Leafy liverworts of Chapada das Mesas National Park: a floristic survey and checklist of the leafy liverworts of Maranhão state, Brazil Author Silva, Jainara Pereira Laboratório de Sistemática Vegetal, Centro de Ciências Agrárias e Ambientais, Universidade Federal do Maranhão, Chapadinha, MA, Brazil Author Oliveira-da-Silva, Fúvio Rubens Programa de Pós-graduação em Ciências Biológicas, Botânica Tropical, Universidade Federal Rural da Amazônia / Museu Paraense Emílio Goeldi, Belém, PA, Brazil • FROS: oliveira. fuvio @ gmail. com https: // orcid. org / 0000 - 0002 - 4871 - 6740 &): 479 - 495. https: // doi. org / 10.15560 / 17.2.479 Author Ilkiu-Borges, Anna Luiza Museu Paraense Emílio Goeldi, Belém, PA, Brazil • ALIB: ilkiu-borges @ museu-goeldi. br https: // orcid. org / 0000 - 0002 - 1266 - 7211 Author Fernandes, Rozijane Santos Laboratório de Sistemática Vegetal, Centro de Ciências Agrárias e Ambientais, Universidade Federal do Maranhão, Chapadinha, MA, Brazil text Check List 2021 2021-03-11 17 1 479 495 journal article 10.15560/17.1.479 1809-127X 5380155 ** Frullania rio-janeirensis (Raddi) Ångstr. Figure 3C Material examined. BRAZIL • Maranhão , município de Carolina , Parque Nacional Chapada das Mesas , Rio Farinha , sobre tronco de árvore próximo à corrente d´água; 06°59′S , 049°09′W ; alt. 197 m ; 6 Jun. 2018 ; JAS Silva 203; CCAA 1645 . Identification. Plants green to brown, irregularly (bi)pinnate branched. Hemiphyll entire associated with a lobule. Leaves subimbricate to imbricate, orbicular, slightly convex, margins entire, dorsal base auriculate, apex rounded, slightly plane; median cells isodiametric, trigones nodulose. Lobules erect, subparallel to the stem, galeate with an oblong laminal portion, rarely laminate, beak long, not decurrent. Stylus filiform, 2‒3 cells in a row, 1‒2 cell wide at base. Underleaves imbricate, orbicular, 1/6 bifid, margins entire, recurved, apex rounded, base rounded to auriculate, plane to recurved, insertion line curved. Widely distributed in Brazil , this species is recognized by lobules galeate with a long beak and an oblong laminal portion (as large as the galeate portion) and perianth 4-keeled. Description and illustration. Schuster (1992: 254–260, fig. 816). Lejeuneaceae