On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini) Author Assing, Volker text Beiträge Zur Entomologie = Contributions to Entomology 2017 2017-06-30 67 1 117 192 journal article 2472 10.21248/contrib.entomol.67.1.117-192 39f2d8c0-d0e4-46ea-8d5d-63e668fe9e43 0005-805X 5742363 FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE Zyras ( Zyras ) ambulans spec. nov. urn:lsid:zoobank.org:pub: FD33C1AE-F7D9-4E3A-A053-A2CAA7261CFE ( Figs 34 , 70 , 107 , 212–215 , Map 10 ) Type material : Holotype ♂: “ Thailand , Satun Prov. , Thale Ban N.P. , 20 km E Satun , 200–400 m , 1.–4.I.1996 , leg. Schulz & Vock / Holotypus ♂ Zyras ambulans sp. n. , det. V. Assing 2016” (cAss). Etymology : The specific epithet is the present participle of the Latin verb ambulare (to walk) and alludes to absence of functional hind wings. Description : Body length 6.2 mm ; length of forebody 2.5 mm . Coloration ( Figs 34 , 70 , 107 ): head blackishbrown; pronotum and elytra dark-brown; abdomen with segments III–V reddish, tergite VI dark-brown with reddish margins, VII and VIII dark-brown with the anterior and posterior margins narrowly paler; legs yellowish with the apices of the meso- and metafemora brown; antennae dark-brown with antennomeres I–II reddish and XI yellow; maxillary palpi dark-yellowish with the terminal palpomere pale-yellowish. Head ( Fig. 70 ) moderately transverse, middle extensively impunctate; punctation in lateral dorsal portions sparse and fine. Eyes large and bulging, much longer than postocular region in dorsal view. Antenna ( Fig. 34 ) 2.3 mm long; antennomeres IV–V oblong, VI weakly oblong, VII as long as broad or weakly oblong, VIII–IX very weakly transverse, X less than 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X. Pronotum ( Fig. 70 ) weakly transverse, approximately 1.1 times as broad as long and 1.2 times as broad as head, broadest anteriorly, distinctly tapering posteriad; lateral margins straight in posterior two-thirds (dorsal view); punctation shallow and sparse; midline broadly impunctate; lateral margins anteriorly with two very long, stout, dark setae. Elytra ( Fig. 70 ) short, approximately 0.7 times as long as pronotum; punctation moderately dense, moderately coarse, and regularly distributed. Hind wings reduced. Metatarsomere I longer than the combined length of II and III, but shorter than the combined length of II–IV. Abdomen ( Fig. 107 ) broader than elytra, with moderately deep anterior impressions on tergites III–V; anterior impressions of tergites III–V without non-setiferous punctures; tergites III–VI with a lateral puncture bearing a long black seta on either side and with a transverse row of approximately 10 setiferous punctures at or near posterior margins, at least the lateral punctures of these rows bearing long black setae; tergite VII anteriorly with a cluster of rather sparse shallow non-setiferous punctures and posteriorly with sparse setiferous punctures, posterior margin with narrow palisade fringe; tergite VIII with sparse setiferous punctures only in posterior portion, posterior margin smoothly convex, without median excision or other modifications. ♂: sternite VIII with convex posterior margin; median lobe of aedeagus 0.65 mm long and shaped as in Figs 212–213 ; paramere ( Fig. 214 ) much longer than median lobe ( 0.9 mm ), with long, flattened, and somewhat club-shaped apical lobe ( Fig. 215 ). Comparative notes : Zyras ambulans is distinguished from all the species previously known from Thailand and Peninsular Malaysia by the reduced hind wings and the morphology of the aedeagus (shape of median lobe; modifications of paramere). It is additionally characterized by the coloration (body, legs, and antennae), a slender pronotum, short elytra, and the punctation pattern of the abdomen. Distribution and natural history : Thale Ban National Park is situated in the extreme south of Thailand at approximately 6°43'N , 100°10'E , close to the border with Malaysia ( Map 10 ). The holotype was collected at an altitude between 200 and 400 m . Since one of the collectors is a myrmecologist, it may have been found associated with ants.