An overview of the extant genera and subgenera of the order Scolopendromorpha (Chilopoda): a new identification key and updated diagnoses Author Schileyko, Arkady A. schileyko1965@gmail.com Author Vahtera, Varpu varpu.vahtera@gmail.com Author Edgecombe, Gregory D. 0000-0002-9591-8011 schileyko1965@gmail.com text Zootaxa 2020 2020-08-10 4825 1 1 64 journal article 8703 10.11646/zootaxa.4825.1.1 5ab5f5c8-481e-4d1a-8643-21e72c367278 1175-5326 4402145 F230F199-1C94-4E2E-9CE4-5F56212C015F Family Plutoniumidae Bollman, 1893 Synonyms. Theatopidae Verhoeff, 1906; Theatopinae Verhoeff, 1906 ; Theatopsidae Verhoeff, 1907. Diagnosis. Eyes absent, light/depigmented spots at their place ( Fig. 34 ). Labrum with a single median tooth. Slender pretarsus of maxillae 2 ( Fig. 35 ) not pectinate, consisting of two well-distinguishable parts (darker basal and a semi-transparent apical ones), curved and pointed apically. Pretarsus (at least in Theatops ) is accompanied by welldeveloped ventral projection ( Fig. 35 ) which is of the same shape as pretarsus but visibly thinner and shorter (sometimes nearly as long as it; see below); it is never accompanied by accessory spine(s). Anterior margin of forcipular coxosternite with well-developed tooth-plates ( Fig. 36 ), forcipular trochantero-prefemur with simple processes of various length. Sternites with a single longitudinal median suture, developed to varying degrees (hardly visible in some Theatops ). 21 LBS; spiracles (figs 1AB in Vahtera et al . 2012b ) on LBS 2–20 or on macrosegments only (if so LBS 7 with or without spiracles). Ultimate LBS considerably elongated with ultimate tergite nearly twice as long the penultimate one. Coxopleuron virtually without process ( Fig. 37 ), sometimes with a single spine at its place. Tarsus of legs 1–19 monopartite; legs with two tibial spurs and one tarsal spur. Ultimate legs forcipulate ( Figs 33, 37 ), swollen, strongly sclerotized, truly “pincer-shaped” (sensu Schileyko 2009 ) with all articles much shortened and enlarged, excluding falcate pretarsus which is considerably elongated being at least as long as corresponding tarsus 2 (or much longer; see below). Dorsal and medial surface of both prefemur and femur flat, these articles with or without ( Fig. 37 ) ventral spines. Edgecombe & Bonato (2011: 395) also wrote: “Poison calyx extending into the forcipular coxosternite… Gizzard with stiff anteriorly/directed projections; projections evenly curved, covered by multifurcating scales that spirally encircle the projection, branching into slender, needle-like spines”. Number of subtaxa. 2 genera. Sexual dimorphism. Unknown. Range. Southern Europe ( Southern Iberian Peninsula , Southern Italian Peninsula , Balkan Peninsula, Sardinia, Sicily); South-West, East and South-East of USA , Northern Mexico ; Central China ( Sichuan , Hunan , Gansu ) . Remarks. Treated as a family in Edgecombe & Bonato (2011: 395) , Edgecombe et al. (2012: 770) , Vahtera et al. (2012a: 9 , 2012b: 229 , 2013: 580 ), Bonato et al. (2017: 2) .