A cladistic revision of Tor top us Needham & Murphy with description of the new genus Tortopsis (Ephemeroptera: Polymitarcyidae) Author Molineri, Carlos text Zootaxa 2010 2481 1 36 journal article 10.5281/zenodo.195431 2df8bd5a-e125-42aa-aa6f-ae3e78b83929 1175-5326 195431 Tortopsis sarae (Domínguez) New combination Tortopus sarae Domínguez 1985 : 71 ; Molineri 2008 : 13 . Material ( IML ): holotype and 16 paratypes female imagos from Argentina : Jujuy, 10 km N Ledesma, río Zora, 14-XII-1983 , E. Domínguez col. Other material reported in Molineri (2008) . New localities: 10 male and 10 female imagos from Argentina : Tucumán, Acheral, río Aranillas, 366 m , S 27º 6' 59.9" - W 65º 27' 43.9" , 29- XII-2007 , C. Molineri & J. Giordano cols.; 1 reared male adult and 15 nymphs from Argentina : Tucumán, ca. Manuela Pedraza, río Balderrama, 335 m , S 27º 12' 0.6" - W 65º 21' 30.3" , 14-XI-2009 , C. Molineri col. FIGURES 87–92. Tortopsis , female adults, head and pronotum. 87, T. bruchianus ; 88, T. limoncocha ; 89, T. obscuripennis ; 90, T. puella ; 91, T. sarae ; 92, T. spatula . Female imago. Head shaded black between ocelli, occiput without marks. Pronotum with sublateral gray lines not completely separated from the rest of the pigmented area. Furcasternal plates basally contiguous ( Fig. 45 ). Wings: veins C, Sc y R1 (and membrane in between) shaded with brownish gray, rest of wing whitish. Vein R3 and short intercalary before it absent in fore wings but some females present a weak vein in zig-zag formed by coalescence of cross veins, with a length of 3 cells or less. Abdominal terga widely pigmented except on sublateral dashes on terga II–VII, black thin median line present on terga I–VIII, tergum IX with a thin pale median line. Sternum VIII with C-shaped sockets ( Figs. 46–47 ). Egg. Suboval, no attachment structures. Length, 355–375 µm; width, 290–305 µm. The chorionic cover is smooth. FIGURES 93–96. Polymitarcyidae , nymphal head: 93, Asthenopus sp.; 94, Campsurus near violaceus ; 95, Tortopus harrisi ; 96, Tortopsis obscuripennis . Discussion and diagnosis. Tortopsis sarae is currently known from all the stages ( Domínguez 1985 , Molineri 2008 ). A short description of the female imago is given to complete its characterization. This species can be recognized by: 1) fore wing length 13.0–13.5 (male), 17.5–18.0 (female); 2) wings whitish translucent except C and Sc areas brownish; 3) parastyli moderately curved; 4) penes cylindrical; 5) female parastylus receptor C-shaped ( Figs. 46–47 ); 6) head shaded black behind median ocellus, occiput very pale, at most with a small grayish netting near hind margin; 7) female pronotum shaded as in Tortopsis obscuripennis .