Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java Author Nurinsiyah, Ayu Savitri 5AFBFEFB-B243-49CC-AACC-248B2F007BC8 Museum Zoologicum Bogoriense, Research Center for Biology, Indonesian Institute of Science, Cibinong, West Java, Indonesia. ayu_nurinsiyah@yahoo.com & ayus002@lipi.go.id Author Neiber, Marco T. 8F6006F5-D880-4841-AC32-B97E18D7BB5C Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany mneiber@hotmail.de Author Hausdorf, Bernhard FEDE92C0-108D-4315-8F86-9F2CFF1CB906 Zoological Museum, Center of Natural History, University of Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany. hausdorf@zoologie.uni-hamburg.de text European Journal of Taxonomy 2019 2019-05-21 526 1 73 journal article 10.5852/ejt.2019.526 2da65c0a-9c48-43f5-ada3-e088de28dec1 2118-9773 3152217 3FFC527D-E9CA-4D9F-BF59-076FA60171AC Landouria smimensis ( Mousson, 1848 ) Figs 11 , 43 , 52 , 67 , 78–80 ; Tables 3–4 Helix smimensis Mousson, 1848: 266 (“ Java ”; more exactly ( Mousson 1849 ): “Smim, 4000′”). Helix smironensis Mousson, 1850: 177 (unjustified emendation). Helix smiruensis Martens, 1867: 268 (unjustified emendation). Plectotropis tenggerica Schepman, 1912: 233 , pl. 10, figs 9–11 (“Nongkodjadjar”). Helix smimensis – Mousson 1849: 21 , pl. 2, fig. 10. Plectotropis sp. – Schepman 1915: 19 . Landouria smironensis – van Benthem Jutting 1950: 467 (in part). Diagnosis Landouria smimensis is characterized by a small, depressed conical shell with hairs and a usually slightly angular body whorl, a long penis, which gradually tapers towards its proximal end, and a slightly crenated, gradually tapering flagellum. Material examined Type material INDONESIA – East Java • holotype of P. tenggerica ; Nongkojajar ; 7°54′ S , 112°49′ E ; ZMA 174453 • 2 syntypes of H. smimensis ; Gunung Semeru ; 8°06′ S , 112°55′ E ; ca 1200 m a.s.l. , ZMZ 508598 . Other material INDONESIA • 23 spec. (vouchers for Martens 1867 ); Java ; ZMB 4062 • 1 spec. (voucher for van Benthem Jutting 1950 ); Java ; ZMB 67043 . – East Java • 3 spec. (vouchers for van Benthem Jutting 1950 ); Nongkojajar , near waterfall ; 7°54′ S , 112°49′ E ; ZMA 407685 • many spec. (vouchers for van Benthem Jutting 1950 ); same data as for preceding; ZMA 407681 • 7 spec. ; Nongkojajar , near Baron ; 7°54′ S , 112°49′ E ; ZMA 407686 • 2 spec. (vouchers for van Benthem Jutting 1950 ) (det. anat.); Tosari ; 7°53′ S , 112°54′ E ; ca 1800 m a.s.l. ; ZMA 390881 . Description SHELL ( Figs 11 , 78–80 ; Tables 3–4 ). Depressed conical, with 4.5–5.5 convex whorls; protoconch almost smooth; teleoconch irregularly striated, with indistinct incised spiral lines; with hair-like processes; not tuberculate around umbilicus; brownish-corneous; body whorl slightly angular or rounded; aperture oval; upper insertion of peristome hardly descending; peristome sharp; umbilicus slightly eccentric, narrow, comprising 18–32% of shell diameter, hardly obscured by columellar edge. GENITALIA ( Figs 43 , 67 ; Table 4 ). Atrium short; penis long, gradually tapering towards its proximal end; without penis coecum; surrounded by a membranaceous sheath connected to proximal epiphallus by tissue; penial retractor runs from diaphragm to middle part of cylindrical epiphallus; flagellum moderately long, narrow, broadest in its middle part, without node, gradually tapering towards its proximal end, slightly crenated; vas deferens narrow and long; vagina broad and short; oviduct short, proximally slightly tapering; peduncle of bursa copulatrix subdivided into a broad abruptly tapering distal part and a narrower and longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina. Remarks The original spelling of the species name was Helix smimensis Mousson, 1848 . This was not an inadvertent error, such as a lapsus calami or printer’s error, because this spelling was also used later by Mousson (1849) . Thus, H. smimensis is the correct original spelling according to ICZN, Article 32. The name was intentionally changed to H. smironensis by Mousson (1850) and to H. smiruensis by Martens (1867) . Both names are unjustified emendations, which are available with these authors and dates, and which are junior objective synonyms of H. smimensis (ICZN, Art. 33). According to ICZN, Article 33.2.3.1, an unjustified emendation may be deemed to be a justified emendation, if it is in prevailing usage. Helix smiruensis was used by Tryon (1887) , Leschke (1914) and Rensch (1932) , L. smironensis by van Benthem Jutting (1950) and Vermeulen & Whitten (1998) , the incorrect subsequent spelling Landouria cf. smiroensis by Boonngam et al. (2008) and the correct original spelling L. smimensis by Köhler et al. (in press) . Thus, there is no prevailing usage of any of the emendations and the correct original spelling should be used. The voucher specimen for L. tenggerica of Rensch (1934) from Gunung Lawu, 2400 m a.s.l. (ZMB 76086) is tuberculate around the umbilicus and is a juvenile of a different species. Distribution Landouria smimensis is known from the Tengger Mountains in East Java (Fig. 52). It has also been recorded from other mountains in Central and East Java ( Rensch 1934 ), Lombok ( Rensch 1932 ), Bali ( Vermeulen & Whitten 1998 ) and Thailand ( Boonngam et al. 2008 ), but these populations probably represent separate species.