Review of the Brazilian species of Omorgus Erichson, 1847 (Coleoptera: Trogidae: Omorginae) Author Costa-Silva, Vinícius Author Strümpher, Werner P. Author Vaz-de-Mello, Fernando Z. text Journal of Natural History 2021 2021-02-16 54 31 - 32 1993 2024 http://dx.doi.org/10.1080/00222933.2020.1833999 journal article 10.1080/00222933.2020.1833999 1464-5262 5028835 Omorgus ( Omorgus ) suberosus ( Fabricius, 1775 ) ( Figures 6 a-d and 11) Trox suberosus Fabricius,1775: 31 (original description); Herbst 1790: 29 (description); Illiger 1802: 332 (catalogue); Castelnau 1840:107 (as ‘ tuberosus ’ – misspelling); Blanchard 1847: 190 (catalogue), Harold 1872: 119 (redescription); Leng 1920: 253 (catalogue); Mutchler 1925: 238 (catalogue of Galapagos Islands); Leng 1928: 422 (catalogue of New York); Denier 1936: 205 (natural history); Hayward 1936: 217 (feeding habit); Blackwelder 1944: 219 (catalogue – as ‘ suberosa ’); Van Dyke 1953: 123 (distribution data); Haaf 1954: 739 (catalogue of Australia ); Vaurie 1955: 60 (redescription); Richter, 1958: 325 (biology); Vaurie 1962: 144 (redescription); Ritcher 1966: 73 (biology); Hatch 1971: 464 (key to the Pacific Northwest beetles); Chalumeau and Gruner 1974: 787 (catalogue of French Antilles); Zimsen 1964: 38 (catalogue) Trox ( Omorgus ) suberosus : Burmeister 1876: 257 (redescription); Arrow 1903: 516 (notes about fauna of the St. Vincent Island ); Arrow 1912: 62 (catalogue); Scholtz 1982: 13 (systematics). Omorgus suberosus : Baker 1968: 42 (description of larvae); Scholtz 1986: 361 (phylogenetics); Scholtz 1990: 1407 (redescription); Zídek 2013: 17 (catalogue); Zídek 2017: 109 (catalogue); Huchet and Costa-Silva 2018: 565 (new distribution records from South America); Hielkema and Hielkema 2019 (checklist of the Guianas). For synonyms see Smith (2017) and Zídek (2017) . Type material Type series could not be traced. Fabricius (1775) stated the specimens he studied for the description of the species came from the Joseph Banks Collection, which is currently housed in the BMNH ( Zimsen 1964 ). Nevertheless, according to Vaurie (1962) , due to inconsistencies with the original description the specimen currently resting under the ‘ suberosus ’ label in the Banks collection certainly does not correspond to the species named by Fabricius (1775) as Trox suberosus . The whereabouts of the actual type series was unknown to Vaurie (1962) and all subsequent authors, including us. Even so, we do not consider it necessary to designate a neotype since Omorgus suberosus is a welldefined species and can be readily recognised from the literature ( e.g . Vaurie, 1962 ; Scholtz 1990; Huchet and Costa-Silva 2018 ). Type locality: BRAZIL , Rio de Janeiro : off the coast of Rio de Janeiro city, Ilha Rasa (originally cited as ‘Brasilia’; see Papavero (1971) for more details). Material examined (additionaly to Huchet and Costa-Silva 2018 ) USA: Texas , ‘Furesthing’ (sic), vii .1950, no collector (1 DZUP ). MEXICO : Chihuahua , no specific locality, 15 .viii .2001, Juan Jesus leg. (1 CVMD ). PUERTO RICO : no specific locality or date, Julio Micheli leg. (1 CVMD ). CUBA : no further data (1 MACN ) . NICARAGUA : León , no specific locality, v .1993, Maes and Tellez leg. (1 CVMD ); no specific locality, v.1990 , B . Garcete leg . (1 CVMD ). Masaya , Las Flores, vi.1993 , C . Levoq and I . Cautanessa leg. (1 CVMD ). ECUADOR : Esmeraldas , San Lorenzo, 18–22 .x .2004, V .M . Diéguez leg. (1 CVMD ). Guayas , El Aromo , 1 .xi .2010, W . Chamorro leg . (1 CVMD ). PERU : Loreto , Maynas, Iquitos , 14 .xii .1988, Pedro Hocking leg. (1 CVMD ). Amazonas , Bagua , Copallín , 748 m , 78°25’S 5° 41 ʹ W , 21 .i .2012, Medina and Razuri (1 CVMD ) . Tumbes , Tumbes , Los Cedros , 24 .iii .2009, Castillo Carillo P . leg . (2 CVMD ). Piura , Road (‘ Corretera’ , sic) from Piura to Paita , 10 .ii .2012, on vulture carcase, D . Saavedra leg . (1 CVMD ); Sullana , 2 .v .2006, D . Saavedra leg . ( 1 ♂ CVMD ) . Lima , Canta , ii .2002, Cristian Mendoza leg . (1 CMVD ); Lima , 18 .ii .1999. M. Beeche leg . ( 1 ♂ , 1 CVMD ); Lima , Cieneguillas , 500 m , 9–10 .xi .2002, V .M . Diéguez leg . (1 CVMD ); Ate , Vitarte , 2 .iv .1999, fluorescent light trap, G . Valencia leg . (2 CVMD ). Ica , Reserva Nacional de Paracas , Bahía de Paracas , 13–15 .iii .2003, V .M . Diéguez leg. (1 CVMD ). BOLIVIA : no locality or date, A . Beyer leg . (1 MACN ). BRAZIL : Roraima , Pacaraima, Surumu , x .1955, P .J . Silva leg. (1 DZUP ). Pará , Jacareacanga , x .1959, M . Alvarenga leg . (4 CEMT ); Óbidos , iii .1954, J . Brazillino leg . (1 DZUP ). Rio Grande do Norte , Natal , iii .1950, M . Alvarenga leg . (1 DZUP ); Natal , ii .1951, M . Alvarenga leg . (1 DZUP ); Natal , iii .1952, M . Alvarenga leg . (1 DZUP ). Paraíba , Campina Grande , Catolé , Paus Brancos [‘Catolé de B. Vista / Paus Brancos’ ], 1 .vii .2000, M . Avany leg . (3 CEMT ); Campina Grande , Catolé , Paus Brancos [‘Catolé de B. Vista /Paus Brancos’], 5–6 .iii .2000, light trap, M . Avany leg . (3 CEMT ); Campina Grande , Catolé , Paus Brancos [‘Catolé de B. Vista /Paus Brancos’], 3 .v .2000, light trap, M . Avany leg . (1 CEMT ); Congo , 19 .i .2010, dry pig carcase, no collector (1 CEMT ); João Pessoa , Jardim Botânico Benjamin Maranhão [‘Mata do Buraquinho’], 7°8’S 34°51 ʹ W , 17 . iii .2009, R .C .A.P. Farias leg . (1 CEMT ); João Pessoa , Jardim Botânico Benjamin Maranhão [‘Mata do Buraquinho’], 7°8’S 34°51 ʹ W , 21 .iii .2010, R .C .A.P. Farias leg . (1 CEMT ); João Pessoa , Jardim Botânico Benjamin Maranhão [‘Mata do Buraquinho’], 7°8’S 34°51 ʹ W , 6 .ix .2009, R .C . A.P. Farias leg . (1 CEMT ); João Pessoa , Jardim Botânico Benjamin Maranhão [‘Mata do Buraquinho’], 7°8’S 34°51 ʹ W , 7 .ix .2009, R .C .A.P. Farias leg . (4 CEMT ); Mataraca , Parque Eólico Millennium , 15 .i .2005, Bruno and Valdez leg . (1 CEMT ); Rio Tinto , Área de ProteÇão Ambiental da Barra do Rio Mamanguape [APA do Mamanguape], 6°46’S 34°55 ʹ W , 26 . iii .2012, D . Dal-bó leg . (2 CEMT ); Rio Tinto , APA do Mamanguape , 6°46’S 34°55 ʹ W , 20 . iii .2012, D . Dal-bó leg . (3 CEMT ); Rio Tinto , APA do Mamanguape , 6°46’S 34°55 ʹ W , 20 . xi .2012, D . Dal-bó leg . (1 CEMT ); Rio Tinto , APA do Mamanguape , 6°46’S 34°55 ʹ W , 25 . iii .2012, D . Dal-bó leg . (2 CEMT ); Rio Tinto , APA do Mamanguape , 6°46’S 34°55 ʹ W , 23 . iii .2012, D . Dal-bó leg . (1 CEMT ); Rio Tinto , APA do Mamanguape , 6°46’S 34°55 ʹ W , 22 . iii .2012, D . Dal-bó leg . (1 CEMT ); Rio Tinto , APA do Mamanguape , 6°46’S 34°55 ʹ W , 12 . iii .2012, D . Dal-bó leg . (2 CEMT ); São José dos Cordeiros , Fazenda Almas , 600 m , 7°28’S 36° 53 ʹ W , 9 .ii .2011, W .E . Santos leg . (1 CEMT ); São José dos Cordeiros , Fazenda Almas , 600 m , 7° 28’S 36°53 ʹ W , 8 .ii .2011, W .E . Santos leg . (2 CEMT ); São José dos Cordeiros , Fazenda Almas , 600 m , 7°28’S 36°53 ʹ W , 18 .ii .2011, W .E . Santos leg . (1 CEMT ); São José dos Cordeiros , Fazenda Almas , 600 m , 7°28’S 36°53 ʹ W , 11 .ii .2011, W .E . Santos leg . (1 CEMT ), São José dos Cordeiros , Fazenda Almas , 600 m , 7°28’S 36°53 ʹ W , 3–4 .ii .2011, Creão & Rothea leg . (4 CEMT ); São José dos Cordeiros , Fazenda Almas , 600 m , 7°28’S 36°53 ʹ W , 8–9 .ii .2011, R . Rothea leg . (1 CEMT ). Bahia , Jequié , 250 m , 24 .ii .1995, C . Sperber leg . (3 CEMT ); Salvador , i .1944 (1 DZUP ). Tocantins , Porto Nacional , 10°31’S 48°17 ʹ W , 12 .x .2015, light trap, A . Specht leg . (1 CEMT ); Porto Nacional , 10°31 ʹ 8”S 48°17 ʹ W , 9 .x .2015, light trap, A . Specht leg . (1 CEMT ); Porto Nacional , 10°31’S 48°17 ʹ W , 8 .x .2015, light trap, A . Specht leg . (1 CEMT ); Santa Maria do Tocantins , Casa de Pedra Cave , 30 .viii .2005, F .J . Silva leg. (3 CEMT ). Goias , AragarÇas , x .1959, M . Alvarenga leg . (1 DZUP ). Mato Grosso , Chapada dos Guimarães , 23 .xi .1983, no collector (1 DZUP ); CotriguaÇu , Fazenda São Nicolau , 20 .viii .2018, R . Stofel leg . (1 CEMT ); Jaciara , 29–30 .xii .2017, S .M . Jacobina leg. (1 CEMT ). Mato Grosso do Sul , Camapuã , x .1967 (16 MGAP ); Chapadão do Sul , 18°46’S 52°31 ʹ W , 9 .x .2015, light trap, A . Specht leg . (1 CEMT ); Rio Verde de Mato Grosso , 400 m , xi .1959 (1 DZUP ); Rio Verde de Mato Grosso , i .1967, A . Maller leg . (1 DZUP ). Espírito Santo , Baixo Guandú, 22–31 .iii .1971 , Tadeu & C . Elias leg . (2 DZUP ); Baixo Guandú , 2–9 .i .1971, Tadeu & C . Elias leg . (1 DZUP ). Minas Gerais , Ipatinga, 11 .xii .1993 (2 CVMD ); Ipatinga , iii.1985 , E . Grossi leg . (4 CVMD ); Itaobim , xii .2007, no collector (1 CVMD ); Passos , xi .1961, Claudionor Elias leg . (2 DZUP ); Sete Lagoas , 20 .x .1969, V .O . Becker leg . (2 DZUP ); Uberlândia , 18 .x .1970, H .N . Espínola leg . (1 DZUP ); Varginha , i .1960, Seabra & Alvarenga leg . (1 DZUP ); Vespasiano , i .1952, A . Machado leg . (2 CEMT ). Rio de Janeiro , Rio de Janeiro , Ilha do Governador , Galeão , i .1943, M . Alvarenga leg . (1 DZUP ); Rio de Janeiro , Ilha do Governador , Galeão , ix .1956, M . Alvarenga leg . (1 DZUP ); Rio de Janeiro , Ilha do Governador , Galeão , x .1956, M . Alvarenga leg . (1 DZUP ); Rio de Janeiro , Ilha do Governador , Galeão , xi .1955, M . Alvarenga leg . (2 DZUP ); Rio de Janeiro , Ilha do Governador , Galeão , xii .1955, M . Alvarenga leg . (1 DZUP ). São Paulo , Ribeirão Preto , 31 .x .1973, Padre Moure leg . (2 DZUP ); São Carlos , 900 m , x .1981, G . Serocchi leg . (2 IFML ); São Carlos , 13 .x .1982, Marinoni and Hoffmann leg. (2 DZUP ). Paraná , Curitiba , 15 .xii .1974, R . Langue leg . (1 DZUP ); Curitiba , 5 .xi .1992, Rosado N . leg . (1 DZUP ); Curitiba , 900 m , 11 .ix .1984, no collector (1 DZUP ); Curitiba , 900 m , 9 .xi .1968, Departamento de Zoologia leg . (1 DZUP ); Francisco Beltrão , 17 .x .1981, R . Dias leg . (1 CVMD ); Francisco Beltrão , 20 .ii .1981, R . Dias leg . (1 CVMD ); Francisco Beltrão , 3 .x .1980, R . Ojeda leg . (2 CVMD ); Jundiaí do Sul , Fazenda Monte Verde , 27 .iii .1987 (1 DZUP ); Londrina , 14 .ii .1975, Pe . Moure leg . (1 DZUP ); Maringá , 8 .xi .1968, J . Parra Muro leg . (1 DZUP ); Maringá , xi .1969, J . Parra Muro leg . (2 DZUP ); Marumbi , xii .1973, Dr . Curial leg . (1 DZUP ); Morretes , 23–24 .xi .1984, light trap, C.I.I.F leg. (1 DZUP ); Piraquara , Banhado Railway Station , xii .1972, P .J . Riehs leg . (2 DZUP ); Ponta Grossa , xii .1943, ‘on the wall’ [possibly referring to one of the famous sandstone formations of Vila Velha State Park] (1 DZUP ); Porecatu , 20 .x .1970, Becker and Hatschback leg. (1 DZUP ); no further data (1 MLPA ). Santa Catarina , Corupá , 60 m , xi .1957, no collector (1 DZUP ); Florianópolis , 23 .xii .1994, P .C . Garcia leg . (1 MCNZ ); Governador Celso Ramos , 10–27 . ii .1993, L . Moura leg . (1 MCNZ ); Itapema , 67 m , i .1993, J . Carlos leg . (1 CEMT ); Itapiranga , i .1954, no collector (1 MGAP ); Itapiranga , ii .1934, no collector (1 MGAP ); JoaÇaba , 2 .i .1967, R . Antoni leg . (1 DZUP ); Pinhalzinho , iv .1951, A . Maller leg . (1 DZUP ). Seara , Nova Teutônia , 300 ‒ 500 m , 27°23’S 51°12 ʹ W , ii .1979, Fritz Plaumann leg . (4 DZUP ); Seara , Nova Teutônia , 300 ‒ 500 m , 27°23’S 51°12 ʹ W , ii .1981 (1 DZUP ); Seara , Nova Teutônia , 300 ‒ 500 m , 27°23’S 51° 12 ʹ W , xi .1977 (3 DZUP ); Seara , Nova Teutônia , 300 ‒ 500 m , 27°23’S 51°12 ʹ W , xii .1975 (1 DZUP ); Seara , Nova Teutônia , no further data (1 MLPA ); Tubarão , 13 .vii .2018, R.B. Goulart (1 CEMT ) . Rio Grande do Sul , CanguÇu , Coxilha do Fogo , i .1996, C .N . Duckett leg . (1 MCNZ ); Cerro Largo , 1949, no collector (1 MGAP ); Cerro Largo , i .1930, no collector (1 MGAP ); Cerro Largo , i .1932, no collector (1 MGAP ); Estrela Velha , Itaúba , 7 .iv .1978, H . Bischoff leg . (2 MCNZ ). Guaíba , 1976, M .H . Galileo leg . (1 MCNZ ); Maquiné , 7–8 .iv .1995, L . Moura leg . (1 MCNZ ); Parecei Novo , 15 .ix .1964, no collector (1 MGAP ); Pareci Novo , 17 .x .1964, no collector (1 MGAP ); Pareci Novo , iv .1933, no collector (1 MGAP ); Porto Alegre , 17 . viii .1927, no collector (1 MGAP ); Porto Alegre , 2 .iv .1932, no collector (2 MGAP ); Porto Alegre , 24 .v .1927, no collector (2 MGAP ); Porto Alegre , 25 .ix .1927, no collector (1 MGAP ); Porto Alegre , 27 .ix .1928, no collector (1 MGAP ); Porto Alegre , 29 .iv .1928, no collector (1 MGAP ); Porto Alegre , 30 .x .1952, no collector (1 MCNZ ); Porto Alegre , 5 .iii .1965, L . Buckup leg . (4 MCNZ ). Porto Alegre , 8 .xi .1927, no collector (2 MGAP ); Porto Alegre , 9 .ix .1927, no collector (1 MGAP ); Porto Alegre , Bairro Vila ConceiÇão , xi .1952, L. Buckup (1 MCNZ ); Porto Alegre , Ipanema , xi .1956, Palova leg . (1 MCNZ ); Porto Alegre , Morro Santana , 28 .xii .1982, light trap , no collector (1 MCNZ ); Porto Alegre , x .1956, L . and E . Buckup leg . (1 MCNZ ); Porto Alegre , xi .1951, L . Buckup leg . (1 MCNZ ). Salto do Jacuí , garden of Companhia Estadual de Energia Elétrica , 25–29 .x .1999, light trap , no collector (1 MCNZ ); Santa Maria , 1 .iii .1974, light trap, D . Link leg . (1 DZUP ); Santa Maria , 6 .iv .1996, D . Link leg . (1 MCNZ ); São Jerônimo , Instituto Rio-Grandense do Arroz [‘IRGA’: Rio-Grande Rice Institute], 11 .i .1989, A . Ferreira leg . (2 MCNZ ); Sapucaia do Sul , 1964, no collector (2 MGAP ); Terra de Areia , São Salvador Beach , 1963, no collector (2 MGAP ) . Ambiguous state: ‘ Matto Grosso’ , no further data (5 MLPA ); ‘ Matto Grosso’ , no date, Koslomshy leg . (4 MLPA ). PARAGUAY : no further data (1 MLPA ; 1 MACN ) . Amambay , Parque Nacional Cerro Corá [Cerro Corá National Park], 280 m , 22°38’S 56°01 ʹ W , 1–6 .ii .2006, Roberto Barrera Medina leg. (9 CVMD ). Concepción , Azotey , x .2001, C . Aguiler leg . (1 CVMD ). San Pedro , Cororo, 20 .xii .1999 , Jorge Jensen leg. (2 CVMD ). Cordillera , Piribebuy , Cerro Naranjo , 27 .ix .1999, C . Aguilar leg . (1 CVMD ). Guayrá , Independencia , 25 km E, 28 .i .1991, pitfall baited with meat, Endödy-Younga leg . (1 TMSA ); Independencia , 25 km E, 30 .i .1991, on dog carcase, Endödy-Younga leg. (8 TMSA ). Boquerón , Loma Plata , 20–30 .i .1994, P . Gerlach leg . (5 CVMD ); Teniente Enciso , 4 .xi .2001, Carlos A . Julio leg . ( 1 ♂ CVMD ) . Central , Potrerito , vi .1988, Carlos A . Julio leg . (1 CVMD ). Distrito Capital , Asunsión, 21 .x .1983 (1 CVMD ); Asunsión , Bahía de Asunción , 5 . vi .1988 , Carlos A . Aguilar leg . (1 CVMD ). ARGENTINA : no further data (2 CVMD ; 2 MLPA ) . Santiago del Estero , Río Salado , Wagner leg . (2 MLPA ); Moreno , Girardet , no further data (22 MLPA ); no precise locality, x .1939, no collector (1 DZUP ; 6 MLPA ). Buenos Aires , La Plata , no date, A .R . Bezzi leg. (5 MLPA ); no specific locality, ii.1899 , C . Bruch leg . (2 MACN ); no further data, C . Bruch leg . (2 MACN ); no specific locality or date, A . Breyer leg . (11 MACN ); Caseros , 15 .i .1946, no collector (4 MACN ); Río Colorado , no further data (1 MACN ; 1 MLPA );, no specific locality, 25 .xi .1919, J.B. leg. (1 MACN ); no specific locality, 27 .vi .1903 , A . Zotta leg . (1 MACN ); no specific locality or date, A . Zotta leg . (1 MACN ); no specific locality or date, A . Pellerano leg . ( MACN ); Miramar , i .1977, S . Roig leg . (1 IADIZA ); no specific locality, 9 .ii .1914 , J.B. leg. (1 MACN ); no specific locality, 20 .xi .1922 , J.B. leg. (1 MACN ); no specific locality, 2 .i .1912 , J.B. leg. (1 MACN ); La Plata , no date, A .R . Bezzi leg . (1 MLPA ); Tres Arroyos , 30 .i .1938, Biraben-Scott leg . (1 MLPA ); San Miguel , Las Delicias , 22 . iii .1925, J .R . Baer leg. (1 MACN ); no specific locality or date, J . Boso leg . (1 MACN ); no specific locality or date, A . Zotta leg . (2 MACN ); La Plata , no further data (2 MLPA ); Isla Martín García , iii .1935, Castillo leg . (1 MACN ); Tigre , Tres Bocas , El Tropezón Hotel , Paraná River Delta , 25–30 .xii .1933, Hector Gavio leg. (1 MACN ); no specific locality or date, G . Pellerano leg . (2 MACN ); Bahía Blanca , no further data (1 MACN ); Tandil , iv .1952, no collector (1 MACN ); Lanús , xi .2002, J .E . Barriga leg. (1 CVMD ); no specific locality or date, H . Richter leg . (2 MLPA ); no specific locality or date, J .E . Barriga leg. (1 CVMD ; 11 MLPA ; 5 MACN ). CIUDAD AUTÓNOMA DE BUENOS AIRES : no further data (3 MACN ); no more precise locality, iii .1941, Arenema leg. (1 DZUP ). Córdoba , Río Segundo , Oncativo , 17 . i .1939, Biraben-Scott leg . (5 MLPA ); Calamuchita , ‘ El Sauce’ , xii .1938, M . Viana leg . (3 DZUP ); Santa María , Alta Gracia , ‘ La Granja’ , xii .1926, C . Bruch leg . (1 MACN ); no specific locality or date, C . Bruch leg . (1 MACN ); no specific locality or date, A . Breyer leg . (1 MACN ); Colón , Jesús María , 4 .iv .1940, Maldonado leg . (1 MLPA ); Cruz de Eje , 21–25 .ii .2000 (1 IADIZA ); Río Cuarto , Río Cuarto , Universidad Nacional de Río Cuarto, IV , 17–18 .xi .2005, E . Ruiz and S . Ferrer leg . (2 IADIZA ); Santa María , Alta Gracia , i .1909, R . Vigel leg . (1 MACN ); Calamuchita , Yacanto de Calamuchita , no date, G . Pellerano leg . (2 MACN ); Cabana [?], 7 . i .1926, no collector (2 MLPA ); Cabana [?], 4 .ii .1926, no collector (1 MLPA ); Cabana [?], 22 . i .1926, no collector (1 MLPA ); Cabana [?], 26 .i .1926, no collector (1 MLPA ); no specific locality, xi.1938 , R . Cabellos leg . (4 MACN ); Río Cuarto , Río Cuarto , Universidad Nacional de Río Cuarto , 13 .xi .2001, on meat, no collector (1 CVMD ); Punilla , La Falda , 9 .ii .1963, M . Viana leg . (1 CVMD ); Calamuchita , Los Reartes , x .1970, M . Viana leg . (1 CVMD ); Calamuchita , ‘ El Sauce’ , x . 1975, M . Viana leg . (5 CVMD ); Calamuchita , ‘ El Sauce’ , xi .1976, M . Viana leg . (2 CVMD ); no further data (6 MLPA ; 3 MACN ). Corrientes , Santo Tomé, v.1926 , G . Pellerano leg . (12 MACN ); Santo Tomé , i .1928, G . Pellerano leg . (1 MACN ); Santo Tomé , viii .1923, G . Pellerano leg . (1 MACN ); Santo Tomé , xi .1925, G . Pellerano leg . (1 MACN ); Paso de los Libres , no date, G . Pellerano leg . (2 MACN ); no further data (1 MLPA ). Jujuy , no specific locality or date, C . Bruch leg . (1 MACN ); no further data (2 MACN ). Río Negro , Valcheta , Aguada Cecilio, no date, C . Bruch leg . (1 MACN ); no specific locality or date, A . Breyer leg . (1 MACN ). San Luis , Ayacucho , Quines , 18 .ii .1940, Biraben leg . (1 MLPA ); San Martín de Pueyrredón , San Luis , ii .1933 (1 MLPA ); San Martín de Pueyrredón , Suyuqye Nuevo , xii .1935 (1 MLPA ); Juan Martín de Pueyrredón , San Gerónimo , i .1972, M . Viana leg . (46 MLPA ; 1 MACN ); Belgrano , Fundo El Molle , 622 m , ii .2010, José Gomez leg . (1 CVMD ); Belgrano , Fundo El Molle , 622 m , 15–25 .i .2007, José Gomez leg. (1 CVMD ). Chaco , no specific locality, i.1898 , C . Bruch leg . (1 MACN ; 2 MLPA ; 8 MACN ). Santa Fe , Rosario (‘ Rosario de Santa Fe’ ), no date, G . Pellerano leg . (1 MACN ); Rosario , 18 .xii .1927, M .P . Gomez leg . (1 MACN ); Garay , Colonia Mascias , xi .1942, M . Viana leg . (2 MACN ); General Obligado , Guadalupe Norte , 27 .xi .1939, Biraben & Bezzi leg. (1 MLPA ); no specific locality, no date, C . Bruch leg . (2 MACN ). La Rioja , no specific locality or date, A . Breyer leg . (9 MACN ); Ruta Nacional 60 (‘ Ruta 60’), 5 .ii .2002, on dear horse over the road, C . Medina leg . (5 UPTC ); Chilecito , no further data (2 MLPA ); General Belgrano , 5 km W Olta , 3 .xii .2003, ‘bovine’ [dung or carcase?], no collector (1 IFML ); Salinas Grandes , 15 .xii .1993, G . Flores leg . (1 IADIZA ); no specific locality, 13 .ii .1923 , no collector (1 MACN ); no further data 9 ( MACN ). Misiones , no specific locality or date, A . Breyer leg . (7 MACN ); Candelaria , Bonpland , no further data (1 MLPA ); San Ignacio , no further data (1 MACN ); Puerto Iguazú , x .1927, G . Pellerano leg . (1 MACN ); no further data 1 ( MACN ). Tucumán , Leales , Tacanas , i .1949, M . Asnau leg . (17 IFML ); no specfic locality or date A . Breyer leg . (7 MACN ); 1 ( MLPA ); 4 ( MACN ). Salta , Orán , 24 .vi .1947, Biraben leg . (2 MLPA ); La Viña , iii .1983, M . Viana leg . (1 CVMD ); Anta , Las Lajitas , xii .1982, M . Viana leg . (1 CVMD ); Rosário de Lerma , xi .1986, M. Viana (1 CVMD ); Cafayate , 1 km N Cafayate , 26°3’S 65°56 ʹ W , 24 .xi .2006, mercury vapour and UV light trap, F . Ocampo, E . Ruiz, G . Sanblas and G . Zalazar leg. (2 IADIZA ). Formosa , no specific locality, 24 .xii .1953 , Denier leg . (4 MLPA ); Laishí , 12 .xi .1942, Biraben and Bezzi (1 MLPA ); Formosa , Estancia Guaycolec , 13–15 .i .2002, R . Gomez leg . (1 CVMD ); ‘ El Bagual’ , 8–9 .xii .2001, Exc. Hum . [ Human feaces] (1 IADIZA ); Bermejo , Laguna Yema , Reserva de Biosfera Riacho Teuquito , 24°21 ʹ 20”S 61°18 ʹ 55”W , 11 .xii .2008, F .C . Ocampo, G . Sanblas , and F . Campón leg . (26 IADIZA ); Pilagás , El Espinillo , 29 .i .1948, Duret leg. (1 CVMD ). Catamarca , no specific locality, 1917, no collector (1 MLPA ); no specific locality, 17 .ii .1908, C . Bruch leg . (1 MLPA ); Andalgalá , no further data (2 MLPA ); Andalgalá , El Pucará (‘ Campos de Pucará’ ), 9 .i .1951, no collector (1 IFML ); Ambato , El Rodeo , 12 .xii .2003, horse carcase, no collector (1 IFML ); El Alto , Guayamba , 1078 m , 28°20’S 65°24 ʹ W , 17–26 .i .2011, F .C . Ocampo, F.M . Ocampo , and P . Valentini leg . (1 IADIZA ); Santa María , 120 km N Belén , 2233 m , 26°59’S 66°18 ʹ W , 3 .ii .2009, on a bovine carcase, V .M . Diéguez and G . Arriagada leg . (2 CVMD ); Campo Posuelo , 60 km S Punta de Balasto , 8 .ii .1997, G . Arriagada leg . (4 CVMD ); Santa María , Punta de Balasto , 16 .i .1995, P . Gerlach leg . (1 CVMD ); no further data (44 MACN ; 11 MLPA ). San Juan , Valle Fértil , i .1972, M . Viana leg . (2 MACN ); Valle Fértil , xi .1970, M . Viana leg . (2 CVMD ). Entre Ríos , Islas del Ibicuy , Villa Paranacito , no further data (1 MACN ); Concordia , no date, Daguerre leg . (1 MACN ); Colón , Pueblo Liebig , xii .1997, Mateo Zelich leg . ( 1 ♂ , 12 CVMD ); Parque Nacional El Palmar , xi .1974, M . Viana leg . (1 CVMD ); Colón , Pueblo Liebig , ii .1996, Mateo Zelich leg . ( 1 ♂ , 7 CVMD ); Colón , Pueblo Liebig , xii .1986, Mateo Zelich leg. (1 CVMD ); no further data (4 MACN ). La Pampa , Lihuel Calel , Cuchillo-có , 1913, A . Copello leg . (1 MACN ); Chalileo , Santa Isabel , 9 .i .2007, no collectr (1 IADIZA ); Capital , Santa Rosa , 22 .iv .2002, Martínez leg . (1 IADIZA ); Capital , Santa Rosa , Facultad de Agronomía , 20 .i .1995, Bárbara Corró Molas leg . (3 IADIZA ); Capital , Santa Rosa , Facultad de Agronomía , 23 .viii .1996, Bárbara Corró Molas leg . (1 IADIZA ); Reserva Provincial Parque Luro , 10 .xii .1997, D . Rojas Lanus leg . (1 CVMD ). Neuquén , no further data (1 MLPA ) . Chubut , no further data (2 MLPA ) . Mendoza , Capital , Mendoza , Centro Regional de Investigaciones Cientificas y Tecnologicas [Fundación CRICYT], 850 m , 32°53’S 68°52 ʹ W , 7 .xi .2008, F . Aballay leg . (4 IADIZA ); Capital , Mendoza , Fundación CRICYT, 850 m , 32°53’S , 68°52 ʹ W , 5 .xi .2008, F . Aballay leg . (1 IADIZA ); Capital , Mendoza , Fundación CRICYT, 850 m , 32°53’S 68°52 ʹ W , 3 .xi .2008, F . Aballay leg . (1 IADIZA ); Capital , Mendoza , Fundación CRICYT, 850 m , 32°53’S , 68°52 ʹ W , 1 .i .2008, F . Aballay leg . (1 IADIZA ); Capital , Mendoza , Fundación CRICYT, 850 m , 32°53’S 68°52 ʹ W , 11 .xi .2008, F . Aballay leg . (4 IADIZA ); Capital , Mendoza , Fundación CRICYT, 850 m , 32°53’S , 68°52 ʹ W , 10 .xi .2008, F . Aballay leg . (2 IADIZA ); Capital , Mendoza , Fundación CRICYT, 850 m , 32°53’S 68°52 ʹ W , 13 .xi .2008, F . Aballay leg . (2 IADIZA ); Capital , Mendoza , Fundación CRICYT, 850 m , 31 .i .2008, F . Aballay leg . (1 IADIZA ); Tunuyán , Reserva Manzano Histórico , 1400 m , 33°36’S 69°19 ʹ W , 31 .i .2016, F . Aballay and F . Jofré leg . (3 IADIZA ); Tunuyán , Reserva Manzano Histórico , 1400 m , 33°36’S 69°19 ʹ W , 31 .xii .2016, F . Aballay and F . Jofré leg . (1 IADIZA ); Tunuyán , Reserva Manzano Histórico , 1400 m , 33° 36’S 69°19 ʹ W , 31 .xii .2015, F . Aballay and F . Jofré leg . (3 IADIZA ); Tunuyán , Reserva Manzano Histórico , 2200 m , 33°36’S 69°19 ʹ W , 31 .i .2016, F . Aballay and F . Jofré leg . (1 IADIZA ); Tunuyán , Reserva Manzano Histórico , 1400 m , 33°36’S 69°19 ʹ W , 31 .i .2015, F . Aballay leg . (1 IADIZA ); Tunuyán , Reserva Manzano Histórico , 1400 m , 33°36’S 69°19 ʹ W , 31 .i .2016, F . Aballay leg . (3 IADIZA ); Tunuyán , Reserva Manzano Histórico , 1800 m , 33°36’S 69° 19 ʹ W , 31 .xii .2015, F . Aballay and F . Jofré leg . (1 IADIZA ); Las Heras , Reserva Natural Villavicencio , 1000 m , 32°36’S 68°54 ʹ W , 31 .xii .2015, F . Aballay and F . Jofré leg . (3 IADIZA ); Las Heras , Reserva Natural Villavicencio , 1000 m , 32°36’S 68°54 ʹ W , 31 .i .2016, F . Aballay and F . Jofré leg . (1 IADIZA ); Las Heras , Reserva Natural Villavicencio , 1400 m , 32°36’S 68°54 ʹ W , 31 .i .2016, F . Aballay and F . Jofré leg . (2 IADIZA ); San Carlos , Pareditas , 18 km S San Carlos , [road to] Laguna del Diamante , 1000 m , 33°57’S 69° 4 ʹ W , 31 .i .2015, F . Aballay leg . (1 IADIZA ); Capital , 25 km W Mendoza , 7 .ii .1997, G . Arriagada leg . ( 1 ♂ , 1 ♀ , 22 CVMD ); Santa Rosa , Reserva da Biosfera de Ñacuñan , 12 .xii .2002, cow [dung or carcase?], R . Gomez leg . (1 CVMD ); Santa Rosa , Reserva da Biosfera Ñacuñan , 11–13 .xii .2002, V .M . Diéguez leg . (16 CVMD ); La Paz , Desaguadero , 28–29 .ii .2005, Roberto Barrera leg . (2 CVMD ); La Paz , Desaguadero , 10–18 .ii .1998, Daniel Vazquez leg . (1 CVMD ); La Paz , Desaguadero , 8–19 .i .1996, D . Escobar leg . (1 CVMD ); Lavalle , Reserva Natural Bosques Telteca , 8–10 .xii .2002, V .M . Diéguez leg. (7 CVMD ); no further data (1 MACN ). URUGUAY : Río Negro , Menafra , 15 .i .1914, no collector (1 MLPA ); Menafra , 4 .x .1914, no collector (1 MLPA ). no specific locality or date, A . Breyer leg . (13 MACN ); no specific locality or date, G . Pellerano leg . (1 MACN ); no specific locality, 6 .i .1899 , no collector (1 MLPA ); no specific locality, 4 .x .1906 , no collector (1 MLPA ); no specific locality, 23 .x .1910 , no collector (1 MLPA ); no specific locality, 23 .v .1922 , underneath a horse skeleton (1 MLPA ). No specified country: South America [‘ Am. Bor. ’] (156 MACN ; 27 MLPA ; 1 IADIZA ) . Soriano , Mercedes , 8 . i .1911, no collector (4 MLPA ); no further data (2 MACN ). Lavalleja , Minas , i .1980, W .D . Roverd leg. (5 NHMP ). Canelones , Las Piedras, 18 .i .1968 , A . Carmenes leg . (9 CVMD ); Solymar , 12 .i .1964, W .D . Roverd leg. (3 NHMP ). Montevideo , Montevideo , 25 .ii .1909, J. B. leg. (1 MACN ); Montevideo , 22 .xi .1921, J . Tremoleras leg . (1 MLPA ); Montevideo , 15 . x .1899, no collector (1 MLPA ); Montevideo , 22 .xii .1921, no collector (1 MLPA ); Montevideo , 18 .xii .1890, M . Montoro leg . (1 MLPA ); Montevideo , 22 .ii .1922, no collector (1 MLPA ); Montevideo , 12 .i .1909, no collector (1 MLPA ); Montevideo , 16 .xii .1906, light trap [‘a la lampada’], no collector (1 MLPA ); Montevideo , 15 .iii .1929, no collector (1 MLPA ); Montevideo , 12 .iii .1927, no collector (1 MLPA ); Montevideo , 15 .i .1929, no collector (1 MLPA ). Figure 7. Holotype of Omorgus ( Omorgus ) triestinae Pittino, 1987 . A, dorsal view; B, lateral view; C, aedeagus in dorsal view and D, holotype labels (Photos by A. Mantilleri, copyright MNHN). Diagnosis Omorgus suberosus resembles O. triestinae . It can be distinguished from the latter by the sculpture of the even numbered elytral costae. In O. suberosus the even numbered costae are barely raised, whereas O. triestinae has the even numbered costae distinctly raised. The male genitalia of the two species are distinctive ( Figures 6b and 7c ). For a more detailed treatment of the species see Pittino (1987) and Huchet and Costa-Silva (2018) . Geographic distribution This ubiquitous New World species is now established in Australia , parts of Europe and northern Africa. It has also been recorded from Fiji , New Caledonia , the Cape Verde Islands and the Canary Islands ( Huchet and Costa-Silva 2018 ). In Brazil , this species has been so far recorded in all biomes and states, except for AP, AL and SE ( Figure 11 ). The absence of actual records notwithstanding, the authors agree that the distribution of the species should contemplate these states.According to Huchet and Costa-Silva (2018) and Hielkema and Hielkema (2019) , this species is present in all countries of South America, with exception of Chile (see Diéguez 2008 ).