Prodromus of a fern flora of Bolivia. XLII. Update I. Author Kessler, Michael Systematic and Evolutionary Botany, University of Zurich, Zollikerstrasse 107, CH- 8008 Zurich, Switzerland. Author Smith, Alan R. University Herbarium, 1001 Valley Life Sciences Bldg. # 2465, University of California, Berkeley, CA 94720 - 2465. Author Øllgaard, Benjamin Department of Biology - Ecoinformatics and Biodiversity, Ny Munkegade 116, building 1137, 8000 Aarhus C, Denmark. Author Matos, Fernando B. Centro de Referência em Informação Ambiental (CRIA), Rua João Carlos do Amaral 500, 13070 - 111, Campinas, SP, Brasil. Author Moran, Robbin C. New York Botanical Garden, 2900 Southern Blvd., Bronx, New York, 10458 - 5126, USA. text Phytotaxa 2023 2023-12-20 630 3 183 210 https://phytotaxa.mapress.com/pt/article/download/phytotaxa.630.3.2/51376 journal article 10.11646/phytotaxa.630.3.2 1179-3163 10409920 Hiya Schwartsburd et al . (2017) treated 12 species of Hypolepis for Bolivia , including Hypolepis nigrescens Hook. More recently, Shang et al . (2018) conducted a phylogenetic analysis of Hypolepis and related genera and found that three species placed in Hypolepis ( H. brooksiae , H. nigrescens , and H. scabristipes ) are more closely related to the genera Blotiella , Paesia , and Histiopteris than to the remainder of Hypolepis . They therefore created the new genus Hiya to accommodate these three species. Hiya is characterized by indeterminate, scandent leaves exhibiting rhythmic growth, provided with recurved black-tipped prickles, and stipule-like pinnules that protect the emerging crozier and young pinna apices, rachis-costa architecture where the adaxial sulcus is confluent with the next lower order, and a base chromosome number of x = 29 ( Shang et al . 2018 ). The name for what we called Hypolepis nigrescens thus is: