possibly Cape Guardafui,
Somalia
(
Philippi 1852a
: 41
;
Brown 1980
: 47
, 1994, 55;
Köhler & Glaubrecht 2006
: 210
)].
Distribution
—“Practically the whole of eastern tropical Africa” (
Alderson 1925: 98
), but restricted to
Somalia
, northern
Kenya
and southeastern
Ethiopia
,
teste
Pain
(1949b: 69)
and
Brown (1965
: 44
, 1980: 47, 1994: 55), followed by
Köhler & Glaubrecht (2006: 211)
.
Remarks
.
Pila
,
teste
Pain
(1949b: 69)
and
Brown (1965
: 44, 1980: 47, 1994: 55), followed by
Köhler & Glaubrecht (2006: 210)
.
sphaerica
Ampullaria sphaerica
Deshayes, 1830a
: 30
.
Type
material
—location(s) unknown [not found in MNHN or NHMUK (cf.
Dance 1986
: 210
, 217)].
Type
locality
—“Pondichéri” [the locality of one of Deshayes’ specimens (that of M. Marmin), which he presumed to be the locality of the others; locality of Sowerby’s shell, referenced by Deshayes, unknown].
Remarks
. In addition to the shells of
Deshayes (1830a: 30)
the shell illustrated by G.B. Sowerby I (1822: pl. [184], figs. 1, 2) and referenced by Deshayes is part of the
type
series. By implication,
rugosa
G.B. Sowerby I
non
Lamarck (considered a species of
Pila
based on Sowerby’s illustration of the calcified operculum) was considered by Swainson (1822a: pl. 119),
Deshayes (1830a: 30)
and
Prashad (1925: 70)
to be a misidentification of
rugosa
Lamarck
(synonym of
urceus
Müller
, a species of
Pomacea
,
teste
Cowie & Thiengo (2003: 74))
rather than the proposal of a new name. Swainson proposed the name
corrugata
for Sowerby’s shell, prior to Deshayes’ proposal of
sphaerica
. Synonym of
corrugata
Swainson
,
teste
Philippi (1851: 7)
and Martens (1857: 209), followed here.
sphaerica
Ampullaria globosa
var.
sphaerica
Hanley & Theobald, 1874: 46
, pl. 113, fig. 4.
Type
material
—
syntype
: UMZC I.104725 (1 spm., with label “figured in Conch. Indica CXIII, f. 4”) [not found in Leeds Museum or NHMUK (
Coan & Kabat 2012: 325
)].
Type
locality
—“Moradabad”.
Remarks
. Junior primary homonym of
sphaerica
Deshayes.
Variety of
globosa
Swainson
,
teste
G.B. Sowerby III (1910: 58) and
Preston (1915: 97)
. Synonym of
globosa
Swainson
,
teste
Nevill (1877: 1)
and
Prashad (1923
: 587, 1925: 71), followed here.
stoliczkana
Ampullaria Stoliczkana
Nevill, 1877
: 11
.
Type
material
—possible
syntypes
: NZSI 2420 (1 spm., considered a
holotype
in NZSI), NZSI M 25079/5 (7 spms., considered
paratypes
in NZSI).
Type
locality
—“Penang”.
Remarks
.
Nevill (1877: 11)
noted 7 spms. but subsequently (
Nevill 1885: 7
) listed only 6; the reason that there are now 8 considered as
type
material in NZSI requires further research. Variety of
conica
Wood
,
teste
G.B. Sowerby III (1910: 50) and
Alderson (1925: 78)
. Synonym of
scutata
Mousson
,
teste
Low
et al
. (2013
: 56) and Ng
et al
. (2014: 34), followed here.
+
stuhlmanni
Ampullaria erythrostoma
Var.
stuhlmanni
Martens, 1897a: 155
–156, text fig.
Type
material
—
syntypes
: ZMB 109527 (1 spm.), ZMB
111742
(1 spm.), the additional
syntypes
not found.
Type
locality
—“Albert-Nyansa, an der Südwestseite bei Kassenye” [southwestern side of Lake Albert near Kassenye], “Kassesse in Karagwe”, “Albert- Edward-See... bei Katarenga an der Südwestseite” [southwestern side of Lake Edward, near Katarenga].
Distribution
—Lakes Albert and Kyoga, Victoria Nile (
Pain
1961: 11–12
) and Lake Edward (
Dartevelle & Schwetz 1947
: 15, 17;
Köhler & Glaubrecht 2006
: 211).
Remarks
. The placement of this taxon by Martens (1897a: 155) as a variety of
erythrostoma
Reeve
is in error as the latter is a South American species (
Cowie & Thiengo 2003
: 63;
Köhler & Glaubrecht 2006
: 211). The two large specimens from “Albert-Nyanza” and the one from “Kassesse in Karagwe” (Martens 1897a: 155) were not found by
Köhler & Glaubrecht (2006: 211)
. The parenthetical “
type
Stuhlmann” in the tables of shell dimensions given by
Pain
(1952: 289, 1961: 13) is here considered as referring to the
type
locality and not to be
lectotype
designations. Treated as a distinct species by
Pilsbry & Bequaert (1927: 182–183)
and
Dartevelle & Schwetz (1947: 32)
, but a subspecies of
ovata
Olivier
,
teste
Pain
(1952: 288, 1961: 13) and
Mandahl-Barth (1954: 45)
, followed here, all in
Pila
. Form of
ovata
Olivier
,
teste
Brown (1980: 45)
, though not so listed by
Brown (1994: 54)
.
subampullacea
Ampullaria turbinis var subampullacea
Nevill, 1885
: 6
.
Type
material
—
syntypes
: NZSI (2 spms., not found in 2014), H. Crosse collection (1 spm., not found in MNHN in 2014).
Type
locality
—“Perak” [Peninsular
Malaysia
].
Remarks
.
Annandale (1920a: 19)
indicated a “Type-specimen” (NZSI “2427 M”) but whether this was one of the two specimens deposited by Nevill has not been definitively determined. Neither it nor other material identifiable as Nevill’s
types
were found in NZSI in 2014. Var iety or race of
turbinis
Lea
,
teste
G.B. Sowerby III (1910: 61) and
Annandale (1920a: 5)
, respectively. Synonym of
ampullacea
Linnaeus
,
teste
Paetel (1888: 481) and
Brandt (1974: 49)
, followed here.
subglobosa
Ampullaria turbinis var. subglobosa
Nevill, 1885
: 6
.
Type
material
—
syntypes
: NZSI (4 spms., not found in 2014).
Type
locality
—“Java (?)”, “Possibly from
Siam
” [
Thailand
].
Remarks
. Synonym or variety of
turbinis
Lea
,
teste
Paetel (1888: 482). Variety of
turbinis
Lea
,
teste
G.B. Sowerby III (1910: 61). Synonym of
turbinis
Lea
,
teste
Annandale (1920a: 15)
. Possibly a senior synonym of
dalyi
Blanford,
teste
Alderson (1925: 65)
. Synonym of
ampullacea
Linnaeus
,
teste
Brandt (1974: 49)
, followed here.
subscutata
Ampullaria subscutata
Mousson, 1882
: 46
, pl. 3, fig. 6.
Type
material
—
syntypes
: RBINS MT 1156/1 (2 spms.), RBINS MT 1156/2 (4 spms.); possible
syntypes
: ZMZ 564075 (2 spms.).
Type
locality
—“
Madagascar
; Maravare, à la latitude de Farafanga, dans la partie orientale de l’île”.
Remarks
. Valid species,
teste
G.B. Sowerby III (1910: 61), Kobelt (1912b: 57) and
Alderson (1925: 91)
. Synonym of
cecillei
Philippi,
teste
Starmühlner (1969: 119)
and
Fischer & Vukadinovic (1974: 349)
, followed here. The names
subsulcata
of Paetel (1888: 482) and of
Kobelt (1909: 91)
are considered incorrect subsequent spellings of
subscutata
Mousson
and therefore unavailable (
Code
, Art. 33.3).
sumatrensis
Ampullaria sumatrensis
Philippi, 1852b
: 28
[1852a: 59, pl. 19, figs. 1, 2].
Type
material
—location unknown.
Type
locality
—“der Fluss Danu-luar in Sumatra” [Danu-luar river, Sumatra]; “Java?”.
Remarks
. Synonym of
ampullacea
Linnaeus
,
teste
G.B. Sowerby III (1910: 56), Kobelt (1912b: 76),
Habe (1964: 49; as “
ampullaria
”)
,
Brandt (1974: 49)
and Ng
et al
. (2014: 32). Recorded from Perak (Paetel 1888: 481), throughout peninsular
Malaysia
(de
Morgan 1885
: 418) and in
Cambodia
(Dautzenberg & Fischer 1905: 427).