An evolutional special case in the lower Orthorrhapha: some attractive fossil flies from the Middle Jurassic of China (Insecta: Diptera: Brachycera) Author Zhang, Kuiyan Author Yang, Ding Author Ren, Dong Author Shih, Chungkun text Zoological Journal of the Linnean Society 2010 2010-02-26 158 3 563 572 http://dx.doi.org/10.1111/j.1096-3642.2009.00552.x journal article 276483 10.1111/j.1096-3642.2009.00552.x 3be82545-1797-46a0-95fc-3e0fc44c9af0 0024-4082 5438122 STRENORHAGIO DEVIATUS ZHANG, YANG & SHIH GEN. ET SP. NOV. ( FIG. 3 ) Etymology: The specific name refers to the Latin ‘d eviatus ’ (abnormal), because of the abnormal position of crossvein r–m. Diagnosis: Hind femur and tibia slender. Crossvein r–m far proximal to beginning of vein R 2+3 ; basal trunk of veins M 1 and M 2 longer than crossvein m–m; vein CuA 1 arising from infall of cells bm and d; mouth of cell sc slightly wider than that of cell r 1 ; mouth of cell m 1 much narrower than that of cell m 2 ; mouth of cell m 3 subequal to that of cell m 1 in width. Holotype : CNU-DIB-NN2007018, an almost complete adult body with wings in dorsal view. Type locality and horizon: Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia , China ; Jiulongshan Formation, the Middle Jurassic (Aalenian–Bajocian). Description: Body length, 14.8 mm ; wing length, 11.0 mm; wing width, 3.3 mm . Body stout. Head elliptical, slightly narrower than thorax. Eyes large, bare; ommatidia clearly visible, with upper area composed of larger facets. Legs pubescent. Hind femur and tibia slender; first tarsomere distinctly longer and wider than following tarsomeres. Wing hyaline; veins thick and strong. Vein Sc ending slightly beyond middle of wing. Veins R 1 and Rs 1 long; vein R 2+3 distinctly and strongly bent upwards at base; vein R 5 ending at wing apex. Crossvein r–m at basal third of cell d, far proximal to beginning of vein R 2+3 ; relative lengths of veins Rs 1 , Rs 2 and Rs 3 about 9.5: 1: 8.5. All branches of vein M long, veins M 1 and M 2 bifurcating distal to crossvein m–m, basal trunk of veins M 1 and M 2 longer than crossvein m–m. Vein CuA 1 arising from infall of cells bm and d; veins CuP and A 2 present. Mouths of cells sc and r 1 wide; mouths of cells r 2+3 and r 4 distinctly narrow. Cell bm about as wide as cell br; cell d slender, far from wing margin. Five posterior cells present and wide open; mouth of cell m 1 much narrower than that of cell m 2 ; mouth of cell m 3 subequal to that of cell m 1 in width, about 1/2 as wide as that of cell cua 1 . Cell cup distinctly open; anal lobe small and narrow. Abdomen cylindrical, robust, and pubescent. Eight segments visible; segment I evidently shortest. Remarks: See remarks for Strenorhagio grimaldi sp. nov.