A review of the New Caledonian Arpactophilus (Hymenoptera: Crabronidae)
Author
Breitkreuz, Laura C. V.
Author
Ohl, Michael
Author
Engel, Michael S.
text
Zootaxa
2016
4063
1
1
66
journal article
37206
10.11646/zootaxa.4063.1.1
9608b695-6310-4c63-bd94-46bda058e6d8
1175-5326
270412
80401ED8-C6BA-4420-9109-854C5CC1E88D
Arpactophilus arhoe
sp. nov.
(
Figs. 29–30
)
Diagnosis.
Arpactophilus arhoe
is the only New Caledonian species in the genus with the combination of a black clypeus (
Fig. 29
), a continuous occipital carina (
Fig. 5
), an imbricate and punctured gena without a carina, a black metasoma, and a bulge on sternum II (
Fig. 30
and as in
Fig. 2
).
Description.
FEMALE: Total length
5.4 mm
; forewing length
3.4 mm
.
Body black, with areas of dark yellow and brown. Dark yellow: labrum; palpi; scape; pedicel; flagellum in front; anterior pronotal margin; tegula; coxae apically; trochanters; most of femora; tibiae; tarsi. Brown: mandible partially; posterior surface of of flagellum; pronotal lobe apically; most of coxae; femora partially (
Fig. 30
). Wings hyaline; pterostigma dark brown.
Head about 0.9 × as long as wide in frontal view. Apical margin of clypeus medially projecting, with shallow, broad medial incision (
Fig. 29
.). Apical margin of labrum with two small lobes. Ventral mandibular tooth about ¼– 1/5 of total mandibular length, not reaching opposite mandibular base. Palpal formula 5:4. Frons imbricate, with dense, coarse punctation, punctures separated by less than a puncture width and dense setae (
Fig. 29
). Frontal carina present from median ocellus to almost apical margin of clypeus, elevated to a distinct ridge on clypeus. Scape 3.9 × as long as wide. Ocellar triangle anterior of tangent between upper posterior orbits of compound eyes, lateral ocelli anterior of tangent by less than their diameter. Occipital carina continuous (
Fig. 5
). Gena imbricate, with dense, coarse punctation and dense setae, without tubercles, spines, or dorsoventral carina. Hypostomal midventral line carinate, with bordering short transverse carinulae on hypostomal integument, not angulate.
Mesosoma about 1.7 × as long as wide in dorsal view. Propodeum about 0.8 × as long as wide in dorsal view. Mesosoma imbricate with additional dense punctation (
Fig. 30
), except dorsal surface of propodeum; lateral surface of propodeum also transversely carinulate; dorsal surface of propodeum coarsely reticulate (similar to pattern in
Fig. 18
). Pitted sulcus present posterior of fine mesoscuto-scutellar sulcus. Hypersternal sulcus long and pitted. Metafemur 2.6 × as long as wide. Metatibia without differently colored area apically. Pretarsal claws without teeth. Forewing with two submarginal cells; anterior border of submarginal cell II slightly shorter than posterior border.
Hind
wing with two distal hamuli.
Metasoma polished, with sparse punctation (
Fig. 30
). Metasomal sternum II swollen medially, forming bulge (as in
Fig. 2
). Pygidium with broad row of silk setae apically.
MALE: Unknown
Remarks.
Arpactophilus arhoe
can be distinguished from all other
Arpactophilus
with a bulge on the metasomal sternum II by the form of its clypeus. In
A. arhoe
, the apical margin of the clypeus is medially projecting, with a shallow, broad medial concave incision. Among species with a bulging metasomal sternum II, the form of the clypeus is closest to that of
A. arha
,
but the two species differ noticeably in coloration, especially of the clypeus and lower frons, which are black in
A. arhoe
(
Fig. 29
) and yellow in
A. arha
(
Fig. 27
).
Arpactophilus pwaamei
and
A. tayo
are also similar in coloration and clypeal form, but they have no bulge on metasomal sternum II.
Material examined.
HOLOTYPE
♀: “Nouvelle Calédonie, Pinda
30m
,
18.IV.1995
, Réc. Chazeau & Jourdan [
MNHN
].
Etymology.
The specific epithet is taken from the name
Arhö
, one of the New Caledonian native languages. It is treated as a noun in apposition.