Revision of the subgenus Phortica (sensu stricto) (Diptera, Drosophilidae) from East Asia, with assessment of species delimitation using DNA barcodes
Author
Huang, Jia
Author
Gong, Lu
Author
Tsaur, Shun-Chern
Author
Zhu, Lin
Author
An, Keying
Author
Chen, Hongwei
text
Zootaxa
2019
2019-09-30
4678
1
1
75
journal article
25341
10.11646/zootaxa.4678.1.1
96c8a18a-e887-4345-b266-d76076e8416b
1175-5326
3468782
91B46647-B441-41F5-BE27-3A67D699788C
Phortica
(
Phortica
)
cardua
(
Okada, 1977
)
(
Figs. 5A, 5B
,
7
)
Amiota
(
Phortica
)
cardua
Okada, 1977: 24
.
Amiota
(
Phortica
)
shillongensis
Singh & Gupta in
Dwivedi
et al
. 1979: 62
.
syn. nov.
Diagnosis.
Male arista without or with 1–3 short ventral branch(es) distally, the longest ventral branch as long as 1/3 the longest dorsal branch (
Fig. 7A
); posterior bands of male abdominal third to fifth tergites connect to lateral margins (
Fig. 5B
); male abdominal sixth tergite narrow laterally (
Fig. 7B
).
Remarks.
P
.
shillongensis
was poorly described by Singh & Gupta in
Dwivedi
et al
. 1979
, which lacks sufficient morphological evidence in supporting it as a different species compared with
P
.
cardua
.
Specimens examined.
CHINA
:
7♂
(
4♂
in
KIZ
,
3♂
in
SCAU
),
Longgong
,
Anshun
,
Guizhou
,
26°05'47''N
,
105°53'08''E
, alt.
750m
,
22–24.vii.2000
, H.W. Chen, J.J. Gao;
19♂
(
7♂
in
KIZ
,
12♂
in
SCAU
),
Tongmu
,
Wuyishan
,
Fujian
,
27°54'33''N
,
117°51'54''E
, alt.
580–800m
,
16–18.viii.2001
, H.W. Chen, M. Nozawa, H. Watabe;
2♂
(
SCAU
),
Shimentai
,
Yingde
,
Guangdong
,
24°25'49''N
,
113°17'50''E
, alt.
630m
,
9.iv.2004
, H.W. Chen;
33♂
,
Jianfengling
,
Ledong
,
Hainan
,
18°41'50''N
,
108°50'04''E
, alt.
700–860m
,
15♂
(
SCAU
) on
20.iv.2004
, H.W.
Chen
,
18♂
(
KIZ
) on 17,18.
iv.2008
,
X.P. Chen
, J.J.
Gao
;
55♂
(
SCAU
),
Nanling
,
Ruyuan
,
Guangdong
,
24°05'22''N
,
113°04'47''E
, alt.
650–1500m
,
19♂
on
1–5.v.2004
, H.W. Chen,
26♂
on
21–24.iv.2005
, H.L. Cao, M.F. Xu,
7♂
on
24.viii.2010
, M.F. Xu,
3♂
on
10.ix.2017
, L. Gong, N.N. Wang;
2♂
(
SCAU
),
Guanshan
,
Yifeng
,
Jiangxi
,
28°01'04''N
,
114°25'02''E
, alt.
580m
, 8,10.
vi.2004
, H.W. Chen, J.J. Gao;
48♂
,
Wuyishan
,
Yanshan
,
Jiangxi
,
27°59'38''N
,
117°49'04''E
, alt.
1200–1400m
,
26♂
(
KIZ
) on
15.vi.2004
,
H.W. Chen
, J.J.
Gao
,
22♂
(
SCAU
) on
3.vi.2005
, H.L.
Cao
;
44♂
,
Jiaoye Park
,
Kunming
,
Yunnan
,
25°04'13''N
,
102°37'13''E
, alt.
2100m
,
20♂
(
KIZ
) on
22.vii.2004
, J.J.
Gao
,
5♂
(
SCAU
) on
19.viii.2006
, J.J.
Gao
,
18♂
(
SCAU
) on
5.viii.2007
, J.J.
Gao
,
1♂
(
SCAU
) on
28.viii.2011
, H.W.
Chen
;
35♂
(
SCAU
),
Samage
,
Weixi
,
Yunnan
,
27°36'20''N
,
99°24'10''E
, alt.
2700–2900m
,
27–29.vii.2004
, H.W. Chen, Y.F. Li;
30♂
,
Grand Canyon
,
Maoershan
,
Xing'an
,
Guangxi
,
25°51'28''N
,
110°25'01''E
, alt.
380–600m
,
4♂
(
SCAU
) on
19.x.2004
,
H.L. Cao
,
H.W. Chen
,
M.F. Xu
, X.L.
Zhang
,
20♂
(
SCAU
) on 21,22.
ix.2006
,
J.J. Jiang
, F.
Zhao
,
6♂
(
KIZ
) on
17–19.iii.2009
, J.J.
Gao
;
8♂
(
SCAU
),
Mangshan
,
Yizhang
,
Hunan
,
24°56'53''N
,
112°54'01''E
, alt.
1200– 1300m
,
25–27.iv.2005
, H.L. Cao, M.F. Xu;
10♂
(
SCAU
),
Baiyunshan
,
Luoyang
,
Henan
,
33°40'19''N
,
111°51'46''E
, alt.
1200m
,
10–12.vii.2005
, J.J. Jiang, T. Li;
5♂
(
SCAU
),
Hongping
,
Dajiuhu
,
Shennongjia
,
Hubei
,
31°40'37''N
,
110°25'28''E
, alt.
1720m
,
27.vii–3.viii.2005
, H.Z. Cao, H.W. Chen, J.J. Jiang, T. Li, M.F. Xu;
30♂
,
Jizushan
, Bin- chuan,
Yunnan
,
25°56'54''N
,
100°23'58''E
, alt.
2100–2500m
,
9♂
(
KIZ
) on
21.viii.2005
, J.J.
Gao
,
16♂
(
SCAU
) on
21–23.vii.2006
,
H.L. Cao
,
H.Z. Cao
, T.
Li
,
5♂
(
KIZ
) on 19,20.
vii.2007
, J.J.
Gao
;
6♂
(
SCAU
),
Huoditang
,
Ningshan
,
Shaanxi
,
33°26'02''N
,
108°26'53''E
, alt.
1500m
,
16–18.vii.2006
, J.J. Jiang, M.F. Xu;
6♂
(
SCAU
), Fop- ing,
Meixian
,
Shaanxi
,
34°07'10''N
,
107°52'39''E
, alt.
1500m
,
18.vii.2006
, J.J. Jiang, M.F. Xu;
3♂
(
SCAU
), Wul- iangshan,
Jingdong
,
Yunnan
,
25.vii.2006
,
H.L. Cao
,
H.Z. Cao
,
T. Li
;
8♂
(
SCAU
),
Atuoling
,
Wuzhishan
,
Hainan
,
18°50'11''N
,
109°30'53''E
, alt.
570m
,
20.iv.2007
, H.Z. Cao, T. Li;
8♂
(
4♂
in
KIZ
,
4♂
in
SCAU
),
Wangtianshu
,
Mengla
,
Yunnan
,
21°36'09''N
,
101°35'06''E
, alt.
660m
,
23–25.iv.2007
, H.W. Chen, J.J. Gao;
4♂
(
2♂
in
KIZ
,
2♂
in
SCAU
), E'meishan,
Leshan
,
Sichuan
,
29°36'14''N
,
103°28'56''E
, alt.
700m
,
7.vii.2007
, H.Z. Cao, J.J. Gao, J.J. Jiang;
5♂
(
SCAU
),
Damingshan
,
Shanglin
,
Guangxi
,
23°41'01''N
,
108°30'33''E
, alt.
900m
,
15.iii.2009
, J.J. Gao;
1♂
(
KIZ
),
Yunmengshan
,
Huairou
,
Beijing
,
40°36'10''N
,
116°47'04''E
, alt.
570m
,
4.vii.2009
, J.J. Gao;
12♂
(
6♂
in
KIZ
,
6♂
in
SCAU
),
Yixiang
,
Pu'er
,
Yunnan
,
22°44'19''N
,
101°07'01''E
, alt.
1400m
,
18.ix.2009
, J.J. Gao;
9♂
(
3♂
in
KIZ
,
3♂
in
SCAU
,
3♂
in
TMNS
),
Motuo
108k,
Xizang
,
29°29'23''N
,
95°26'44''E
, alt.
843m
,
3.x.2010
, L. Wang;
7♂
(
SCAU
),
Longtan Park
,
Ximeng
,
Yunnan
,
22°38'44''N
,
99°35'39''E
, alt.
1170m
,
3♂
on
31.iii.2011
, Z.F. Shao, S.J. Yan,
4♂
on 26,27.
x.2017
, H.W. Chen, L. Gong, B.X. Li;
30♂
(
SCAU
),
Muyiji Park
,
Ximeng
,
Yunnan
,
22°37'15''N
,
99°35'42''E
, alt.
1200m
,
14♂
on
2.iv.2011
, J.M. Lu, Z.F. Shao, Y.R. Su, S.J. Yan,
16♂
on
3.v.2016
, J.
Huang
,
Y.Q. Liu
,
Y.L. Wang
, L.
Zhu
;
42♂
,
Hesong
,
Menghai
,
Yunnan
,
21°50'17''N
,
100°05'48''E
, alt.
1700–1900m
,
16♂
(
SCAU
) on
6–8.iv.2011
,
J.M. Lu
,
Z.F. Shao
, Y.R.
Su
,
26♂
(
KIZ
) on
7–12.v.2012
,
H.W. Chen
, J.J.
Gao
;
3♂
(
SCAU
),
Tianmushan
,
Lin'an
,
Zhejiang
,
30°19'32''N
,
119°26'35''E
, alt.
640m
,
25.vii.2011
, Z.F. Shao, S.J. Yan;
3♂
(
SCAU
),
Sanchahe
,
Xishui
,
Guizhou
,
28°30'03''N
,
106°24'16''E
, alt.
1000m
,
7.viii.2011
, J. Huang, Y.R. Su;
13♂
(
7♂
in
KIZ
,
6♂
in
SCAU
),
Hutiaoxia
,
Diqing
,
Yunnan
,
27°11'24''N
,
100°02'55''E
, alt.
1800m
, 22,23.
viii.2011
, H.W. Chen, J.J. Gao;
3♂
(
SCAU
),
Cangshan
,
Dali
,
Yunnan
,
25°40'22''N
,
100°08'03''E
, alt.
1800m
,
26.viii.2011
, H.W. Chen, J.J. Gao;
6♂
(
SCAU
),
Zhuque Forest
Park,
Xi'an
,
Shaanxi
,
33°54'26''N
,
108°30'50''E
, alt.
1500m
, 12,13.
vii.2012
, J. Huang, S.J. Yan;
7♂
(
SCAU
),
Haoping
,
Meixian
,
Shaanxi
,
34°05'22''N
,
107°42'24''E
, alt.
1200m
, 12,13.
vii.2012
, H.W. Chen, J.J. Gao;
26♂
(
13♂
in
KIZ
,
13♂
in
SCAU
),
Baihualing
,
Baoshan
,
Yunnan
,
25°17'50''N
,
98°48'12''E
, alt.
1400–1700m
,
23–26.viii.2013
, Q.S. Gao, K. Liu;
41♂
(
SCAU
),
Mengdong
,
Cangyuan
,
Yunnan
,
23°10'08''N
,
99°13'52''E
, alt.
1270–1430m
,
40♂
on 6,7.
v.2016
, J.
Huang
,
Y.Q. Liu
,
Y.L. Wang
, L.
Zhu
,
1♂
on
19.iv.2018
, Y. Lin;
1♂
(
SCAU
),
Nabang
,
Yingjiang
,
Yunnan
,
24°39'51''N
,
97°35'31''E
, alt.
700m
,
16.v.2016
, J. Huang;
1♂
(
SCAU
),
Tongbiguan
,
Yingjiang
,
Yunnan
,
24°37'04''N
,
97°40'03''E
,
1313m
,
19.v.2016
, J. Huang;
8♂
(
SCAU
),
Langping
,
Tianlin
,
Guangxi
,
24°27'43''N
,
106°22'52''E
, alt.
1490m
,
30.vii.2016
, H.W. Chen, L. Gong, Y.Q. Liu;
2♂
(
SCAU
),
Qimaba
,
Lüchun
,
Yunnan
,
22°49'08''N
,
102°17'25''E
, alt.
1280m
,
3.viii.2016
, H.W. Chen;
1♂
(
SCAU
), Mengyu- an,
Mengla
,
Yunnan
,
21°47'29''N
,
101°22'59''E
, alt.
995m
,
7.viii.2016
, H.W. Chen;
2♂
(
SCAU
),
Nanling
(Shang- songshan),
Lancang
,
Yunnan
,
22°47'57''N
,
99°55'11''E
, alt.
1870m
,
12.viii.2016
, L. Gong, Y.Q. Liu;
3♂
(
SCAU
),
Wenquan
,
Changning
,
Yunnan
,
24°35'34''N
,
99°55'18''E
, alt.
1530m
,
16.viii.2016
, H.W. Chen, L. Gong, Y.Q. Liu;
1♂
(
SCAU
),
Shengtangshan
,
Jinxiu
,
Guangxi
,
23°58'23''N
,
110°06'45''E
, alt.
1300m
,
31.vii.2017
, H.W. Chen;
1♂
(
SCAU
),
Sipaishan
,
Shuangjiang
,
Yunnan
,
23°28'30''N
,
99°39'54''E
, alt.
1569m
,
29.x.2017
, L. Gong;
1♂
(
SCAU
),
Zhongshan
,
Mangshi
,
Yunnan
,
24°07'49''N
,
98°31'27''E
, alt.
1480m
,
3.xi.2017
, B.X. Li;
1♂
(
SCAU
),
Bangdong
,
Ruili
,
Yunnan
,
24°06'09''N
,
97°50'55''E
, alt.
1370m
,
6.xi.2017
, H.W. Chen, L. Gong;
1♂
(
SCAU
),
Moli Forest
Park,
Ruili
,
Yunnan
,
24°07'05''N
,
97°58'51''E
, alt.
920m
,
8.xi.2017
, H.W. Chen;
2♂
(
SCAU
),
Bangda
,
Mangshi
,
Yunnan
,
24°07'55''N
,
98°13'42''E
, alt.
1710m
,
8.xi.2017
, L. Gong, B.X. Li.
Distribution.
China
(
Beijing
*
,
Henan
,
Shaanxi
,
Zhejiang
*
,
Anhui
,
Hubei
,
Jiangxi
,
Hunan
,
Fujian
,
Taiwan
, Guangdong, Hainan, Guangxi, Guizhou, Yunnan, Xizang
*
),
Vietnam
,
India
(
Meghalaya
).