Five new ocellus-bearing species of the cusk-eel genus Neobythites (Ophidiidae, Ophidiiformes) from the West Pacific, with establishment of three new species groups
Author
Uiblein, Franz
Institute of Marine Research, P. O. Box 1870 Nordnes, N- 5817 Bergen, Norway. Corresponding author; & National Research Foundation - South African Institute for Aquatic Biodiversity, Makhanda / Grahamstown, South Africa & Vietnam National Museum of Nature, Hanoi, Vietnam
Author
Nielsen, Jørgen G.
Natural History Museum of Denmark, University of Copenhagen, Denmark
text
Zootaxa
2023
2023-08-22
5336
2
179
205
http://dx.doi.org/10.11646/zootaxa.5336.2.2
journal article
10.11646/zootaxa.5336.2.2
1175-5326
8272417
7BD6357E-BB67-44E9-B028-4CA7A2769660
Neobythites ocellatus
Günther, 1887
Figure 4
,
Tables 1
,
3
Neobythites ocellatus
Günther, 1887: 103
, pl. 21 (fig. B) (off
Pernambuco
, northeastern
Brazil
,
09°05′S
,
34°50′W
, 350 fathoms;
holotype
: BMNH 1887.12.7.43).
Diagnosis.
No or one thin and flat spine on hind margin of preopercle; dorsal-fin rays 95–100; anal-fin rays 81–85; pectoral-fin rays 24–27; precaudal vertebrae 12; total vertebrae 56–58; pseudobranchial filaments 3–4; long rakers on anterior gill arch 14–16; head length 21–24% SL; pelvic-fin length 16–22% SL, fins not extending beyond anus; orbit length 5.3–6.1% SL and 22–28% HL; longest gill filament 1.7–2.6% SL and 7.3–8.4% HL; dorsal fin with two ocelli, one smaller ocellus placed closer to fin origin, spot distance 28–30% SL and spot covers 5–7 dorsal-fin rays, and slightly larger central ocellus placed behind line through anus, spot distance 45–50% SL and spot covers 8–11 dorsal-fin rays; both ocellus spots not extending ventrally onto body; preserved specimens with two black ocellus spots; no vertical bars on body; otolith length 5.6–5.8% SL, sulcus length 4.0–4.3% SL, and ostium height 18–21% sulcus length and 28–34% ostium length.
Distribution and size.
SW Atlantic, off
Bahia
, E
Brazil
at
233–641 m
depth. Known up to
148 mm
SL.
Comparisons.
Neobythites japonicus
n. sp.
differs from
N. kenyaensis
and
N. ocellatus
in having two vs. 0 or 1 reduced preopercular spines, fewer dorsal-fin rays, anal-fin rays, and vertebrae, more pseudobranchial filaments, smaller eyes, and shallower otolith ostium height; furthermore, it differs from
N. kenyaensis
in having both ocelli placed more anterior on the dorsal fin and longer head and pelvic fin; and it differs from
N. ocellatus
in having fewer gill rakers and shorter gill filaments.
Neobythites kenyaensis
differs from
N. ocellatus
in having more precaudal vertebrae, fewer gill rakers, shorter upper jaw and pelvic fin, and more posteriorly placed ocelli.
Remarks.
Okamoto
et al.
2011
considered the colour structure close to the dorsal-fin origin to be merely a black blotch. However, our re-examination revealed a clear ring surrounding the spot (
Fig. 4A
).
Neobythites ocellatus
only has posterior spots, but no posterior ocelli (
Nielsen
et al.
2009
)