A revision of the spider genus Raveniola (Araneae, Nemesiidae). II. Species from Central Asia
Author
Zonstein, Sergei L.
urn:lsid:zoobank.org:author:EADD3607-30FF-49AE-93F5-8410630469BE
Steinhardt Museum of Natural History, Tel-Aviv University, 69978 Tel-Aviv, Israel
znn@tauex.tau.ac.il
text
European Journal of Taxonomy
2024
2024-10-24
967
1
185
https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2699/12459
journal article
10.5852/ejt.2024.967.2699
2118-9773
13990819
C08B8027-50CC-417E-BCD4-5183B9FF6738
Raveniola karategensis
sp. nov.
urn:lsid:zoobank.org:act:
631DA123-F0BE-4C2B-A58E-F073BB22E75B
Figs 44
,
119
,
176
,
238
,
316–317
,
510–512
,
576
, 659–666, 675, 678–679, 753
Diagnosis
The new species shares with
Raveniola afghana
sp. nov.
,
R. alajensis
sp. nov.
and
R
.
hirta
sp. nov.
the presence of the modified long hairs on the female tibia and metatarsus IV. Females of
R. karategensis
sp. nov.
can be distinguished from those of
R. afghana
and
R
.
hirta
by having a fuzzily ornamented (vs uniformly coloured) abdomen (see
Fig. 44
cf.
Figs 37, 42
), and from females of
R. alajensis
in possessing a less pronounced abdominal pattern and the distinctly configured spermathecae with wider bases, broadly spaced stalks and clearly wider inner branches (
Figs 44
,
510–512
cf.
Figs 38
,
491–493
).
Etymology
The specific epithet is a toponym referring to the range of this species, confined to the historical area Karategin (also Karategen), an eastern province of the Bukhara Emirate in the 19
th
century, which included the Karategin Mts and Peter I Mts.
Material examined
Holotype
TAJIKISTAN
•
♀
;
Peter I Mts
(southern slope),
Harvikush Canyon
,
35 km
ENE of Tavildara Village
;
38°52′ N
,
70°49′ E
;
1900–2200 m
a.s.l.
;
10 Jul. 2019
;
S. Zonstein
and
A. Hakimov
leg.;
SMNH
.
Paratypes
(
10 ♀♀
)
TAJIKISTAN
•
4 ♀♀
; same collection data as for preceding;
SMNH
•
3 ♀♀
; same collection data as for preceding;
9 Jul. 1978
;
V
.
I. Ovcharenko
leg.;
SMNH
•
3 ♀♀
; same collection data as for preceding;
ZISP
.
Description
Female
(
holotype
)
HABITUS
. See
Fig. 44.
MEASUREMENTS
. TBL 20.95, CL 7.46, CW 6.10, LL 0.59, LW 1.37, SL 3.88, SW 3.45.
COLOUR
. Carapace dull reddish brown with clypeus and eye tubercle even darker brown, eyes encircled with partially fused wide blackish brown rings; chelicerae dark cherry red brown; sternum, labium, maxillae, epigastrum, book-lungs and spinnerets light yellowish brown; palps and legs light to medium brownish orange; abdomen medium brown, dorsally with several small light brownish spots and diffuse dark brown chevron-like pattern.
CEPHALOTHORAX
. Carapace and chelicerae as shown in
Fig. 119
. Clypeus and eye group as in
Fig. 176
. Eye diameters and interdistances: AME 0.16(0.20), ALE 0.28, PLE 0.28, PME 0.19; AME–AME 0.21(0.17), ALE–AME 0.20(0.18), ALE–PLE 0.22, PLE–PME 0.09, PME–PME 0.55. Weak rastellum composed of 30–40 slightly thickened spikes on anterior cheliceral edge. Each cheliceral furrow with 9 promarginal teeth and 6–7 mesobasal denticles. Sternum, labium and maxillae as shown in
Fig. 238
. Maxillae with 26–29 cuspules each.
LEGS
. Tibia and metatarsus IV densely covered with long modified hairs, as shown in
Fig. 316
. Scopula: entire and distal on metatarsi I–II; entire on palpal tarsus and tarsi I–II; sparse and widely divided by setae on tarsi III–IV. Trichobothria: 2 rows of 8–9 each on tibiae, 15–17 on metatarsi, 17–20 on tarsi. Palpal claw with 4 promarginal teeth. PTC I–IV with 6–7 teeth on each margin.
LEG
MEASUREMENTS
.
|
Femur
|
Patella
|
Tibia
|
Metatarsus
|
Tarsus
|
Total
|
| Palp |
3.74 |
1.98 |
2.47 |
– |
2.47 |
10.66 |
| Leg I |
5.19 |
2.97 |
3.63 |
3.34 |
2.35 |
17.48 |
| Leg II |
4.61 |
2.26 |
2.73 |
2.92 |
2.36 |
14.88 |
| Leg III |
4.26 |
2.23 |
2.51 |
3.41 |
2.19 |
14.60 |
| Leg IV |
5.83 |
3.12 |
4.42 |
5.91 |
2.96 |
22.24 |
SPINATION
. All femora with one basodorsal spine and 3–4 median and/or apical bristles; tarsi I–IV aspinose. Palp: femur pd1; patella p1; tibia p1(0), v6; tarsus v2. Leg I: femur pd2; patella p1; tibia p1(0), v4; metatarsus v4. Leg II: femur pd3; patella p1; tibia p2, v6; metatarsus v7(6). Leg III: femur pd4(3), rd3; patella p3, r2(1); tibia d1, p2, r3, v7; metatarsus p2, r3, v7. Leg IV: femur rd3(2); patella r1; tibia r3, v7; metatarsus p2, r4, v9.
SPERMATHECAE
. Each of paired spermathecae U-shaped with a relatively low and wide base carrying two more or less broadly spaced and unevenly shaped branches: a longer and wider trapezoidal inner branch and a more slender, shorter and club-like outer one (
Fig. 510
).
SPINNERETS
. See
Fig. 576
. PMS: length 0.89, diameter 0.36. PLS: length of basal, medial and apical segments 1.55, 0.73, 0.42; total length 2.70; apical segment triangular.
Male
Unknown.
Variation
Carapace length in
paratype
females (n=7) varies from 6.71 to 7.88. All the examined females are habitually very similar to each other. Variations in the structure of the eye group, tibia and metatarsus IV, and the spermathecae as shown in
Figs 177
,
317
,
511–512
.
Ecology
The spiders were found in the midland zone at an altitude of
1900–2000 m
a.s.l. inhabiting open woodland dominated by
Juniperus seravschanica
. Females of
Raveniola karategensis
sp. nov.
live in open burrows of
30–40 cm
depth, provided with a weakly silk-lined entrance rim, walls and living chamber (
Figs 659–666
,
675, 678–679
).
Distribution
Tajikistan
: Peter I Mts. See
Fig. 753
.