Revision of genus-group taxa in the family Scorpiopidae Kraepelin, 1905, with description of 15 new species (Arachnida Scorpiones) Author Kovařík, František Author Lowe, Graeme Author Stockmann, Mark Author Šťáhlavský, František text Euscorpius 2020 325 1 140 journal article 10.5281/zenodo.5741842 1536-9307 5741842 DCAC2354-0168-4A66-AC36-87F1BB19EAA2 Scorpiops solegladi sp . n . ( Figures 129 , 729–753 , 799 , Tables 7 , 9) http : //zoobank . org / urn : lsid : zoobank . org : act : 1272AAE1- 35F4-4BAE-96F8-DD47ECB61D9B TYPE LOCALITY AND TYPE REPOSITORY . Vietnam , Lào Cai Province , ca. 15 km from Sa Pa ( 22.01°N 102.81°E ), 600 m a. s. l. ; FKCP . TYPE MATERIAL ( FKCP ). Vietnam , Lào Cai Province , ca. 15 km from Sa Pa ( 22.01°N 102.81°E ), 600 m a. s. l. III .2010 , 1♀ ( holotype ), leg. Z. Bříza ; Lai Châu Province , Than Uyen ( 21.97°N 103.86°E ) , III .2010 , 1♀ ( paratype ), leg. V . Fura . ETYMOLOGY. The specific epithet honors Michael E. Soleglad ( USA ) for his friendship, contribution to understanding Scorpiopidae , and lifelong dedication to scorpions. DIAGNOSIS ( ♀ ). Total length 32–35 mm . Base color reddish brown to black with yellow spots. Pectine teeth number 4–6 in females, fulcra absent; pectines from one compact unit with an incomplete furrow between areas where marginal and middle lamellae are usually delimited. Patella of pedipalp with 18 (5 eb , 2 esb , 2 em , 4 est , 5 et ) external and 10–11 ventral trichobothria. Chela of pedipalp with 4 V series trichobothria located on ventral surface. Chelal trichobothrium Eb 3 located in middle of manus between trichobothria Dt and Est . Fingers of pedipalps straight in females. Chela length to width ratio 4.1 in female. Pedipalp movable finger with 50–60 IAD which create a second row, parallel with MD ( 67–70 in number) and there are also 4 ID and 12–13 OD present. Tarsomere II of legs with row of 5–7 stout median ventral spinules and two pairs of flanking setae. Telson elongate and very finely granulate, length to depth ratio 3.62 in female; annular ring present. DESCRIPTION ( ♀ ). Total length 32–35 mm . Male unknown. Habitus as shown in Figs. 729–730 . For position and distribution of trichobothria on pedipalps, see Figs. 735–741 . Fingers of pedipalps are straight in females ( Fig. 736 ). Coloration ( Figs. 729–730 ). Base color uniformly reddish black to black with yellow spots. Chelicerae reddish brown and reticulate, anterior part and fingers are black. Carapace and mesosoma ( Figs. 729–730 , 751–753 ). Entire carapace covered with both large and minute granules; carinae absent. Anterior margin of carapace with a deep median notch. Carapace with 3 pairs of lateral eyes of which two are normal and one is reduced. Mesosoma granulated, with one median carina. Tergite VII is pentacarinate. Sternites smooth with two parallel furrows except sternite VII which is finely granulated without carina developed. Pectine teeth number 4–6 in females, fulcra absent. Pectines create one compact unit with an incomplete furrow between areas of marginal and middle lamellae, or with two marginal lamellae indicated. Metasoma and telson ( Figs. 731–734 ). Metasoma sparsely hirsute and granulated, with sparse, minute granules. Metasomal segment I with 10 carinae, II–IV with 8 carinae, and V with 5 carinae. Median lateral carinae of metasoma II and V indicated by isolated granules that may coalesce into carinae. Dorsolateral carinae of the segment IV terminate posteriorly in a pronounced tooth. Telson elongate and very finely granulate with annular ring developed. Pedipalps ( Figs. 735–750 ). Pedipalps very sparsely hirsute. Patella with 18 (5 eb , 2 esb , 2 em , 4 est , 5 et ) external and 10–11 ventral trichobothria. Chela with 4 V series trichobothria located on the ventral surface. Femur and patella are finely granulated. Femur with 5 granulose carinae, and patella with 5 carinae with dorsal and ventral patellar spurs pronounced. Manus dorsally with fine, rounded granules, which in the central area form an irregular dorsal secondary carina. External surface of chela covered by minute granules which indicate an external secondary carina. Movable fingers bear 50–60 IAD which have the same size as MD ( 67–70 in number) and create a second parallel row. There are also 4 ID and 12–13 OD present. Legs . Tibia and tarsomeres of legs with several setae not arranged into bristle combs on dorsal surfaces, but with rows of spinules on dorsolateral surfaces and on legs I – II also on ventrolateral surface. Tarsomere II of legs I–IV with 5–7 stout median ventral spinules and two pairs of flanking setae. Femur with 3–4 and patella 4–5 carinae; both femur and patella granulated. Measurements . See Table 7 . Figures 729–734 : Scorpiops solegladi sp . n . Figures 729–730 , 732–734 . Female holotype, dorsal (729) and ventral (730) views. Metasoma and telson lateral (732), ventral (733), and dorsal (734) views. Figure 731 . Female paratype, telson lateral. Scale bars: 10 mm (729–730, 732–734). Figures 735–753 : Scorpiops solegladi sp . n . , female paratype (735–742, 749–752), and female holotype (743–748, 753). Figures 735–750 . Pedipalp segments. Chela dorsal (735, 743), external (736, 744) and ventral (737, 745) views. Patella dorsal (738, 746), external (739, 747) and ventral (740, 748) views.Femur and trochanter dorsal (741), and ventral (742) views.Movable finger dentition under white (749) and UV light (750).Trichobothrial pattern is indicated by white circles (735–741). Figure 751 . Coxosternal area and sternites. Figures 752–753 . Carapace and tergites I–III (IV).

S. thailandus sp . n .	S. thailandus sp . n .	S. affinis	S. anthracinus
Dimensions (mm)	♂ holotype	♀ paratype	♂ LT , ZMUH	♂ FKCP
Carapace	L / W	4.03 / 3.93	3.90 / 3.92	4.66 / 4.73	13.10 / 13.16
Mesosoma	L	5.90	12.28	9.89	33.38
Tergite VII	L / W	1.24 / 2.52	1.92 / 2.90	1.56 / 3.87	6.64 / 9.07
Metasoma + telson	L	13.40	11.52	18.05	44.24
Segment I	L / W / D	1.39 / 1.54 / 1.26	1.11 / 1.42 / 1.27	1.92 / 2.37 / 2.01	4.03 / 4.99 / 4.03
Segment II	L / W / D	1.51 / 1.40 / 1.27	1.31 / 1.24 / 1.17	2.22 / 2.17 / 1.85	4.71 / 4.45 / 4.12
Segment III	L / W / D	1.78 / 1.28 / 1.25	1.47 / 1.15 / 1.17	2.49 / 2.05 / 1.72	5.07 / 4.22 / 4.01
Segment IV	L / W / D	2.11 / 1.21 / 1.16	1.81 / 1.05 / 1.14	2.77 / 1.86 / 1.67	6.08 / 3.81 / 3.96
Segment V	L / W / D	3.28 / 1.32 / 1.09	2.92 / 1.11 / 1.10	4.18 / 1.65 / 1.55	11.58 / 3.73 / 3.80
Telson	L / W / D	3.33 / 1.41 / 1.19	2.90 / 1.13 / 0.97	4.47 / 1.84 / 1.61	12.77 / 3.80 / 3.59
Pedipalp	L	18.22	14.33	16.91	46.97
Femur	L / W	5.07 / 1.51	3.81 / 1.37	4.13 / 1.85	11.25 / 4.58
Patella	L / W	5.16 / 1.53	3.90 / 1.51	4.44 / 1.80	10.69 / 4.61
Chela	L	7.99	6.62	8.34	25.03
Manus	W / D	2.18 / 1.66	2.06 / 1.58	3.21 / 2.92	8.12 / 6.55
Movable finger	L	3.28	2.93	4.28	13.44
Total	L	23.33	27.70	32.6	90.72

Table 8 . Comparative measurements of adults of Scorpiops thailandus sp . n . , S. affinis Kraepelin, 1898 , and S. anthracinus Simon, 1887 Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D), lectotype (LT). AFFINITIES. The combination of six characters (chela trichobothrium Eb 3 located in distal half of manus between trichobothria Dt and Est ; patella of pedipalp with 7 ventral and 18 external trichobothria; fingers of pedipalps straight in females; pedipalp movable finger with 50–60 IAD; and total length 32–35 mm ) is unique in the entire genus Scorpiops . DISTRIBUTION. Vietnam ( Fig. 799 ).