The family Onychiuridae (Collembola) from karst caves of the Basque biospeleologic district, with description of four new species Author Beruete, Enrique 0000-0002-3240-8946 University of Navarra, Faculty of Sciences, Department of Environmental Biology, University Campus, 31080 Pamplona, Spain. eberuete @ yahoo. es; https: // orcid. org / 0000 - 0002 - 3240 - 8946 eberuete@yahoo.es Author Arbea, Javier I. c / Ría de Solía 3, ch. 39. 39610 El Astillero (Cantabria). Author Baquero, Enrique 0000-0002-3240-8946 University of Navarra, Faculty of Sciences, Department of Environmental Biology, University Campus, 31080 Pamplona, Spain. eberuete @ yahoo. es; https: // orcid. org / 0000 - 0002 - 3240 - 8946 & University of Navarra, Institute of Biodiversity and Environment (BIOMA), University Campus, 31080, Pamplona, Spain * corresponding author: ebaquero @ unav. es; https: // orcid. org / 0000 - 0002 - 2145 - 8606 eberuete@yahoo.es Author Jordana, Rafael 0000-0002-3240-8946 University of Navarra, Faculty of Sciences, Department of Environmental Biology, University Campus, 31080 Pamplona, Spain. eberuete @ yahoo. es; https: // orcid. org / 0000 - 0002 - 3240 - 8946 eberuete@yahoo.es text Zootaxa 2021 2021-09-21 5040 2 151 194 journal article 10.11646/zootaxa.5040.2.1 1175-5326 5531192 CC1289B1-4FF9-4369-A596-9FF260C8F314 Protaphorura prolata ( Gisin, 1956 ) sensu Jordana et al ., 1997 Onychiurus prolatus Gisin, 1956 . Mitt. Schweiz . Entomol. Ges ., 29(4): 340 Onychiurus gisini : Gama, 1964 . Mem. Estud. Mus. Zool. Univ. Coimbra , 292: 174 (non Haybach, 1960) Protaphorura prolata (Gisin) : Salmon, 1964 . Bull. R. Soc. New Zealand , 7(2): 174 Onychiurus cancellatus : Jordana & Beruete, 1983 . Mitt. Schweiz . Entomol. Ges ., 56(3-4): 308 (non Gisin, 1956 ) Studied Material. Loc. 9. 7 ex. , 25.X.1985 ; 5 ex. , 3.XI.1985 ; 1 ex. , 22.V.1989 ; 2 ex. , 13.VIII.1989 ; 2 ex. , 12.X.1989 . Loc. 17. 1 ex. , 15.VI.1986 . Loc. 21. 3 ex. , 12.XII.1987 . Loc. 26. 5 ex. , 27.X.1985 ; 4 ex. , 4.VIII.1987 . Loc. 29. 1 ex. , 8.V.1980 . Troglophilous and euedaphic species that has been found in forest humus of the Basque Country and Sierra de Gredos and in caves of Araba and Navarra ( Jordana & Beruete 1983 as O. cancellatus , Arbea & Jordana 1991 , Jordana et al. 1997 , Beruete 2000a , 2000b , Beruete et al. 2002 ) and from Gipuzkoa ( Galán 1993 as O. cancellatus ). In these caves it appears on the surface of gours and puddles, on highly decomposed plant remains and on insectivore droppings.