Hydroids of the family Halopterididae (Cnidaria, Hydrozoa) collected in the western Pacific by various French expeditions
Author
Agís, José Ansín
Facultade de Ciencias do Mar, Universidade de Vigo, Departamento de Ecoloxía e Bioloxía Animal, Campus Lagoas-Marcosende, E- 36310 Vigo (Spain) agis @ uvigo. es
agis@uvigo.es
Author
Vervoort, Willem
Nationaal Natuurhistorisch Museum, P. O. Box 9517, NL- 2300 RA Leiden (The Netherlands) vervoort @ naturalis. nnm. nl
Author
Ramil, Fran
Facultade de Ciencias do Mar, Universidade de Vigo, Departamento de Ecoloxía e Bioloxía Animal, Campus Lagoas-Marcosende, E- 36310 Vigo (Spain) framil @ uvigo. es
framil@uvigo.es
text
Zoosystema
2009
2009-03-31
31
1
33
61
http://www.bioone.org/doi/abs/10.5252/z2009n1a3
journal article
10.5252/z2009n1a3
1638-9387
5398547
Halopteris infundibulum
Vervoort, 1966
(
Fig. 8
;
Table 8
)
TABLE 8. — Measurements of
Halopteris infundibulum
Vervoort, 1966
in μm. * Gonothecae are still immature.
Halopteris infundibulum
Vervoort, 1966:133
, fig.36a-d.—
Schuchert 1997: 121
, fig. 44a-d. —
Vervoort & Watson 2003: 359
, fig. 87C-J.
MATERIAL
EXAMINED. —
New Caledonia
.
BIOCAL 1, stn DW 36, 23°08.647’-
23°08.900’S
, 167°10.994’-
167°11.296’E
,
650-680 m
,
29.VIII.1985
, 1 colony with
2 immature
gonothecae (
MNHN
).
DISTRIBUTION. — This species is known from the
Tasman
Sea (
42°10’S
,
170°10’E
) at
610 m
depth (
Vervoort 1966
), and from
New Zealand
waters south of
43°S
, depth
512-1006 m
. Our material comes from a single station in the
New Caledonia
area, collected between 650 and
680 m
depth.
DESCRIPTION
Colony with a hydrorhiza composed of perisarcal fibres attached to substratum, from which rises a monosiphonic hydrocaulus. Nevertheless, basal part of colony shows several secondary tubes.
Hydrocaulus unbranched, with basal part divided into ahydrothecate internodes by transverse nodes as the result of regeneration processes after damage; last node with 14 frontal nematothecae in two longitudinal rows, separated from remain- der of stem by a hinge-joint. First hydrothecate internode, delimited by hinge-joints, bearing 10 nematothecae: one infracalycine, two pairs of laterals and five supracalycine in two rows; without lateral apophyses. Remainder of stem with hydrothecae, nematothecae and lateral apophyses but not divided into internodes. Cauline hydrothecae not strictly in one line, but slightly displaced towards hydrocladial apophyses, not as deep as hydrocladial hydrothecae and flanked by two pairs of lateral nematothecae; between two consecutive hydrothecae there are two mesial nematothecae. Apophyses short, without nematothecae, placed besides cauline hydrothecae and alternately directed left and right, except in second cauline hydrotheca where they are opposite
Hydrocladia inserted on lateral apophyses next to cauline hydrothecae. First hydrocladial internode short, ahydrothecate, without nematothecae, with basal transverse and distal oblique node. Remainder of hydrocladium unsegmented; only a weak trans-
BIOCAL 1 stn DW 36
Height of colony (in mm) 30
Axial segments:
Length
Diameter at node 220-380
Cauline hydrotheca:
Length 180-210
Diameter at rim 220-230
Hydrocladial internode:
Length 760-940
Diameter at node 90-150
Hydrotheca:
Length abcauline wall 330-370
Length adnate part adcauline wall 230-270
Length free portion adcauline wall 100-130
Diameter at rim 280-310
Lateral nematotheca (long pair):
Length 200-270
Diameter at rim 50-55
Lateral nematotheca (short pair):
Length 80-110
Diameter at rim 50-60
Gonotheca*:
Length 420-450
Maximum diameter 220-240
verse node above a hydrotheca can occasionally be found, usually in distal parts of hydrocladia. Each hydrocladial internode, when visible, with one hydrotheca and six nematothecae: two median inferior and two pairs of lateral nematothecae. Hydrothecae almost cylindrical, deep, with ¾ of adcauline wall adnate and abcauline wall almost straight. Mesial inferior nematothecae with apical chamber (slightly) lowered on adcauline side. First pair of lateral nematothecae on well-developed apophyses, long, slightly curved and surpassing the hydrothecal rim; basal chamber much longer than distal one, rim of apical chamber slightly lowered at side close to hydrotheca. Second pair of lateral nematothecae inserted on base of apophyses, small and with the rim of apical chamber deeply scooped. All nematothecae are bithalamic and movable.
Two immature gonothecae inserted at hydrothecal base by one-segmented pedicel, cone-shaped, and with one or two nematothecae on basal part; apical chamber of nematothecae lowered at side close to gonotheca.
FIG. 8. —
Halopteris infundibulum
Vervoort, 1966
, BIOCAL 1, stn DW 36:
A
, colony, frontal view;
B
, fragment of colony with immature gonotheca, latero-frontal view;
C
, basal internodes of stem, latero-frontal view;
D
, internodes from distal part of hydrocladium, lateral view;
E
, internodes, frontal view;
F
, immature gonotheca. Scale bars: A, 1 cm; B-E, 0.2 mm; F, 0.1 mm.
VARIABILITY
Once three nematothecae were observed between two successive axial hydrothecae.
REMARKS
Material from
New Caledonia
shows the first axial internode limited by oblique nodes (hinge-joints),but it agrees with the
type
material of
Halopteris infundibulum
in all morphological characters of trophosome, gonosome and also with the measurements.