A review of the New Caledonian Arpactophilus (Hymenoptera: Crabronidae)
Author
Breitkreuz, Laura C. V.
Author
Ohl, Michael
Author
Engel, Michael S.
text
Zootaxa
2016
4063
1
1
66
journal article
37206
10.11646/zootaxa.4063.1.1
9608b695-6310-4c63-bd94-46bda058e6d8
1175-5326
270412
80401ED8-C6BA-4420-9109-854C5CC1E88D
Arpactophilus tayo
sp. nov.
(
Figs. 99–100
)
Diagnosis.
Arpactophilus tayo
is the only New Caledonian species in the genus with the combination of a black clypeus with a medially projecting apical margin that has a shallow V-shaped incision (
Fig. 99
), a brown mandible, a continuous occipital carina (
Fig. 5
), two submarginal cells in the forewing, and a flat sternum II without a bulge (
Fig. 100
, and as in
Fig. 1
).
Description.
FEMALE: Total length
4.6 mm
; forewing length
2.9 mm
.
Body black, with areas of yellow and brown. Yellow: (labrum not visible; anterior pronotal margin not visible); legs, except coxae. Brown: antenna; tegula. Wings hyaline; pterostigma brown.
Head about 0.9 × as long as wide in frontal view. Apical margin of clypeus projecting medially, medioapically with a shallow V-shaped incision (
Fig. 99
). Labrum not visible. Ventral mandibular tooth about ¼–1/5 of total mandibular length, not reaching opposite mandibular base. Palpal formula 4:3. Frons imbricate with dense and coarse prominent punctation and short setae (
Fig. 99
). Frontal carina present from median ocellus to upper half of clypeus, slightly elevated as a ridge on upper half of clypeus. Scape 4.5 × as long as wide. Ocellar triangle anterior of tangent between upper posterior orbits of compound eyes, lateral ocelli anterior of tangent by approximately their diameter. Occipital carina continuous (
Fig. 5
). Gena imbricate, with dense punctation and associated setae, without tubercles, spines, or dorsoventral carina. Hypostomal midventral line carinate with bordering short transverse carinulae on hypostomal integument, not angulate.
Mesosoma about 1.5 × as long as wide in dorsal view. Propodeum about 0.6 × as long as wide in dorsal view. Mesosoma imbricate, with dense punctation and associated setae (
Fig. 100
), except on dorsal surface of propodeum; lateral surface of propodeum also coarsely reticulate; dorsal surface of propodeum coarsely reticulate (similar to pattern in
Fig. 18
, but on much smaller surface). Slightly pitted sulcus present posterior to fine mesoscuto-mesoscutellar sulcus. Hypersternal sulcus long and pitted. Metafemur 2.8 × as long as wide. Metatibia without differently colored area apically. Pretarsal claws without teeth. Forewing with two submarginal cells; anterior border of submarginal cell II much shorter than posterior border.
Hind
wing with five distal hamuli.
Metasoma polished, with sparse punctation (
Fig. 100
). Metasomal sternum II without bulge (as in
Fig. 1
). Pygidium with narrow row of silk setae apically.
MALE: Difference from female aside from genitalic structures: clypeus yellow on apical half, edges of apical margin of clypeus blunt; mandible and antenna yellow. Metasomal sternum VIII narrow; forming upwardly curved ‘pseudo sting’ [form appears similar to ovipositor of female].
Remarks.
Arpactophilus tayo
is similar to
A. caac
and
A. pwaamei
as all have a similar body form and coloration. They can be distinguished by the basally dark brown coxae in
A. tayo
which is yellow in
A. caac
and
A. pwaamei
. The surface sculpture of the frons is similar in
A. tayo
,
A. pwaamei
, and
A. arhoe
but in
A. arho
the metasomal sternum II is flat without a bulge (as in
Fig. 1
) and in
A. pwaamei
the dorsal surface of the propodeum is longer and partially transversally carinate in addition to the aforementioned coxal coloration difference.
Material examined.
HOLOTYPE
♀: “Nouvelle Calédonie, Mont Koghis.
350m
,
25.IV.1995
, Réc. Chazeau & Jourdan” [
MNHN
].
PARATYPES
2♂
: “Nouvelle Calédonie, Pinda
30m
,
18.IV.1995
, Réc. Chazeau & Jourdan [
CAS
,
MNHN
].
Etymology.
The specific epithet is taken from the name
Tayo
, one of the New Caledonian native languages. It is treated as a noun in apposition.