A review of Mexican Stamnodes (Lepidoptera: Geometridae) with the description of 16 new species Author Matson, Tanner A. AD85C521-0781-451F-92F6-D32B08C21395 Department of Ecology and Evolutionary Biology, University of Connecticut, Storrs, Connecticut 06269 – 3043, USA. National Museum of Natural History, Smithsonian Institution, Washington D. C., USA. MatsonT@SI.edu text European Journal of Taxonomy 2023 2023-12-14 911 1 79 https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2371/10397 journal article 10.5852/ejt.2023.911.2371 2118-9773 10376790 DB29E6F1-7925-46DB-8C9E-055C639203CE Stamnodes matrona sp. nov. urn:lsid:zoobank.org:act: 3C7D2A28-E94B-4E2B-9914-3CFDB1570B2B Figs 9 , 48 , 66 , 79 , 95 Diagnosis Stamnodes matrona sp. nov. is not easily confused with other Stamnodes , but as this species is likely sympatric with S. favilla sp. nov. , a brief diagnosis with respect to the latter is warranted. While S. favilla has a dark red outlining of the white patterning of the forewing underside apex and hindwing underside, such is absent in S. matrona . Furthermore, the male phallus altogether lacks cornuti, which should separate this species from S. favilla and all other similar species. Etymology The specific name matrona , a noun in the nominative case, is derived from the Latin ‘ matrona ’, meaning ‘lady, married woman, wife’. Material examined Holotype MEXICO • ♂ ; Nuevo León , 4 mi. W of Iturbide ; [ 24.73° N , 99.97° W ]; elev. 5500 ft ; 22 Sep. 1975 ; J. Powell , J. Chemsak and T. Friedlander leg.; at light; EMEC1748430 . Paratypes ( 6 ♂♂ , 1 ♀ ) MEXICO – Nuevo León • 3 ♂♂ ; same collection data as for holotype; EMEC1748430 to EMEC1748432 • 1 ♂ ; 4 mi W Iturbide ; [ 24.74° N , 99.97° W ]; elev. 5500 ft ; 13–14 Sep. 1976 ; J. Powell and J. Chemsak leg.; black light; EMEC1748433 • 1 ♂ ; same collection data as for preceding; 22 Sep. 1975 ; J. Powell , J. Chemsak , T. Friedlander leg.; at light; AMNH _ IZC 00353019 • 1 ♂ ; same collection data as for preceding; 22 Sep. 1975 ; J. Powell , J. Chemsak , T. Friedlander leg.; at light; AMNH _ IZC 00353019 • 1 ♀ ; same collection data as for preceding; 22 Sep. 1975 ; J. Powell , J. Chemsak , T. Friedlander leg.; at light; genitalia: TAM-2023-284 ( USNM 154214 ); USNMENT01771226 • 1 ♂ ; highway 60, 4 mi. E of Galeana Jct. ; [ 24.83° N , 100.15° W ]; elev. 6000 ft ; 23 Sep. 1975 ; J. Powell , T. Friedlander leg.; at light; genitalia: TAM-2023-280 ( USNM 154215 ); USNMENT01771225 . Description Male FOREWING LENGTH . 15–16 mm (n = 7). HEAD . Antenna filiform, fuscous. Vertex mostly pink. Frons mostly black, but white and pink dorsally, and thinly white along lateral and ventral margins. Labial palpus short, slightly porrect, subequal to diameter of eye, mostly fuscous. Cephalic collar pink and white. THORAX . Patagium pink and white; tegula fuscous. Mesothorax fuscous above, whitish below. Legs mostly fuscous, though coxa and proximal half of femur given more toward white; tibial spur formula 0–2–4; epiphysis well developed. FOREWING . Ground colour yellow-orange. Costa faintly grey; apical area black with yellow-orange and white patch below costa. Underside like upperside but apical patch white, costa darker grey, and costal area with scarlet suffusion. Fringe starkly checkered along outer margin and given toward pink along inner margin. HINDWING . Concolourous with forewing above. Underside much different than upperside, predominantly dark grey, blotted with large irregular white patches. White patches as follows: irregular patches at base; subovate patches at costal median and apex; large, angled patch extending from median of outer margin to median of inner margin; and small irregular patch about the tornus (see Fig. 9b ). Patches not finely outlined as in other Stamnodes . Fringe pink. ABDOMEN . Fuscous above, white below. GENITALIA ( Fig. 48 ). Uncus long, slender, and tapering. Subscaphium well developed. Juxta wide and U-shaped with posterolateral, long, conical processes. Inner face of valva with two hair tufts: smaller tuft arising basally from digitate tubercle; second, larger hair tuft more laterally widened, positioned in slight depression, and extending to subapical area of valva. Costal sclerite with medial swelling and terminating just short of apex. Vesica without cornuti. Female Outwardly undifferentiated from male. FOREWING LENGTH . 17 mm (n = 1). GENITALIA ( Fig. 66 ). Anterior apophysis two-thirds length of posterior apophysis. Ductus bursae with prominent sclerite flattened on ventral surface and dorsolaterally rolled toward median. Corpus bursae ovoid with inwardly, depressed circular signum bearing numerous minute papillae. Distribution ( Fig. 79 ) Mexico : Stamnodes matrona sp. nov. is known from the Sierra Madre Oriental pine-oak forests of Nuevo León . Biology Stamnodes matrona sp. nov. flies in September. The immature stages remain unknown but are likely hosted by mints ( Lamiaceae ). Molecular characterization This species has not been sequenced.