Taxonomic revision of the Afrotropical species of the tribe Eudorylini (Diptera, Pipunculidae) Author FÖLDVÁRI, MIHÁLY text Zootaxa 2013 2013-05-22 3656 1 1 121 http://dx.doi.org/10.11646/zootaxa.3656.1.1 journal article 20775 10.11646/zootaxa.3656.1.1 d2c3e582-04bb-4d44-8261-b62f1b0db73a 1175­5334 6039126 73E59AED-B16D-430C-B611-EB5CFEAF7FDD Eudorylas wittei ( Hardy, 1950 ) ( Figs 37G–L , 40U ) Dorilas (Eudorylas) wittei Hardy 1950: 39 . Diagnosis: Scutellum shining black, 6–9 pairs of short, pale hairs (as long as width of t3 at widest). Abdominal tergites shining black. Hairs dense, whitish, longest on T5 and distal part of T4. Lateral spines on first tergite present as a patch of 8–10 long hairs. Surstyli highly asymmetrical, with numerous hairs ventrally; ST8 enlarged, with narrow m.a.; gonopods asymmetrical; hypandrium short, hypandrial apodeme well developed; phallic guide pointed at tip and with protuberance on the right side covered with thick hairs; in lateral view with membranous part; phallus trifid, branches long, not separating from each other; ejaculatory apodeme flat, broad at tip; sperm pump with two wing-like projections. Type material: Rwanda : 1♂ , HT, Congo belge : Ruanda, Kundhuru ya Tsuve , (Col Gahinga-Sabinyo), 2600 m , (Bambous), 15-ix-1934 , G.F. de Witte : 601 ; 1♀ , AT, same data as HT, 18-ix-1934 , [ No. ] 611 ; 2♂ , 2♀ , PTs, same data as HT, Rutabagwe , 13-14-ix-1934 , [ No. ] 595 ; 1♀ , PT, Congo belge : Ruanda , Kansenze (pied Volc. Karisimbi ), 2400 m , 4-iii-1935 , G.F. de Witte : 1209 (all MRAC ) ; 1♂ , PT, same data as HT, Rutabagwe , 13-14-ix-1934 , 2600 m , [ No. ] 595 ( AMNH ) . Democratic Republic of Congo : 2♂ , PTs, Congo belge : PNA, Nyakibumba , (près Kikere ), 2226 m , 9-vii-1934 [as “ 2250 m 5.VII.1934 ” in Hardy 1950 ], G.F. de Witte : 478 ; 3♂ , PTs, Congo belge : P.N.A., Nyarusambo ( Kikere ), 2226m , 28–29.vi.1934 [as “VII” in Hardy 1950 ], G.F. de Witte : 453 (all USNM ) ; 1♂ , PT, same data as the specimen before ( AMNH ) . Other material examined: Kenya : 1♂ , 1♀ , Mt. Kenya. , Kathita river ., 9,900ft. , 7.viii.1949 ., J.A. Riley. ; " O.U.E.C. Exp , Mt Kenya, B.M. 1949-562."; "56" [hand-written] ; 1♂ , 1♀ , Mt. Kinangop , 8,000ft. , F.W. Edwards. ; Aberdare Range. , x.1934 ., B.M.E. Afr. Exp. , B.M. 1935-203. (all BMNH ) ; 1♂ , Mt. Kenya. , Naro Moru , 9,000ft. , 31.viii.1949 ., J.A. Riley. ; " O.U.E.C. Exp , Mt Kenya, B.M. 1949-562." ; 1♀ , Mt. Kenya. , Kathita river ., 9,900ft. , 7.viii.1949 ., J.A. Riley. ; " O.U.E.C. Exp , Mt Kenya, B.M. 1949-562."; "56" [hand-written] (both HNHM ) ; 1♂ , Ngong Forestry Sta [tion]. IV-13-20-1968 , Malaise trap , Paul J. Spangler ; 1♂ , 1♀ , same data, 13–18 Jan. 1968 , Krombein & Spangler ( ♂ HNHM , USNM ) . Rwanda : 1♀ , PT, Congo belge: Ruanda , Nyabitsindi (entre Volc. Bishoke-Musule ), 2400 m , 18-II-1935 , G.F. de Witte : 1159 ( MRAC ) , [the specimen is not listed in Hardy (1950) , therefore it is not paratype ]. South Africa : 1♂ , RSA: Cape Prov. Nature’s valley , at Groot Rivier , 33° 58´S , 23° 33´E , 15– 17.10.1994 , loc 21, leg. Roy Danielsson ( ZML ) ; 1♂ , Blaauwkrantz Pass , Nr. Plettenberg Bay, indigenous forest, E. Cape 11 Oct 1959 , B & P Stuckenberg, “ Pipunculus (Eudorylas) n.sp. freak spec. right wing Cephalosphaera, Det. 1961, D.E. Hardy ” ( NMSA ) . Uganda : 1♂ , Mt. Sabinio. , 8,000ft. , F.W. Edwards. ; Kigezi Distr. , xi.1934 ., B.M.E. Afr. Exp. , B.M. 1935- 203. ( BMNH ) . Zimbabwe : 1♂ , Rhodes-Inyanga National Park ., S. Rhodesia , 13 Jan. 1955 , B.R.S. P.G., ” Pipunculus (Eudorylas) n.sp? runs near P.(P.) pallidipleura Curran, Det. 1961 D.E. Hardy ” ( NMSA ) . Male Head . Third antennal segment acuminate; yellow, arista shining black. Face silvery pollinose. Frons, upper part shining black, lower part silvery pollinose; eyes touching for distance equal to 2 times ocellar triangle. Occiput, upper half subshining black, lower half silvery pollinose. Thorax . Humeri yellow. Mesonotum (viewed obliquely from front) subshining black, faintly greyish pollinose; silver grey from the side. Scutellum shining black, 6–9 pairs of short, pale hairs (as long as width of t3 at widest). Dorsocentral hairs distinct. Halter yellow-brown. Legs. Trochanters, femora brown (may be more yellow, in this case dorsal surface of femora always brown), knees, tibiae, tarsal segments yellow, last segment brown. Ventroapical row of 2–4 short black spines on first four femora; no spines on 3rd femur. Subapical (distal) spines on first four tibiae distinct. No anteromedial hairs on 3rd tibia. Hind trochanter covered with short, white hairs on ventral side. Pulvilli shorter than last tarsal segment. Wing. Fourth costal section 0.7–1.1 times as long as third costal section. Cross-vein R-M at 1/4 to 2/5 of discal cell. Pterostigma fully coloured. Hairs on tegula absent. Abdomen . Viewed obliquely from front tergites shining black. Hairs dense, whitish, longest on T5 and distal part of T4. Lateral spines on first tergite present as a patch of 8–10 long hairs. Postabdomen in dorsal view: T5 1.1 times as long as ST8. Genitalia without dissection: shining black with whitish hairs dorsally, SS and EP yellow, m.a. oval. Genitalia. Surstyli highly asymmetrical, with numerous hairs ventrally; ST8 enlarged, with narrow m.a. ( Fig. 37H ). SES uncertain; gonopods asymmetrical, outer longer; hypandrium short, hypandrial apodeme well developed; phallic guide pointed at tip and with protuberance on the right side covered with thick hairs ( Fig. 37L ); in lateral view with membranous part ( Fig. 37G ); phallus trifid, branches long, not separating from each other; ejaculatory apodeme flat, broad at tip; sperm pump with two wing-like projections ( Fig. 37J ). Female As male except for the following characters. Frons, eyes separated; silver-grey pollinose on lower half, shining black on upper half. Pulvilli only slightly longer than last tarsal segment. Female ovipositor as in Fig. 40U . Remarks— Specimens can be found among the material examined bearing the label “P.(P.) pallidipleura” by Hardy, referring to Cephalops (Cephalops) pallidipleura. Distribution— Democratic Republic of Congo, Kenya, Rwanda, South Africa, Uganda, Zimbabwe.