Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae) Author Johnson, Andrew J. School of Forest Resources and Conservation, University of Florida, Gainesville, FL 32611, andrewjavanjohnson@gmail.com Author Hulcr, Jiri School of Forest Resources and Conservation, University of Florida, Gainesville, FL 32611, Author Knížek, Miloš Department of Forest Protection Service, Forestry and Game Management Research Institute, Strnady, Jíloviště, Praha 5, Zbraslav CZ- 15600, Czechia, Author Atkinson, Thomas H. Texas Natural History Collections, Integrative Biology, University of Texas at Austin, Austin, TX 78712, Author Mandelshtam, Michail Yu. Department of Forest Protection, Wood Science and Game Management, Saint-Petersburg State Forest Technical University named after S. M. Kirov, Institutskii per., 5, 194021 Saint-Petersburg, Russia, Author Smith, Sarah M. Department of Entomology, Michigan State University, East Lansing, MI 48824, Author Cognato, Anthony I. Department of Entomology, Michigan State University, East Lansing, MI 48824, Author Park, Sangwook Research Institute of Forest Insect Diversity, Namyangju 12113, South Korea, Author Li, You School of Forest Resources and Conservation, University of Florida, Gainesville, FL 32611, Author Jordal, Bjarte H. University Museum of Bergen, University of Bergen, P. O. 7800, 5020 Bergen, and text Insect Systematics and Diversity 2020 2020-05-31 4 3 1 1 81 journal article 22148 10.1093/isd/ixaa002 be792f86-9e77-414c-9e2f-767704ff704e 2399-3421 3826789 Coriacephilus Schedl, 1939b: 339 ( Fig. 9 ) Type of genus Stephanoderes coriaceus Eichhoff, 1878 . Figure 7. Phylogeny of Cryphalini , Coriacephilini, Xyloterini , andTrypophloeini, Part 2. Diagnosis Coriacephilus can be separated from other Cryphalini by the combination of cylindrical third tarsal segments, deeply emarginated eye, antennae with sutures and septum, and bifurcating setae on the hypomeron. Female Eye deeply emarginated. Antennae with procurved sutures, the first darker indicating a complete septum. Antennae with four funicular segments. Hypomeron with bifurcating setae in most species. Pronotal disc present and prominent in all species with a mixture of hair-like setae pointing anteriorly. Elytra with flattened interstrial bristles and sparse hair-like interstrial ground vestiture. Proventriculus not examined. Aedeagus not examined. Male Unknown. Distribution All species are known from Southeast Asia. Remarks Five species known. No material was available for the study of internal morphology. All species are known from very few specimens. However, the validity of this genus being separate from Cryphalus and Cosmoderes is strongly supported with genetic evidence, placing Coriacephilus coriaceus as sister to Xyloterini . They are morphologically very similar to Cryphalus , most easily distinguished by the third tarsal segment which is not emarginated in Coriacephilus . A study of the internal morphology, or better genetic resources are needed to provide better diagnostic characters for the reliable determination of this genus. Figure 8. Phylogeny of Ernoporini.Tree is made in RAxML, constrained to the tree by Johnson et al. (2018) . Support values are not included due to artificial inflation by constraints.The constraint tree is indicated by a bold line. Outgroup taxa are omitted from the figure. Name pre- and post-revision are displayed, with changes marked in bold. Color boxes indicate genera.The type species of a genus is marked with underlined text. Type material examined Holotype of Stephanoderes coriaceus Eichhoff, 1878 ( NHMW ); holotype of Coriacephilus exiguus Beaver, 2004 ( QM ); paratype of Coriacephilus exiguus Beaver, 2004 ( BMNH ); holotype of Coriacephilus xyloctonoides Schedl, 1939 ( NHMW ). Included species Coriacephilus coriaceus ( Eichhoff, 1878b: 494 ) ( Stephanoderes ) . = Cryphalus birmanus Eggers, 1925: 153 (syn: Schedl, 1940c ). Coriacephilus cribripennis Schedl, 1943: 40 . Coriacephilus exiguus Beaver, 2004: 57 . Coriacephilus proximus ( Eggers, 1925: 156 ) ( Cryphalus ) . Coriacephilus xyloctonoides Schedl, 1939b: 340 .