External egg structure of the Pentatomidae (Hemiptera: Heteroptera) and the search for characters with phylogenetic importance
Author
Matesco, Viviana Cauduro
Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gançalves, 9500, Bloco IV, Prédio 43435, 91501 - 970. Porto Alegre, RS, Brazil
vimatesco@yahoo.com.br
Author
Bianchi, Filipe Michels
Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gançalves, 9500, Bloco IV, Prédio 43435, 91501 - 970. Porto Alegre, RS, Brazil
bianchi.fm@hotmail.com
Author
Fürstenau, Brenda Bianca Rodrigues Jesse
Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gançalves, 9500, Bloco IV, Prédio 43435, 91501 - 970. Porto Alegre, RS, Brazil
brendabianca@gmail.com
Author
Silva, Priscila Poock Da
Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gançalves, 9500, Bloco IV, Prédio 43435, 91501 - 970. Porto Alegre, RS, Brazil
pripoock@terra.com.br
Author
Campos, Luiz Alexandre
Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gançalves, 9500, Bloco IV, Prédio 43435, 91501 - 970. Porto Alegre, RS, Brazil
luiz.campos@ufrgs.br
Author
Grazia, Jocelia
Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gançalves, 9500, Bloco IV, Prédio 43435, 91501 - 970. Porto Alegre, RS, Brazil
jocelia@ufrgs.br
text
Zootaxa
2014
2014-02-26
3768
3
351
385
journal article
46347
10.11646/zootaxa.3768.3.5
d8748b65-5cf8-489c-972a-85f4329f07d2
1175-5326
252047
076F3E92-066C-46A0-9ADE-09D3AA8F8707
Analysis of
Nezara
The analysis including three discrete and three continuous egg characters resulted in one most parsimonious tree with 63 steps (
Fig. 82
). The characters and states for eggs are as follows.
Character 35. Egg, color prior to embryonic development: (0) light brown, (1) yellow
Character 36. Egg, chorion, pigmentation: (0) present, (1) absent
Character 37. Egg, chorion, sculpture pattern: (0) spinose, (1) smooth
Character 38. Egg, length (range, in millimeters) (continuous)
Character 39. Egg, width (range, in millimeters) (continuous)
Character 40. Egg, aero-micropylar processes, number (range) (continuous)
None of the eggs’ characters were informative for the analysis. Characters 35 to 37 had ambiguous reconstruction due to missing data. The spinose chorion (character 37, state 0) is plesiomorphic but with uncertain evolution of the apomorphic smooth chorion. All values of egg length (character 38) overlap; the plesiomorphic state ranges from 1.12 to 1.20 mm. The egg width (character 39) values did not overlap, so the plesiomorphic condition for the entire group ranges from 0.95 to 1.00 mm and from 0.84 to 0.95 mm for the genus
Nezara
. The number of AMPs (character 40) also overlaps among species (except in
C. purpureipennis
); the plesiomorphic condition of the clade including
A. chloris
and
Nezara
spp. is 27 to 30 AMPs, whereas the plesiomorphic condition for the entire group is 25 to 30 AMPs. The availability of egg measures and AMPs number for other species of
Nezara
would be interesting to test these assumptions and to minimize the missing data in the analysis. Information on the eggs of
N. yunnana
Zheng
would be especially desirable to aid in solving the only polytomy of the cladogram.
Ferrari
et al.
(2010)
mentioned that a close relationship between
N. viridula
(Linnaeus)
and
N. antennata
Scott
has been suggested based on different sets of characters (e.g., interspecific mating behavior), but the lack of studies with
N. yunnana
prevented further conclusions. This highlights the importance of searching for the greatest diversity in number and source for characters but is also a reminder that for rare or poorly studied species, information will be missing on several sets of characters.