A review of the New Caledonian Arpactophilus (Hymenoptera: Crabronidae)
Author
Breitkreuz, Laura C. V.
Author
Ohl, Michael
Author
Engel, Michael S.
text
Zootaxa
2016
4063
1
1
66
journal article
37206
10.11646/zootaxa.4063.1.1
9608b695-6310-4c63-bd94-46bda058e6d8
1175-5326
270412
80401ED8-C6BA-4420-9109-854C5CC1E88D
Arpactophilus haveke
sp. nov.
(
Figs. 59–60
)
Diagnosis.
Arpactophilus haveke
is the only New Caledonian species in the genus that has a stalked triangular submarginal cell II in the forewing, and particularly when taken in combination with the light reddish-brown metasoma further distinguishes itself from its congeners. Another distinctive feature is the apical margin of thee clypeus which is projecting medially and medioapically forming a point, without a medial incision (
Fig. 59
).
Description.
FEMALE: Total length
3.8 mm
; forewing length
2.3 mm
.
Body black, with areas of light yellow and light reddish-brown. Light yellow: apical margin of clypeus (
Fig. 59
); mandible; palpi; scape in front; pronotal lobe; tegula; coxae apically; trochanters; femora partially; tibiae, except apical area on metatibia; tarsi. Light reddish-brown: posterior of scape; pedicel; flagellum; most of coxae; most of femora; apical area on metatibia; metasoma (
Fig. 60
). Wings hyaline; pterostigma light brown. Labrum and anterior pronotal margin not visible.
Head about as long as wide in frontal view. Apical margin of clypeus projecting medially, medioapically forming a point (not incised or with a straight margin as in some other species) (
Fig. 59
). Labrum not visible. Ventral mandibular tooth about ¼–1/5 of total mandibular length, not reaching opposite mandibular base. Palpal formula 4:3. Frons imbricate with sparse punctation and short setae. Frontal carina present from median ocellus to apical margin of clypeus. Scape 3.7 × as long as wide. Ocellar triangle anterior of tangent between upper posterior orbits of compound eyes, lateral ocelli anterior of tangent by approximately their diameter. Occipital carina interrupted dorsally (
Fig. 6
). Gena imbricate with sparse punctation and sparse setae, without tubercles, spines, or dorsoventral carina. Hypostomal midventral line faint, not angulate.
Mesosoma about 1.7 × as long as wide in dorsal view. Propodeum about 0.8 × as long as wide in dorsal view. Mesosoma imbricate except on propodeum (
Fig. 60
); lateral and dorsal surface of propodeum transversely carinulate; dorsal surface of propodeum also finely reticulate with transverse carinulae apically (similar to pattern in
Fig. 20
). No groove present posterior to mesoscuto-mesoscutellar sulcus. Hypersternal sulcus absent. Metafemur 2.6 × as long as wide. Metatibia with a light brown area apically. Forewing with two submarginal cells; submarginal cell II triangular, not reaching media but connected to it by a single crossvein (i.e., cell is stalked).
Hind
wing with four distal hamuli.
Metasoma polished, punctation and associated setae sparse (
Fig. 60
). Metasomal sternum II without bulge (as in
Fig. 1
). Pygidium with narrow row of silk setae.
MALE: Difference from female aside from genitalic structures: yellow marking on clypeus larger; yellow markings present on lower frons and malar space; flagellum yellow; metasoma brown. Metasomal sternum VIII neither broadened nor narrowed.
Remarks.
Arpactophilus haveke
can easily be distinguished from the other species with a reddish metasoma by its stalked second submarginal cell. The only species that has a similarly triangular (but not stalked) second submarginal cell is
A. tiri
, which has a much more slender body form (
Fig. 102
).
Material examined.
HOLOTYPE
♀: “Nouvelle Calédonie, Pinda
30m
,
18.IV.1995
, Réc. Chazeau & Jourdan [
MNHN
].
PARATYPE
1♂
: “Nouvelle Calédonie, Pinda
30m
,
18.IV.1995
, Réc. Chazeau & Jourdan [
MNHN
].
Etymology.
The specific epithet is taken from the name
Haveke
, one of the New Caledonian native languages. It is treated as a noun in apposition.