Five new ocellus-bearing species of the cusk-eel genus Neobythites (Ophidiidae, Ophidiiformes) from the West Pacific, with establishment of three new species groups
Author
Uiblein, Franz
Institute of Marine Research, P. O. Box 1870 Nordnes, N- 5817 Bergen, Norway. Corresponding author; & National Research Foundation - South African Institute for Aquatic Biodiversity, Makhanda / Grahamstown, South Africa & Vietnam National Museum of Nature, Hanoi, Vietnam
Author
Nielsen, Jørgen G.
Natural History Museum of Denmark, University of Copenhagen, Denmark
text
Zootaxa
2023
2023-08-22
5336
2
179
205
http://dx.doi.org/10.11646/zootaxa.5336.2.2
journal article
10.11646/zootaxa.5336.2.2
1175-5326
8272417
7BD6357E-BB67-44E9-B028-4CA7A2769660
The four species groups of
Neobythites
(see also
Table 1
)
-
australiensis
group: species with a single ocellus, two pre-opercular spines, and pelvic fins not reaching anus. Four species belong to this group:
Neobythites superocellatus
n. sp.
,
N. australiensis
,
N. nigriventris
, and
N. unimaculatus
.
-
kenyaensis
group: species with two ocelli, a central ocellus and an ocellus placed close to dorsal-fin origin, with zero, one, or two preopercular spines, and pelvic fins not reaching anus. Three species belong to this group:
Neobythites japonicus
n. sp.
,
N. kenyaensis
, and
N. ocellatus
.
-
longiventralis
group: species with one or two ocelli on dorsal fin, zero, one, or two preopercular spines, and pelvic fins extending beyond anus. Four species belong to this group:
Neobythites jonathan
n. sp.
,
N. zora
n. sp.
,
N. longiventralis
, and
N. longipes
. The latter species was moved to this group after having originally been included in the
steatiticus
group (
Uiblein & Nielsen 2018
).
-
steatiticus
group: species with single ocellus, zero or one pre-opercular spine, and pelvic fins not reaching anus. This group includes nine species:
Neobythites pako
n. sp.
,
N. gloriae
Uiblein & Nielsen
,
N. lombokensis
Uiblein & Nielsen
,
N. malayanus
Weber
,
N. malhaensis
Nielsen
,
N. meteori
Nielsen
,
N. monocellatus
,
N. steatiticus
Alcock
, and
N. stefanovi
Nielsen & Uiblein.
Key to the four species groups
1
One or several ocelli on dorsal fin........................................................................ 2
-
No ocelli on dorsal fin...... 30 non-ocellus bearing species (28 species listed in
Uiblein & Nielsen 2005
, plus
N. machidai
Ohashi, Nielsen & Yabe
and
N. solomonensis
Uiblein & Nielsen
)
2
A central ocellus with no posterior ocellus on dorsal fin or with a posterior ocellus and pelvic fins extending beyond anus.. 3
-
At least one posterior ocellus and pelvic fins not extending beyond anus................. 10 ungrouped species (
Table 1
)
3
Pelvic fins not extending beyond anus.................................................................... 4
-
Pelvic fins extending beyond anus.................................................
longiventralis
species group
4
A central ocellus and no anterior or posterior ocellus......................................................... 5
-
A central ocellus and an anterior ocellus, no posterior ocellus...............................
kenyaensis
species group
5
2 preopercular spines.............................................................
australiensis
species group
-
0–1 preopercular spines.............................................................
steatiticus
species group