A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions Author Shi, Hongliang Author Sciaky, Riccardo Author Liang, Hongbin Author Zhou, Hongzhang text Zootaxa 2013 3664 2 101 135 journal article 10.11646/zootaxa.3664.2.1 a50b71f5-48a6-464f-aea1-f29bbc8ee322 1175-5326 220503 105D178E-6DBF-401D-85BB-CDB041B67E91 Pterostichus ( Wraseiellus ) kambaiti (Andrewes, 1947) ( Figures 17, 18 , 27, 28 , 40 , 78 , 106 ) Andrewes 1947: 22 (Original: Feronia ; holotype deposited in NHRS); Lorenz 2005: 287 ( Pterostichus ( incert. sed. )). Synonym: kambaiti Jedlička 1965: 204 ( Pterostichus ( incert. sed. ); holotype deposited in NHRS), Junior secondary homonym of Pterostichus kambaiti (Andrewes 1947) ; Lorenz 1998: 266 (catalogue; synonymized with kambaiti Andrewes ). Type locality . Myanmar : Kambaiti , 7000 ft . ( N25.40º , E98.13º ). Type materials examined . Holotype of Feronia kambaiti Andrewes (examined by photo): female (NHRS), body length = 10.3 mm , board mounted, "N. E. BURMA / Kambaiti , 7000 ft / 19/4 1934 / R. MALAISE"; " Feronia / Kambaiti / Type Andr . / H.E.Andrewes det."; " Typus " [red label]; "5534 / E91 +" [blue label]; "NHRS- COLE / 000008537"; " kambaiti / Andr ." [box label] [ Figs. 17 , 27 ]. Holotype of Pterostichus kambaiti Jedlička : male (NHRS), body length = 9.9 mm , board mounted, genitalia dissected and glued on mounting board together with specimen, "N. E. BURMA / Kambaiti , 7000 ft . / 19/4 1934 / R. MALAISE"; " 4 "; " HOLOTYPE " [red label]; " Pterostichus / kambaiti / sp.n. / det. Ing. Jedlička" [pink label]; "5679 / E91 +" [blue label] [ Figs. 18 , 28 , 40 , 78 , 106 ]. Notes on types . Feronia kambaiti Andrewes : The original description indicated that this species was published on two females, and in the preface noted that all the " types " were in Stockholm Museum and the " paratypes " were in the collection of Andrewes and British Museum. So, the one we examined by photo with the help of Dr. J. Bergsten is the holotype of this species ( Figs. 17 , 27 ). Pterostichus kambaiti Jedli č ka : The original description clearly indicated that this species was described on a single male. Therefore, the one we borrowed from NHRS is certainly the holotype of this species ( Figs. 18 , 28 ). Diagnosis. In the external characters, this species is most similar to Pt. stictopleurus s.str. These two species can be distinguished from other members of this subgenus by the combination of the following characters: (1) 5th tarsomere glabrous beneath; (2) males with isodiametric microsculpture on elytra; (3) pronotum with area between outer basal foveal groove and lateral margin slightly convex, not forming a distinct ridge; (4) lateral margins straight before hind angle, hind angles obtuse, not pointed. Pt. kambaiti is different from Pt. stictopleurus s.str. in: (1) scutellar striae located between 1st and 2nd striae (scutellar striae located between 1st stria and elytral suture in Pt. stictopleurus s.str. ); (2) metaepisterna impunctate (distinctly punctate in Pt. stictopleurus s.str. ); (3) body size smaller, 9.9–10.3 mm (usually much larger in Pt. stictopleurus s.str. , 10.8–13.1 mm ); (4) right paramere much wider than in Pt. stictopleurus s.str. ; (5) apex of median lobe of male genitalia more bent to the dorsal side than in. Pt. stictopleurus s.str. Description . Body length 9.9–10.3 mm ; dorsal side black, elytra somewhat lighter, shining in males. Males with elytral microsculpture isodiametric. Head . Frons and vertex impunctate, a few punctures present in frontal furrows; antennae reaching elytra basal eighth; 3rd antennomere glabrous except apical setae. Pronotum rounded, widest at about middle, PW/PL = 1.30; lateral margins fully rounded near the middle and then gradually curved posteriorly, almost straight before hind angles; hind angles obtuse, not pointed; one mid-lateral seta present at about anterior fourth; basal fovea shallow, faintly defined; inner groove of basal fovea weak, subparallel to median line, gradually disappeared posteriorly, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea about half length as inner one, somewhat more distinct than the inner one, reaching the basal margin of pronotum; ridge between outer groove and lateral margin slightly convex; area between inner and outer basal foveal grooves hardly convex; some faint punctures present in basal fovea area and the area between the two basal foveae. Elytra with base slightly depressed between 3rd and 5th intervals; elytral shoulders slightly narrowed, humeral teeth small; intervals slightly convex; striae distinct, without punctures; scutellar striae nearly complete, located between 1st and 2nd striae; umbilical pore series on 9th interval sparse in the middle, composed of 15 pores (5, 2, 8). Ventral side . Proepisterna and metaepisterna impunctate; terminal sternum of male without special structure. Legs . 5th tarsomere without seta beneath. Male genitalia . Median lobe of male genitalia bent at a 90 degrees angle at about basal two-fifth; ventral margin almost straight in the middle, apical part slightly bent to the ventral side and then the apical lamella gradually bent to the dorsal side ( Fig. 40 A); apical orifice large, slightly turned to the left side, apical lamella twisted to the left ( Fig. 40 B); apical lamella laminar, about 1.5 times as its basal width, apex rounded. Right paramere short, apical part triangular and curved, narrowed apically, apex moderately wide ( Fig. 78 B). Female genitalia not studied. Distribution . Only known from the type locality, Kambaiti , Myanmar ( Fig. 110 ). Notes on synonym . From the collecting data label, the holotypes of Pt. kambaiti Jedlička 1965 and Feronia kambaiti Andrewes 1947 were collected at the same day in the locality of Kambaiti by Malaise. We did not find any significant difference between the two holotypes . So, there is no doubt that Pt. kambaiti Jedlička 1965 is both synonym and homonym of Pt. kambaiti (Andrewes 1947) .