Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java Author Nurinsiyah, Ayu Savitri 5AFBFEFB-B243-49CC-AACC-248B2F007BC8 Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia. ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id Author Neiber, Marco T. 8F6006F5-D880-4841-AC32-B97E18D7BB5C Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany mneiber@hotmail.de Author Hausdorf, Bernhard FEDE92C0-108D-4315-8F86-9F2CFF1CB906 Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. hausdorf@zoologie.uni-hamburg.de text European Journal of Taxonomy 2019 2019-05-21 526 1 73 journal article 10.5852/ejt.2019.526 2da65c0a-9c48-43f5-ada3-e088de28dec1 2118-9773 3152217 3FFC527D-E9CA-4D9F-BF59-076FA60171AC Landouria zonifera sp. nov. urn:lsid:zoobank.org:act: 81A2A657-B12B-43BE-B19D-22AE36E2525C Figs 44 , 54 , 68 , 82 ; Tables 3–4 Diagnosis Landouria zonifera sp. nov. is characterized by a keeled, corneous shell with a brownish zone at the periphery, a wide umbilicus and a flagellum that is more than twice as long as the short epiphallus. Etymology Landouria zonifera sp. nov. is named after the brownish zone at its shell periphery (Latin: ʻzonaʼ = ʻzoneʼ, ʻferreʼ = ʻto carryʼ; used as an adjective). Material examined Holotype INDONESIA • holotype (det. anat.); Yogyakarta , Kulonprogo , Sibolong , snakefruit plantation ; 7°44′44″ S , 110°08′02″ E ; 716 m a.s.l. ; 9 Jan. 2013 ; A.S. Nurinsiyah and T. Setiadi leg.; D = 13.2 mm , H = 7.6 mm ; MZB 17727 . Paratypes INDONESIA • 1 spec. ; Yogyakarta , Kulonprogo , Sibolong , Sibolong Hill , agroforestry ; 7°45′00″ S , 110°07′00″ E ; 721 m a.s.l. ; MZB 17762 • 13 spec. ; same data as for holotype; MZB 17720 • 1 spec. ; same data as for holotype; ZMH 133420 • 4 spec. ; Kulonprogo , Jonggrangan ; 7°44′20″ S , 110°08′32″ E ; 730 m a.s.l. ; ZMH 140031 . Description SHELL (Fig. 82; Tables 3–4 ). Depressed conical, with 5.5 –5.75 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles and indistinct incised spiral lines; scaly processes sparse, mainly at periphery; tuberculate around umbilicus; corneous with a brownish zone at periphery; body whorl keeled; aperture rounded rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 40% of shell diameter, hardly obscured by columellar edge. GENITALIA ( Figs 44 , 68 ; Table 4 ). Atrium short; penis long, cylindrical, slightly dilated at proximal end, without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to proximal part of short, cylindrical epiphallus; flagellum more than twice as long as epiphallus, broadest near its proximal end, then abruptly tapering, slightly crenated, without node; vas deferens narrow and long; vagina very long, cylindrical; oviduct shorter, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina. Remarks Landouria zonifera sp. nov. clusters in the molecular phylogeny ( Fig. 1 ) with L. abdidalem sp. nov. (Fig. 56) and L. tholiformis sp. nov. (Fig. 51). Landouria zonifera sp. nov. and L. abdidalem sp. nov. have overlapping ranges. Landouria zonifera sp. nov. differs from L. abdidalem sp. nov. in the smaller (D 12.5–13.2 mm vs 13.1–16.4 mm in L. abdidalem sp. nov. ), more depressed (D/H 1.73–1.81 vs 1.93–2.14 in L. abdidalem sp. nov. ), corneous shell with a brownish zone at the periphery, the shorter penis and a flagellum that is more than twice as long as the short epiphallus. Furthermore, Landouria zonifera sp. nov. occurs sympatrically with L. menorehensis sp. nov. ( Fig. 88 ) from which it can easily be distinguished by the keeled, depressed conical, corneous shell with a brownish zone at the periphery. Distribution Landouria zonifera sp. nov. is endemic to the Menoreh karst area (Fig. 54).