A revised phylogenetic classification of the ant subfamily Formicinae (Hymenoptera: Formicidae), with resurrection of the genera Colobopsis and Dinomyrmex Author Ward, Philip S. Author Blaimer, Bonnie B. Author Fisher, Brian L. text Zootaxa 2016 4072 3 343 357 journal article 51665 10.11646/zootaxa.4072.3.4 032c9ceb-f905-482c-a67e-9f70facbb902 1175-5326 264543 A358F7A0-12B8-401D-B6FE-ADBF1469B786 Tribe Camponotini Forel 1878 = Polyrhachidini Ashmead 1905 Genera: Calomyrmex , Camponotus , † Chimaeromyrma , Colobopsis , Dinomyrmex , Echinopla , Opisthopsis , Overbeckia , Polyrhachis , † Pseudocamponotus . Comments. The composition of this tribe remains unchanged, although some generic boundaries have been modified (see below). Overbeckia , not sequenced in this study, is likely a junior synonym of Camponotus ( Bolton 2003 ) . All members of this tribe have a unique, vertically inherited bacterial symbiont, Blochmannia , whose evolutionary history mirrors that of the ants ( Wernegreen et al. 2009 ). Morphologically the workers of Camponotini can be recognized by the combination of distinctive mandibular dentition (5–8 teeth, with the third tooth from apex not reduced in size), antennal insertions well separated from the posterior clypeal margin, and twelve antennal segments ( Bolton 1994 , 2003 ).