A new species of the East Asian endemic subgenus Sadonebria Ledoux et Roux (Coleoptera: Carabidae: Nebria) ground beetles from Japan Author Sasakawa, K. text Russian Entomological Journal 2022 2022-03-31 31 1 10 14 http://dx.doi.org/10.15298/rusentj.31.1.02 journal article 10.15298/rusentj.31.1.02 0132-8069 10920153 Nebria ( Sadonebria ) hakusana Sasakawa , sp.n. Figs 2–7 . TYPE MATERIALS : Holotype , ♂ (KS), Shinmata Pass , Nomuki , Katsuyama , Fukui Prefecture , Japan , 1.vi.2020 , S. Inoue leg.; paratypes , 1♂ (KS), Mt. Sanpôkuzure , ca 1,200 m , Shirakawa City , Gifu Prefecture , Japan , 23–24.vi.2005 , H.Ikeda leg.; 1♂ 2♀♀ (EUM), “KAGA-Hakusan Nakahanba ~ Bettonozoki (1,500~ 1,750 m ) 1 VIII. 1973 M. Miyatake ” [ Nakahanba ( 1,500 m ) ~ Bettonozoki ( 1,500 m ), Mt. Hakusan , Shiramine , Hakusan , Ishikawa Prefecture , Japan ] . DESCRIPTION. External characters ( Figs 2–3 ). Body length (including mandibles): ♂ , 12.5–13.6 mm ; ♀ , 12.6– 12.8 mm . PL / PW : 0.73–0.76; PAW/ PW : 0.73–0.76; PPW/ PW : 0.64–0.68. EL/EW: 1.59–1.71. Habitus slender. Hind wings atrophied. Dorsal surface shiny and almost black; head with a dark-brown patch between eyes; pronotal and elytral margins dark-brown; mouth appendages and antennae reddish to dark brown; legs dark brown except for femora, which are blackish brown. Pronotum cordate and convex; lateral margins reflexed throughout; hind angle acute; laterobasal impression large and deep; median line distinct in the middle, absent near the anterior margin, rudimentary near the posterior margin; surface of central part and near the anterior margin almost smooth; surface near the posterior margins very sparsely punctate; surface of the lateral margins very sparsely punctate and/or shallowly, transversely wrinkled; two marginal setae on each lateral side, anterior setae near the widest pronotal point, and posterior setae near hind angles. Elytra oblong, widest behind the middle; 7–10 setigerous pores on interval 3. Abdominal sterna 4–7 usually with a pair of setae, but in the male from Mt. Sanpôkuzure, additional setae on left side of sternum 4 and on both sides of sternum 7. 4 5 6 Figs 2–9. Nebria ( Sadonebria ) spp.: 2–7 — N. hakusana sp.n. ; 8 — N. kiso ; 9 — N. mikawa ; 2–3 — habitus; 4 — aedeagus; 5–6 — endophallus; 7–9 — pronotum; 2–3, 7–9 — dorsal view; 4–5 — right lateral view; 6 —ventral view; 2, 5–9 — holotype, male; 3 — paratype, female; 4 — paratype, male from Mt. Sanpôkuzure. Abbreviations: da — dorsoapical lobe; db — dorsobasal lobe; go — gonopore; lb — laterobasal lobe; va — ventroapical lobe. Scales: 5.0 mm for 2–3; and 0.5 mm for 4–9. Рис. 2–9. Nebria ( Sadonebria ) spp.: 2–7 — N. hakusana sp.n. ; 8 — N. kiso ; 9 — N. mikawa ; 2–3 — габитус; 4 — Эдеагус; 5–6 — Эндофаллус; 7–9 — переднеспинка; 2–3, 7–9 —сверху; 4–5 — справа; 6 — сниЗу; 2, 5–9 — голотип, самец; 3 — паратип, самка; 4 — паратип, самец с горы СанпокудЗуре. СокраЩениЯ: da — дорсоапикальнаЯ долЯ; db — дорсобаЗальнаЯ долЯ; go — гонопор; lb — латеробаЗальнаЯ долЯ; va — вентропапикальнаЯ долЯ. МасШтаб: 5,0 мм длЯ рис. 2–3; 0,5 мм длЯ рис. 4–9. Male genitalia ( Figs 4–6 ). Aedeagus slender and strongly bent at apical 2/5; ventral contour of subapical part from lateral view arcuate ventrally; apex simple and widely rounded.Endophallus straight, elongate, directed posterodorsally; laterobasal lobes large, hemispherical; lateroapical lobes absent; dorsobasal lobe bifid from lateral view, with the subapex of basal one smaller than that of apical one; dorsoapical lobe widely swollen, and smaller than the laterobasal and dorsobasal lobes; ventroapical lobe barely swollen. Both right and left parameres spatulate, with the former larger than the latter. DIAGNOSIS. Distinguished from N. quinquelobata Sasakawa, 2016 , N. kiso , and N. mikawa , which are relatively closely distributed ( Fig. 1 ), by the following characters: from N. quinquelobata by dark brown legs [versus yellowish brown legs in N. quinquelobata ; Sasakawa, 2016 ], and from N. kiso and N. mikawa by less cordate and less punctate pronotum ( Figs 7–9 ). ETYMOLOGY. Named after the Hakusan Mountains , where the type materials were collected .