Taxonomic review of the Dichotomius (Luederwaldtinia) assifer (Eschscholtz) species-group (Coleoptera: Scarabaeidae) Author Nunes, Rafael V. Author Carvalho, Marcela S. G. Author Vaz-De-Mello, Fernando Z. text Zootaxa 2016 4078 1 230 244 journal article 46800 10.11646/zootaxa.4078.1.21 3e49efd7-cb51-4214-8a47-0286df6e3dfa 1175-5326 264975 9CCA098E-8430-47A2-8100-00500D49C65D Dichotomius ( Luederwaldtinia ) machadoi Martínez & Pereira, 1967 ( Figures 6 A–E) Dichotomius machadoi Martínez & Pereira, 1967 , p. 55 (description); Dichotomius machadoi , Vaz-de-Mello 2000, p. 193 (checklist) Material examined : Paratypes : BRASIL : Pernambuco . Serra Negra. V-1960 . ABM Machado.leg [ 1♂ , 1♀ MZSP ]; Additional specimens: Floresta. Serra Negra. Cow dung. Wet Forest. VII-2012 . RV Nunes & GM De-la- Torre [ 3♂ , 2♀ CEMT ]; Floresta. Reserva Biológica Serra Negra. XII-2012 . RV Nunes & GM De-la-Torre [ 4♂ CEMT ]. Diagnosis. This species is separated of the other species in the group by the following combination of characters: cephalic carina very low, with a weak pair of tubercles ( Figs 6 A–C); males excavation at the anterior portion of the hypomeron bearing few sparse setae; females excavation at the hypomeron very weak, almost absent; pronotal disc wrinkled and bearing deep punctures ( Figs 6 A–C) and elytral interstriae strongly wrinkled and lacking distinct punctures ( Figs 1 E, 6A–B). This species has a detailed description in the work of Martínez and Pereira (1967) and therefore does not need a re-description. Distribution. Since its description, D. machadoi has been collected only in "Brejo de Altitude" forests at the Biological Reserve of Serra Negra, municipalities of Floresta, Inajá and Tacaratu Pernambuco, Brazil ( Fig. 7 ). The Brejo de Altitude forests are moist forests found on the higher parts of hills and plateaus in the Brazilian Caatinga, a semi-arid brazilian biome.