Staphylinidae (Insecta: Coleoptera) in Latin America: synopsis, annotated catalog, diversity and distribution Author Asenjo, Angélico Author Irmler, Ulrich Author Klimaszewski, Jan Author Chandler, Donald S. Author Fierros-López, Hugo Eduardo Author Vieira, Juliana S. text Zootaxa 2019 2019-06-21 4621 1 1 406 journal article 26396 10.11646/zootaxa.4621.1.1 a381190d-7f50-4a66-bc33-3a0417c9f89f 1175-5326 3257228 3D77F329-9BB8-4231-90E4-6A9E0250FCDF Subtribe Bolitocharina Thomson, 1859 0445. Bolitochara Mannerheim, 1830: 75 Type species: Aleochara pulchra Gravenhorst, 1806 Agaribiota Bierig, 1937b: 279 [synonymized by Blackwelder 1952: 41 ] Type species: Bolitochara cinctigastra Bierig, 1937 Ditropalia Casey, 1906: 263 [synonymized by Seevers 1978: 164 ] Type species: Bolitochara bella Märkel, 1844 3 species are included in this genus from the West Indies and South America. Characters/Description: Fenyes 1920: 114 (characters as Ditropalia ); Coiffait & Sáiz 1967: 65 (characters in key), 79 (characters); Ashe 1992: 365 (characters in key), 367 (characters). Key: Coiffait & Sáiz 1967: 80 (Chilean species). Distribution. West Indies: Cuba . South America: Colombia , Argentina . Other regions: Palaearctic, Oriental , Australian, Oceanic. 0446. Habramicrada Pace, 2007: 225 ( Fig. 58 ) Type species: Habramicrada boliviensis Pace, 2007 4 species are known from and are restricted to South America. Distribution. South America: Peru , Bolivia , Brazil . 0447. Leptusa Kraatz, 1856b: 60 Type species: Bolitochara pulchella Mannerheim, 1830 Subgenera: Adexiopisalia Pace, 1982c: 585 Type species: Leptusa hummleri Bernhauer, 1910 Adoxopisalia Pace, 1989: 130 Type species: Leptusa opaca Casey, 1893 Afroleptusa Pace, 1983c: 62 Type species: Leptusa sparsa Fauvel, 1898 Akratopisalia Pace, 1996a: 29 Type species: Leptusa cribrata Pace, 1996 Amalopisalia Pace, 1983c: 97 Type species: Leptusa janetscheki Scheerpeltz, 1956 Anisopisalia Pace, 1989: 214 Type species: Leptusa luigionii Bernhauer, 1935 Anomopisalia Pace, 1983c: 82 Type species: Leptusa granulicauda Eppelsheim, 1890 Anosiopisalia Pace, 1995: 247 Type species: Leptusa nemoricultrix Pace, 1995 Aphaireleptusa Pace, 1996a: 30 Type species: Leptusa anmashanensis Pace, 1996 Aleteleptusa Pace, 1997a: 753 [ Smetana 2004: 429 ] Type species: Leptusa chinensis Pace, 1997 Mimumenepisalia Pace, 1997a: 752 [ Smetana 2004: 429 ] Type species: Leptusa gonggamontis Pace, 1997 Baldopisalia Pace, 1983e: 130 Type species: Leptusa veronensis Pace, 1983 Boreoleptusa Pace, 1989: 82 Type species: Leptusa norvegica Strand, 1941 Bothrydiopisalia Scheerpeltz, 1966b: 20 Type species: Leptusa bosnica Eppelsheim, 1892 Bryopisalia Pace, 1989: 239 Type species: Leptusa pilatensis Scheerpeltz, 1972 Chondrelytropisalia Scheerpeltz, 1976c: 55 Type species: Leptusa nepalica Scheerpeltz, 1976 Chondridiopisalia Pace, 1983c: 85 Type species: Leptusa granulifera Pace, 1983 Chondridiopisalia Scheerpeltz, 1966b: 19 [ nomen nudum , Newton & Thayer 2003 ] Type species: Leptusa granulifera Scheerpeltz, 1966 Chondridiopisalia Lohse, 1974: 53 [ Newton & Thayer 2003 ] Chondropisalia Scheerpeltz, 1966b: 14 Type species: Leptusa schaschli Ganglbauer, 1897 Cidnopisalia Pace, 1989: 235 Type species: Leptusa cavallensis Pace, 1980 Comphopisalia Scheerpeltz, 1966b: 16 Type species: Leptusa brucki Scriba, 1867 Craspedopisalia Scheerpeltz, 1966b: 19 Type species: Leptusa carinata Bernhauer, 1909 Cryptopisalia Pace, 1983c: 88 Type species: Leptusa montisparii Scheerpeltz, 1953 Cryptopisalia Pace, 1983e: 129 [ Newton & Thayer 2003 ] Cyllopisalia Pace, 1982a: 40 Type species: Leptusa difformis Mulsant & Rey, 1853 Cyllopisalia Pace, 1983c: 79 [ Smetana 2004: 431 ] Type species: Leptusa difformis Mulsant & Rey, 1853 Parapisalia Scheerpeltz, 1966b: 18 [synonymized by Pace 1982a: 40 ] Type species: Leptusa difformis Mulsant & Rey, 1853 Sipalia Mulsant & Rey, 1853b: 32 [synonymized by Pace 1982a: 40 ] Type species: Homalota difformis Mulsant & Rey, 1853 Dendroleptusa Pace, 1983c: 65 Type species: Leptusa fumida Erichson, 1839 Drepanoleptusa Pace, 1982d: 93 Type species: Leptusa annapurnensis Pace, 1982 Dysleptusa Pace, 1982c: 580 Type species: Oxypoda fuliginosa Aubé, 1850 Ectinopisalia Scheerpeltz, 1966b: 18 Type species: Leptusa baldensis Ganglbauer, 1895 Emmelopisalia Pace, 1983c: 92 Type species: Leptusa norica Pace, 1979 Entomophallopisalia Pace, 1983c: 90 Type species: Leptusa asturiensis Eppelsheim, 1880 Eospisalia Pace, 1982c: 583 Type species: Leptusa ishizuchiensis Pace, 1982 Eremopisalia Pace, 1989: 196 Type species: Leptusa marianii Pace, 1978 Eucryptusa Casey, 1906: 345 Type species: Silusa nanula Casey, 1893 Evaniopisalia Pace, 1982a: 38 Type species: Leptusa ultracollensis Pace, 1982 Exaeretopisalia Pace, 1989: 233 Type species: Leptusa cansigliensis Pace, 1975 Gallopisalia Pace, 1989: 179 Type species: Leptusa lapidicola Brisout de Barneville, 1863 Gatineauleptusa Klimaszewski in Klimaszewski et al ., 2018: 363 Type species: Leptusa gatineauensis Klimaszewski & Pelletier in Klimaszewski et al ., 2004 Geopisalia Pace, 1984: 214 Type species: Leptusa peyerimhoffi Bernhauer, 1935 Gnopheropisalia Pace, 1983c: 76 Type species: Leptusa granulipennis Eppelsheim, 1880 Habropisalia Pace, 1989: 216 Type species: Leptusa laticeps Scheerpeltz, 1972 Halmaeusa Kiesenwetter in Kiesenwetter & Kirsch, 1877: 160 [*] Type species: Halmaeusa antarctica Kiesenwetter, 1877 Antarctophytosus Enderlein, 1909: 377 [synonymized by Steel 1964: 366 ] Type species: Phytosus atriceps Waterhouse, 1875 Paraphytosus Cameron, 1917c: 125 [synonymized by Steel 1964: 366 ] Type species: Phytosus atriceps Waterhouse, 1875 Austromalota Brèthes, 1925: 170 [synonymized by Steel 1964: 366 ] Type species: Austromalota rufimixta Brèthes, 1925 Hemipasilia Pace, 1989: 90 Type species: Leptusa oertzeni Eppelsheim, 1888 Heteroleptusa Pace, 1989: 160 Type species: Sipalia frontalis Casey, 1893 Heterotyphlopasilia Pace, 1983c: 66 Type species: Leptusa serbica Scheerpeltz, 1935 Homopisalia Pace, 1982c: 584 Type species: Leptusa bengalensis Pace, 1982 Ischnopisalia Pace, 1989: 226 Type species: Leptusa poggii Pace, 1978 Kochliodepisalia Pace, 1996a: 35 Type species: Leptusa spirarum Pace, 1996 Koilokoryphepisalia Pace, 1999a: 210 Type species: Leptusa argentierensis Pace, 1999 Lasiopisalia Pace, 1983c: 76 Type species: Leptusa crenulata Bernhauer, 1899 Lathripisalia Pace, 1983e: 135 Type species: Leptusa montispasubii Pace, 1975 Leptopasilia Scheerpeltz, 1966b: 13 Type species: Leptusa hopffgarteni Eppelsheim, 1881 Leptusa Kraatz, 1856b: 60 Type species: Bolitochara pulchella Mannerheim, 1830 Sipalia Mulsant & Rey, 1853a: 45 [ Newton & Thayer 2003 ] Liopisalia Pace, 1989: 177 Type species: Leptusa serullazi Sainte-Claire Deville, 1919 Megacolypisalia Pace, 1983e: 133 Type species: Leptusa laevicauda Scheerpeltz, 1958 Melopisalia Pace, 1983c: 88 Type species: Leptusa pasubiana Scheerpeltz, 1953 Metaplasmopasilia Pace, 1989: 104 Type species: Leptusa apennina Holdhaus, 1924 Microcolypisalia Pace, 1989: 196 Type species: Leptusa occulta Pace, 1975 Micropisalia Scheerpeltz, 1948: 164 Type species: Leptusa alpestris Scheerpeltz, 1935 Myopopisalia Scheerpeltz, 1966b: 14 Type species: Leptusa bonvouloiri Brisout de Barneville, 1867 Nanoglossa Fauvel, 1868a: 379 [*] Type species: Microglossa chilensis Fauvel, 1866 Microglossa Fauvel, 1866b: 282 [synonymized by Fauvel 1868a: 379 ] Type species: Microglossa chilensis Fauvel, 1866 Microglossa Fauvel, 1866a: 33 [ Newton & Thayer 2003 ] Type species: Microglossa chilensis Fauvel, 1866 Pareuryusa Scheerpeltz, 1972d: 131 [synonymized by Pace 1989: 110 ] Type species: Euryusa parallela Fairmaire & Germain, 1862 Nanopisalia Pace, 1983c: 94 Type species: Homalota piceata Mulsant & Rey, 1853 Micropisalia Scheerpeltz, 1966b: 20 [synonymized by Pace 1983c: 94 ] Type species: Homalota piceata Mulsant & Rey, 1853 Nanopisalia Pace, 1983e: 132 [ Newton & Thayer 2003 ] Type species: Homalota piceata Mulsant & Rey, 1853 Neopisalia Scheerpeltz, 1966b: 16 Type species: Leptusa microphthalma Reitter, 1887 Nesopisalia Pace, 1992b: 112 Type species: Leptusa centralis Pace, 1992 Stenoleptusa Scheerpeltz, 1966b: 10 [synonymized by Assing 2009c: 1286 ] Type species: Bolitochara laeviuscula Hochhuth, 1849 Oligopisalia Scheerpeltz, 1966b: 20 Type species: Leptusa flavicornis Brancsik, 1874 Oncopisalia Pace, 1982c: 589 Type species: Leptusa monachorum Bernhauer, 1912 Oreopisalia Pace, 1983c: 82 Type species: Leptusa hoelzeli Scheerpeltz, 1948 Pachygastropisalia Scheerpeltz, 1966b: 15 Type species: Leptusa lativentris Sharp, 1873 Pachygluta Thomson, 1858: 34 Type species: Oxypoda ruficollis Erichson, 1839 Parapisalia Scheerpeltz, 1948: 159 Type species: Leptusa puellaris Hampe, 1863 Pasilia Mulsant & Rey, 1871: 226 Type species: Homalota nubigena Kiesenwetter, 1861 Pasilia Mulsant & Rey, 1872b: 316 [ Newton & Thayer 2003 ] Pasilia Mulsant & Rey, 1873c: 73 [ Newton & Thayer 2003 ] Pisalia Mulsant & Rey, 1871: 234 Type species: Homalota globulicollis Mulsant & Rey, 1853 Pisalia Mulsant & Rey, 1872b: 324 [ Newton & Thayer 2003 ] Pisalia Mulsant & Rey, 1873c: 73 [ Newton & Thayer 2003 ] Tropidiopasilia Scheerpeltz, 1966b: 11 [synonymized by Pace 1989: 190 ] Type species: Leptusa tricolor Scriba, 1870 Planepisalia Pace, 1989: 217 Type species: Leptusa kochiana Bernhauer, 1936 Protoleptusa Semenov in Semenov et al. , 2001: 156 Type species: Leptusa defuncta Semenov, Perkovsky & Petrenko, 2001 Rhombopisalia Scheerpeltz, 1953: 66 Type species: Leptusa knabli Bernhauer, 1909 Roubaliusa Scheerpeltz, 1966b: 13 Type species: Leptusa storkani Roubal, 1916 Scelopisalia Scheerpeltz, 1966b: 18 Type species: Leptusa gracilipes Krauss, 1899 Scoliophallopisalia Pace, 1989: 232 Type species: Leptusa pascuorum Pace, 1975 Scoliophallopisalia Pace, 1981: 115 [ nomen nudum , Newton & Thayer 2003 ] Scoliophallopisalia Anonymous, 1984: 249 [ Newton & Thayer 2003 ] Type species: Leptusa blesioi Pace, 1981 Scolioplatypisalia Pace, 1989: 86 Type species: Leptusa munelensis Apfelbeck, 1907 Stictopisalia Scheerpeltz, 1966b: 16 Type species: Leptusa eximia Kraatz, 1856 Synpisalia Scheerpeltz, 1966b: 16 [synonymized by Pace 1983c: 56 ] Type species: Leptusa caucasica Eppelsheim, 1877 Toxophallopisalia Pace, 1983c: 99 Type species: Leptusa austriaca Scheerpeltz, 1935 Trichopasilia Scheerpeltz, 1966b: 12 Type species: Leptusa major Bernhauer, 1900 Typhlopasilia Ganglbauer, 1895: 278 Type species: Leptusa kaufmanni Ganglbauer, 1895 Ulitusa Casey, 1906: 347 Type species: Ulitusa cribratula Casey, 1906 Yunnaleptusa Assing, 2008b: 256 Type species: Leptusa curvata Assing, 2006 16 species are included in this genus from South America. Characters/Description: Lynch-Arribálzaga 1884: 87 (characters); Fenyes 1920: 120 (characters), 122 (characters subgenera Typhlosilia and Leptusa ), 123 (characters subgenera Eucryptusa and Pachygluta ), 124 (characters subgenera Oreusa and Pasilia ), 125 (characters subgenera Pisalia ); Pace 1989: 25 (characters), 110 (characters of the subgenus Nanoglossa ), 244 (characters of the subgenus Halmaeusa ). Key: Bernhauer 1900a: 402 (Subgenera and some species); Fenyes 1920: 121 (subgenera); Pace 1989: 25 (subgenera), 110 (species of the subgenus Nanoglossa ), 246 (species of the subgenus Halmaeusa ); Pace 2001a: 12 (Chilean species). Distribution. South America: Chile , Argentina , Falkland Isl. Other regions: Nearctic, Palaearctic, Ethiopian, Oriental , Australian, Antarctic. 0448. Meoticaphaena Pace, 2015e: 72 Type species: Meotica cordobensis Bernhauer, 1934 Only one species is known from and is restricted to South America. Distribution. South America: Argentina . 0449. Phymatura J. Sahlberg, 1876: 85 Type species: Bolitochara brevicollis Kraatz, 1856 Venusa Casey, 1906: 272 [synonymized by Ashe 1992: 362 ] Type species: Venusa picta Casey, 1906 3 species are included in this genus from South America. Characters/Description: Fenyes 1920: 116 (characters); Ashe 1992: 366 (characters in key), 371 (characters). Distribution. South America: Brazil , Argentina . Other regions: Nearctic, Palaearctic, Ethiopian, Madagascan, Oriental , Australian. 0450. Prosoponotha Pace, 1999c: 146 Type species: Prosoponotha chilensis Pace, 1999 5 species are known from and are restricted to South America. Distribution. South America: Brazil , Chile . 0451. Tanystetha Pace, 2002: 137 Type species: Tanystetha brasiliensis Pace, 2002 Only one species is known from and is restricted to South America. Distribution. South America: Brazil . Subtribe Dinardopsina Bernhauer & Scheerpeltz 1926 0452. Dinardopsis Bruch, 1917: 257 ( Fig. 52 ) Type species: Dinardopsis solenopsidicola Bruch, 1917 2 species are known from and are restricted to South America. Distribution. South America: Brazil , Argentina .