Revision of the Bark Beetle Genera Within the Former Cryphalini (Curculionidae: Scolytinae)
Author
Johnson, Andrew J.
School of Forest Resources and Conservation, University of Florida, Gainesville, FL 32611,
andrewjavanjohnson@gmail.com
Author
Hulcr, Jiri
School of Forest Resources and Conservation, University of Florida, Gainesville, FL 32611,
Author
Knížek, Miloš
Department of Forest Protection Service, Forestry and Game Management Research Institute, Strnady, Jíloviště, Praha 5, Zbraslav CZ- 15600, Czechia,
Author
Atkinson, Thomas H.
Texas Natural History Collections, Integrative Biology, University of Texas at Austin, Austin, TX 78712,
Author
Mandelshtam, Michail Yu.
Department of Forest Protection, Wood Science and Game Management, Saint-Petersburg State Forest Technical University named after S. M. Kirov, Institutskii per., 5, 194021 Saint-Petersburg, Russia,
Author
Smith, Sarah M.
Department of Entomology, Michigan State University, East Lansing, MI 48824,
Author
Cognato, Anthony I.
Department of Entomology, Michigan State University, East Lansing, MI 48824,
Author
Park, Sangwook
Research Institute of Forest Insect Diversity, Namyangju 12113, South Korea,
Author
Li, You
School of Forest Resources and Conservation, University of Florida, Gainesville, FL 32611,
Author
Jordal, Bjarte H.
University Museum of Bergen, University of Bergen, P. O. 7800, 5020 Bergen, and
text
Insect Systematics and Diversity
2020
2020-05-31
4
3
1
1
81
journal article
22148
10.1093/isd/ixaa002
be792f86-9e77-414c-9e2f-767704ff704e
2399-3421
3826789
Eidophelus
Eichhoff, 1876: 200
(
Figs. 23–26
)
Synonymy
=
Idophelus
Rye, 1877: 362
(unavailable name).
=
Lepicerus
Eichhoff, 1878a: 388
syn. nov.
=
Scolytogenes
Eichhoff, 1878a: 387
syn. nov.
=
Lepicerus
Eichhoff, 1878b: 501
syn. nov.
=
Scolytogenes
Eichhoff, 1878b: 497
syn. nov.
=
Lepidocerus
Rye, 1880: 103
syn. nov.
=
Cryphalomorphus
Schaufuss, 1891: 12
syn. nov.
=
Letznerella
Reitter, 1913: 68
syn. nov.
=
Ernoporides
Hopkins, 1915: 34
syn. nov.
=
Hypothenoides
Hopkins, 1915: 11
syn. nov.
=
Ptilopodius
Hopkins, 1915: 11
syn. nov.
=
Ernoporicus
Berger, 1917: 242
syn. nov.
=
Neocryphalus
Eggers, 1922: 169
syn. nov.
=
Negritus
Eggers, 1923a: 141
syn. nov.
=
Lepicerinus
Hinton, 1936: 473
syn. nov.
=
Cylindrotomicus
Eggers, 1936c: 633
syn. nov.
(unavailable name). =
Eocryphalus
Kurentsov, 1941: 161
syn. nov.
=
Ernopocerus
Balachowsky, 1949: 211
syn. nov.
(unavailable name). =
Ernoporoides
Balachowsky, 1949: 201
syn. nov.
(unavailable name).
=
Ernopocerus
Wood, 1954: 986
syn. nov.
=
Phellodendrophagus
Krivolutskaya, 1958: 107
.
=
Cryphalophilus
Schedl, 1970c: 358
syn. nov.
=
Xylocryptus
Schedl, 1975a: 352
syn. nov.
=
Cryphalogenes
Wood, 1980: 91
syn. nov.
=
Ernpocerus
Wood, 1982: 858
syn. nov.
(unavailable name). =
Nigritus
Wood and Bright, 1992: 858
syn. nov.
(unavailable name).
Type of genus
Eidophelus imitans
Eichhoff, 1876
Diagnosis
Eidophelus
can be diagnosed from superficially similar genera by the entire eye, by the lateral and basal margins of the pronotum marked by a carina, and by the antennal club which has a partial or absent septum, and by the proventriculus characters.
Female
Frons simple, convex. Eye oval shaped, rarely broadly emarginated. Antennae typically with four funicle segments. Antennal club large and flat, with a septum and sutures sometimes present. Pronotum sometimes with asperities on anterior margin. Pronotal summit rarely pronounced (i.e., protruding higher than most of the disc). Elytra with variable setae, from almost glabrous to dense scale-like or hair-like ground vestiture. Apex of elytra nearly or distinctly vertical. Mesocoxae separated, often much more than metacoxae. Metatibiae with denticles distributed over at least apical quarter. Proventriculus with apical plate shorter than rest of proventriculus (except
Eidophelus darwini
), with rows of granules, occasionally less sclerotized around the median suture (i.e., appearing open). Crop with thick spines or granules, in sclerotized clusters.
Male
Typically indistinguishable externally from the female. Some members of this genus have a sexually dimorphic frons (Johnson, unpublished), highly enlarged mandibles, or males have featherlike setae on the protarsi. Penis apodemes are much shorter than penis body, fused at apex. Tegmen open dorsally, of a similar thickness to penis apodemes. Median tegminal apodeme present, typically hooked at apex. Spiculum gastrale of a similar thickness or thicker than penis apodemes, with a fork. Basal sclerites visible.
Distribution
Worldwide except
Antarctica
, but most diversity is in Africa, Asia, Oceania, and Europe.
Remarks
In total, 152 species known. An extraordinarily diverse genus. The morphological variation in this genus has led to the description of many genera based on a limited number of characters, particularly in the antennae. However, there are no distinct groupings based on morphological characters, and many of the current genera are polyphyletic or represent smaller clades nested within the genus. The genera
Cryphalogenes
,
Ernoporicus
,
Ptilopodius
, and
Scolytogenes
are, therefore, all considered synonyms of
Eidophelus
.
Type material examined
Holotype
of
Ernoporicus alniphagus
Nobuchi, 1975
(
ITLJ
)
;
holotype
of
Ernoporicus ater
Nobuchi, 1975
(
ITLJ
)
;
holotype
and
paratype
of
Ptilopodius bambusae
Browne, 1983
(
BMNH
)
;
2 paratypes
of
Cryphalomorphus bangensis
Eggers, 1927
(
NHMW
)
;
holotype
of
Cryphalomorphus brimblecombei
Schedl, 1972
(
QM
)
;
holotype
of
Euptilius papuanus
Browne, 1983
(
BMNH
)
;
holotype
of
Cryphalomorphus buruensis
Eggers, 1926
(
NHMW
)
;
holotype
of
Cryphalomorphus camelliae
Nobuchi, 1975
(
ITLJ
)
;
holotype
of
Cryphalomorphus candidus
Nobuchi, 1975
(
ITLJ
)
;
holotype
(?) of
Lepicerinus coccotrypanoides
Schedl, 1939
(
NHMW
)
;
syntype
(?) of
Lepicerinus coccotrypanoides
Schedl, 1939
(
BMNH
)
;
holotype
of
Cryphalophilus concentralis
Schedl, 1975
(
NHMW
)
;
holotype
of
Margadillius concentralis
Schedl, 1975
(
NHMW
)
;
holotype
of
Cryphalomorphus confragosus
Sampson, 1914
(
BMNH
)
;
holotype
of
Cryphalomorphus scolytomimoides
Nobuchi, 1975
(
ITLJ
)
;
paratype
of
Ptilopodius dubiosus
Wood, 1960
(
USNM
)
;
holotype
and
paratype
of
Cryphalogenes euphorbiae
Wood, 1980
(
USNM
)
;
holotype
and
paratype
of
Cryphalogenes exiguus
Wood, 1980
(
USNM
)
;
holotype
[potentially mislabeled or misplaced] of
Hypothenemus expers
Blandford, 1894
(
BMNH
)
;
holotype
of
Lepicerinus fijianus
Schedl, 1950
(
NHMW
)
;
holotype
of
Ptilopodius formosanus
Browne, 1981
(
BMNH
)
;
holotype
of
Cryphalomorphus fujisanus
Nobuchi, 1975
(
ITLJ
)
;
holotype
of
Cryphalomorphus fulgidus
Schedl, 1975
(
NHMW
)
;
holotype
of
Cryphalomorphus fulvipennis
Nobuchi, 1975
(
ITLJ
)
;
holotype
of
Cryphalophilus ater
Schedl, 1971
(
QM
)
;
paratype
of
Cryphalomorphus granulatus
Wood, 1960
(
USNM
)
;
holotype
of
Cryphalomorphus grossepunctatus
Browne, 1974
(
BMNH
)
;
holotype
of
Cryphalomorphus hirtus
Wood, 1974
(
USNM
)
;
holotype
of
Cryphalomorphus hylesinopsis
Schedl, 1975
(
NHMW
)
;
holotype
of
Scolytogenes indicus
Wood, 1989
(
USNM
)
;
holotype
of
Hypothenemus inermis
Browne, 1984
(
BMNH
)
;
holotype
of
Hypothenemus insignis
Browne, 1984
(
BMNH
)
;
holotype
of
Cryphalomorphus insularis
Nobuchi, 1975
(
ITLJ
)
;
holotype
of
Ericryphalus elongatus
Nobuchi, 1975
(
ITLJ
)
;
lectotype
of
Ptilopodius javanus
Schedl, 1942
(
NHMW
)
;
paratype
of
Hemicryphalus kinabaluensis
Bright, 1992
(
MK
collection)
;
holotype
of
Cryphalomorphus leprosulus
Browne, 1974
(
BMNH
)
;
holotype
of
Cryphalomorphus longipennis
Eggers, 1936
(
BMNH
)
;
paratype
of
Hemicryphalus minutus
Bright, 1992
(
MK
collection)
;
paratype
of
Cryphalomorphus nanulus
Wood, 1960
(
USNM
)
;
Two
paratypes
of
Cryphalomorphus nubilus
Wood, 1960
(
USNM
)
;
Likely unlabeled
holotype
of
Ptilopodius pacificus
Schedl, 1941
(
BMNH
)
;
holotype
of
Cryphalus parvus
Browne, 1984
(
BMNH
)
;
paratype
of
Hemicryphalus podocarpi
Bright, 1992
(
MK
collection)
;
holotype
of
Ptilopodius praeda
Browne, 1978
(
BMNH
)
;
holotype
of
Scolytogenes punctatulus
Nobuchi, 1976
(
ITLJ
)
;
holotype
and
paratype
of
Hypothenemus quadridens
Browne, 1983
(
BMNH
)
;
holotype
and
paratype
of
Cryphalomorphus rusticus
Wood, 1974
(
USNM
)
;
holotype
and
paratype
of
Cryphalomorphus setifer
Wood, 1974
(
USNM
)
;
holotype
of
Ptilopodius spirostachius
Schedl, 1958
(
BMNH
)
;
holotype
of
Lepicerinus squamosus
Schedl, 1942
(
NHMW
)
;
paratype
of
Hemicryphalus squamosus
Bright, 1992
(
MK
collection)
;
holotype
of
Cylindrotomicus squamulosus
Eggers, 1936
(
BMNH
)
;
holotype
of
Ptilopodius stephegynis
Hopkins, 1915
(
USNM
)
;
holotype
of
Ernoporicus takahashii
Nobuchi, 1975
(
ITLJ
)
;
holotype
and
paratype
of
Cryphalomorphus trucis
Wood, 1974
(
USNM
)
.
Figure 23.
Images of
Eidophelus
spp.: Dorsal and lateral photos of A)
E. darwini
, B)
E. euphorbiae
, C)
E. jalappae
, D)
E. hylesinopsis
, E)
E. puerarae
.
Figure 24.
Images of
Eidophelus
spp.: Dorsal and lateral photos of A)
E. quadridens
, B)
E. fagi
, C)
E. semenovi
, D)
E. spessivtzevi
, E)
E. squamosus
.
Figure 25.
Images of
Eidophelus
spp.:Aedeagus of A)
E. darwini
, B)
E. fagi
, C)
E. incultus
, D)
E. squamosus
, and E)
E. fulvipennis
.
Figure 26.
Images of
Eidophelus
spp.: Proventriculus of A)
E. darwini
, B)
E. fagi
, C)
E. hylesinopsis
, D)
E. jalappae
, E)
E. quadridens
,
and F)
E. spessivtzevi
.
Included species
Eidophelus absonus
(
Schedl, 1975a: 344
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus afer
(
Schedl, 1970c: 359
)
(
Cryphalophilus
)
comb. nov.
[
Scolytogenes
].
Eidophelus africanus
(
Schedl, 1977a: 396
)
(
Stephanorhopalus
)
comb. nov.
[
Ernoporicus
].
Eidophelus aitutakii
(
Beaver and Maddison, 1990: 1367
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus alniphagus
(
Nobuchi, 1975: 43
)
(
Ernoporicus
)
comb. nov.
[
Ernoporicus
].
Eidophelus alstoniae
Johnson
nom. nov.
[
Scolytogenes
].
=
Chiloxylon sumatranus
Schedl, 1970c: 358
syn. nov.
(secondary homonym).
Remarks: A replacement name is proposed for
Chiloxylon sumatranus
Schedl, 1970c
because the name is already occupied by
Eidophelus sumatranus
Schedl, 1961a
. The replacement name is a noun in the genitive case, named after the host plant, described as ‘Pulai logs’ =
Alstonia
sp.
Eidophelus alternans
(
Schedl, 1975a: 345
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus amanicus
(
Eggers, 1919: 239
)
(
Cryphalus
)
comb. nov.
[
Scolytogenes
].
Eidophelus ankius
(
Schedl, 1979c: 96
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus apicalis
(
Schedl, 1971c: 11
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus approximatus
(
Schedl, 1975a: 346
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus aspericollis
(
Eichhoff, 1878a: 388
)
(
Lepicerus
)
comb. nov.
[
Scolytogenes
].
=
Lepicerus aspericollis
Eichhoff, 1878b: 501
syn. nov.
=
Cryphalus stierlini
Eggers, 1911: 121
syn. nov.
(syn:
Eggers, 1929c
).
Eidophelus ater
(
Eggers, 1923a: 142
)
(
Negritus
)
comb. nov.
[
Scolytogenes
].
Eidophelus australis
(
Schedl, 1942c: 175
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
Eidophelus badius
(
Nobuchi, 1975: 44
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus bambusae
(
Browne, 1983b: 69
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus bangensis
(
Eggers, 1927c: 75
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus basilaris
(
Wood, 1960: 30
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus birosimensis
(
Murayama, 1958: 935
)
(
Ernocryphalus
)
comb. nov.
[
Scolytogenes
].
Eidophelus borneensis
Browne, 1984a: 153
.
Eidophelus braderi
(
Browne, 1965: 191
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
=
Cryphalomorphus orientalis
Schedl, 1971c: 11
(syn:
Wood, 1992b
).
Eidophelus brighti
Johnson
nom. nov.
[
Hemicryphalus
].
=
Hemicryphalus minutus
Bright, 1992: 187
syn. nov.
Remarks:
Hemicryphalus minutus
Bright, 1992
is transferred to
Eidophelus
based on the eye shape, and shape of the pronotum. A replacement name for
Hemicryphalus minutus
Bright, 1992
is proposed because the name is already occupied by
Eidophelus minutus
Blandford, 1894b
. The replacement name is a noun in the genitive case, honoring the original author.
Eidophelus brimblecombei
(
Schedl, 1972a: 146
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus brownei
Johnson
nom. nov.
[
Ernoporus
].
=
Euptilius papuanus
Browne, 1983b: 70
syn. nov.
(secondary homonym).
Remarks:
Euptilius papuanus
Browne, 1983b
was previously recognized as a junior synonym of
Ernoporus antennarius
Schedl, 1974
. This was likely a mistake since the two species differ greatly.
Wood (1989)
probably intended to synonymize
Ernoporus antennarius
Schedl, 1974
and
Margadillius papuanus
Schedl, 1973b
. Subsequently, the release from synonymy and the transfer of
Euptilius papuanus
Browne, 1983b
to
Eidophelus
requires a replacement name because the name is already occupied by
Cryphalomorphus papuanus
Schedl, 1974
. The replacement name is a noun in the genitive case, honoring the original author.
Eidophelus buruensis
(
Eggers, 1926a: 300
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus camelliae
(
Nobuchi, 1975: 45
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus candidus
(
Nobuchi, 1975: 47
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus capucinus
(
Schedl, 1971a: 283
)
(
Cryphalops
)
comb. nov.
[
Ernoporicus
].
Eidophelus caucasicus
(
Lindemann, 1877: 373
)
(
Ernoporus
)
comb. nov.
[
Ernoporicus
].
=
Cryphalus schreineri
Eichhoff, 1881: 185
.
Eidophelus ceylonicus
(
Schedl, 1959a: 475
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus cicatricosus
(
Schedl, 1942c: 176
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
Eidophelus coccotrypanoides
(
Schedl, 1939b: 343
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
Eidophelus communis
(
Schaufuss, 1891: 12
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus concentralis
(
Schedl, 1975a: 342
)
(
Cryphalophilus
)
comb. nov.
[
Scolytogenes
].
=
Margadillius concentralis
Schedl, 1975a: 344
syn. nov.
Eidophelus confragosus
(
Sampson, 1914: 386
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus corni
(
Kurentsov, 1941: 162
)
(
Hypothenemus
)
comb. nov.
[
Ernoporicus
].
Eidophelus corpulentus
(
Schedl, 1965a: 54
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus corrugatus
(
Schedl, 1950a: 19
)
(
Stephanorhopalus
)
comb. nov.
[
Scolytogenes
].
Eidophelus creber
(
Schedl, 1975a: 346
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus crenatus
(
Sampson, 1914: 385
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus cylindricus
(
Schedl, 1959a: 476
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus darwini
(
Eichhoff, 1878a: 387
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
=
Scolytogenes darwinii
Eichhoff, 1878b: 497
(syn: Alonso- Zarazaga and Lyal, 2009).
=
Negritus similis
Eggers, 1923a: 142
(syn:
Wood, 1985
).
=
Negritus major
Eggers, 1927c: 69
(syn:
Wood, 1985
).
=
Scolytogenes cryptolepis
Schedl, 1951b: 55
(syn:
Wood, 1985
). =
Cryphalomorphus scolytomimoides
Nobuchi, 1975: 50
.
Eidophelus devius
(
Schedl, 1975c: 280
)
(
Cryphalophilus
)
comb. nov.
[
Scolytogenes
].
Eidophelus dubiosus
(
Wood, 1960: 19
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus eggersi
(
Schedl, 1962e: 490
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
=
Cryphalomorphus minor
Eggers, 1927c: 76
(syn:
Schedl, 1962e
).
Eidophelus euphorbiae
(
Wood, 1980: 91
)
(
Cryphalogenes
)
comb. nov.
[
Cryphalogenes
].
Eidophelus excellens
(
Schedl, 1979c: 97
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus exiguus
(
Wood, 1980: 92
)
(
Cryphalogenes
)
comb. nov.
[
Cryphalogenes
].
Eidophelus exilis
(
Yin, 2001: 327
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus eximius
(
Schedl, 1942d: 9
)
(
Erischidias
)
comb. nov.
[
Ernoporicus
].
Eidophelus expers
(
Blandford, 1894b: 85
)
(
Hypothenemus
)
comb. nov.
[
Scolytogenes
].
Eidophelus fagi
(
Fabricius, 1798: 157
)
(
Apate
)
comb. nov.
[
Ernoporicus
].
=
Bostrichus serratus
Panzer, 1795: 288
.
=
Cryphalus (Ernoporus) thomsoni
Ferrari, 1867: 14
(syn:
Ferrari, 1869
).
Eidophelus fijianus
(
Schedl, 1950b: 42
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
Eidophelus formosanus
(
Browne, 1981: 129
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus fugax
(
Schedl, 1975a: 347
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus fujisanus
(
Nobuchi, 1975: 46
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus fulgens
(
Schedl, 1975a: 348
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus fulgidus
(
Schedl, 1975a: 348
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus fulvipennis
(
Nobuchi, 1975: 47
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus furvus
Johnson
nom. nov.
[
Ptilopodius
].
=
Cryphalophilus ater
Schedl, 1972a: 146
syn. nov.
(secondary homonym).
Remarks: A replacement name is proposed for
Cryphalophilus ater
Schedl, 1972a
because the name is already occupied by
Negritus ater
Eggers, 1923a
. The replacement name is an ajective, maintaining the meaning of the original name, meaning black, dark or obscure.
Eidophelus ghanaensis
(
Schedl, 1977b: 281
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus glabratus
(
Yin, 2001: 330
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus gracilis
(
Schedl, 1950b: 44
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
Eidophelus granulatus
(
Wood, 1960: 31
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus grobleri
(
Schedl, 1962b: 68
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus hirtus
(
Wood, 1974a: 18
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus hobohmi
(
Schedl, 1955c: 215
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus hylesinopsis
(
Schedl, 1975a: 349
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus imitans
Eichhoff, 1876: 150
.
=
Phellodendrophagus elegans
Krivolutskaya, 1958: 150
(syn:
Wood, 1989
).
=
Ptilopodius nitidus
Schedl, 1959a: 475
(syn:
Wood, 1989
).
Eidophelus incultus
(
Yin, 2001: 325
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus indicus
(
Wood, 1989: 184
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus inermis
(
Browne, 1984d: 94
)
(
Hypothenemus
)
comb. nov.
[
Hypothenemus
].
Eidophelus insignis
(
Browne, 1984a: 151
)
(
Hypothenemus
)
comb. nov.
[
Hypothenemus
].
Eidophelus insularis
(
Nobuchi, 1975: 48
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus insularum
(
Krivolutskaya, 1968: 56
)
(
Hypothenemus
)
comb. nov.
[
Ernoporicus
].
=
Ericryphalus elongatus
Nobuchi, 1975: 42
(syn:
Mandelshtam, 2006
).
=
Hypothenemus krivolutskayae
Wood, 1992b: 79
(syn: Alonso- Zarazaga, 2005).
Eidophelus jalappae
(
Letzner, 1849: 99
)
(
Bostrichus
)
comb. nov.
[
Scolytogenes
].
=
Ernoporides floridensis
Hopkins, 1915: 34
(syn:
Wood, 1966
).
=
Ernoporides knabi
Hopkins, 1915: 34
(syn:
Wood, 2007
).
=
Hypothenemus ritchiei
Sampson, 1918: 295
(syn:
Wood, 1966
).
=
Cryphalomorphus caraibicus
Schedl, 1951e: 96
(syn:
Wood, 1966
).
=
Cryphalomorphus minutissimus
Schedl, 1951e: 97
(syn:
Wood, 1977b
).
=
Cryphalomorphus subtriatus
Schedl, 1952c: 360
(syn:
Wood, 1966
).
=
Cryphalomorphus alienus
Schedl, 1976: 65
(syn:
Wood, 1989
).
Eidophelus javanus
(
Schedl, 1942d: 10
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus kanawhae
(
Hopkins, 1915: 35
)
(
Ernoporus
)
comb. nov.
[
Ernoporicus
].
Eidophelus kinabaluensis
(
Bright, 1992: 186
)
(
Hemicryphalus
)
comb. nov.
[
Hemicryphalus
].
Eidophelus levis
Johnson
nom. nov.
=
Eidophelus gracilis
Browne, 1984a: 152
syn. nov
(secondary homonym).
Remarks: A replacement name is proposed for
Eidophelus gracilis
Browne, 1984a
following the genus synonymy of
Eidophelus
and
Scolytogenes
, because the name is already occupied by
Lepicerinus gracilis
Schedl, 1950b
. The name is an adjective, meaning smooth.
Eidophelus landolphiae
(
Schedl, 1961b: 133
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus leprosulus
(
Browne, 1974: 45
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus longipennis
(
Eggers, 1936a: 30
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus lucidus
Johnson
nom. nov.
[
Scolytogenes
].
=
Lepicerinus pacificus
Schedl, 1959a: 477
syn. nov.
(secondary homonym).
Remarks: A replacement name is proposed for
Lepicerinus pacificus
Schedl, 1959a
following the genus synonymy of
Ptilopodius
and
Eidophelus
, because the name is already occupied by
Ptilopodius pacificus
Schedl, 1941b
. The name is an adjective, with reference to its shiny appearance.
Eidophelus magnocularis
(
Yin, 2001: 326
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus marquesanus
(
Beeson, 1935a: 101
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
=
Ptilopodius zimmermani
Schedl, 1951d: 143
(syn:
Beaver, 1991
).
Eidophelus mauritianus
(
Schedl, 1965a: 56
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus micans
(
Eggers, 1927b: 396
)
(
Stephanoderes
)
comb. nov.
[
Scolytogenes
].
Eidophelus minor
(
Eggers, 1927c: 69
)
(
Negritus
)
comb. nov.
[
Scolytogenes
].
Eidophelus minusculus
Johnson
nom. nov.
=
Eidophelus minutissimus
Schedl, 1962a: 191
syn. nov.
(secondary homonym)
Remarks: A replacement name is proposed for
Eidophelus minutissimus
Schedl, 1962a
because the name is now occupied by
Ptilopodius minutissimus
Schedl, 1943
. The replacement name is an adjective, and maintains the original meaning, referencing the very small size.
Eidophelus minutissimus
(
Schedl, 1943: 34
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus minutus
Blandford, 1894b: 88
.
Eidophelus mus
(
Schedl, 1975a: 349
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus nanulus
(
Wood, 1960: 29
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus nigellatus
(
Schedl, 1950b: 44
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
Eidophelus niger
Johnson
nom. nov.
[
Ernoporicus
].
=
Ernoporicus ater
Nobuchi, 1975: 43
syn. nov.
(secondary homonym).
Remarks: A replacement name for
Ernoporicus ater
Nobuchi, 1975
is proposed because the name is already occupied by
Negritus ater
Eggers, 1923a
. The name is an adjective, following the meaning of the original name, meaning black.
Eidophelus nitidus
Schedl, 1965b: 26
.
Eidophelus nubilus
(
Wood, 1960: 30
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus ocularis
(
Schedl, 1965c: 368
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus onyanganus
(
Schedl, 1941a: 391
)
(
Letznerella
)
comb. nov.
[
Scolytogenes
].
=
Cryphalomorphus similaris
Schedl, 1965d: 8
(syn:
Browne, 1973
).
Eidophelus opacus
(
Schedl, 1959a: 477
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus pacificus
(
Schedl, 1941b: 111
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus papuanus
(
Schedl, 1974: 459
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus papuensis
(
Wood, 1989: 179
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
=
Xylocryptus papuanus
Schedl, 1975a: 352
(secondary homonym).
Eidophelus paradoxus
(
Wood, 1992b: 80
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
=
Cryphalomorphus papuanus
Schedl, 1979c: 97
(secondary homonym).
Eidophelus parvulus
Johnson
nom. nov.
[
Cryphalus
].
=
Cryphalus parvus
Browne, 1984a: 152
syn. nov.
(secondary homonym).
Remarks:
Cryphalus parvus
Browne, 1984a
is transferred to
Eidophelus
based on the cylindrical third tarsal segment, on the entire eye, and on the near-vertical apex of declivity. A replacement name is subsequently proposed due to the name being occupied by
Hypothenoides parvus
Hopkins, 1915
. The replacement name is the diminutive of the adjective
parvus
(=small), emphasizing that particularly small body size.
Eidophelus parvus
(
Hopkins, 1915: 11
)
(
Hypothenoides
)
comb. nov.
[
Scolytogenes
].
Eidophelus philippinensis
(
Schedl, 1967a: 126
)
(
Erioschidias
)
comb. nov.
[
Cosmoderes
].
Eidophelus pityophthorinus
(
Schedl, 1943: 39
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
Eidophelus pleiocarpae
(
Schedl, 1957b: 51
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus podocarpi
(
Bright, 1992: 188
)
(
Hemicryphalus
)
comb. nov.
[
Hemicryphalus
].
Eidophelus polisquamosus
(
Yin, 2001: 329
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus praeda
(
Browne, 1978: 589
)
(
Ptilopodius
)
comb. nov.
[
Scolytogenes
].
Eidophelus puerarae
(
Choo and Woo, 1989: 58
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus pumilionides
(
Schedl, 1977c: 500
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus pumilus
(
Wood, 1960: 29
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus punctatulus
(
Nobuchi, 1976: 72
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
=
Cryphalomorphus punctatus
Nobuchi, 1975: 49
.
=
Cryphalomorphus nobuchii
Schedl, 1980: 118
(syn:
Choo and Woo, 1989
).
Eidophelus punctatus
(
Schedl, 1951b: 55
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus puncticollis
(
Schedl, 1950b: 43
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
=
Cryphalomorphus grossepunctatus
Browne, 1974: 63
(syn:
Beaver, 1991
).
Eidophelus pygmaeolus
(
Schedl, 1971c: 12
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus quadridens
(
Browne, 1983b: 71
)
(
Hypothenemus
)
comb. nov.
[
Scolytogenes
].
Eidophelus ramosus
(
Beeson, 1935b: 115
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus rhododendri
Johnson
nom. nov.
[
Hemicryphalus
].
=
Hemicryphalus squamosus
Bright, 1992: 188
syn. nov.
(secondary homonym).
Remarks: A replacement name for
Hemicryphalus squamosus
Bright, 1992
is proposed because of the transfer to
Eidophelus
, where the name is already occupied by
Lepicerinus squamosus
Schedl, 1942c
. The replacement name is a noun in the genitive case, named after the host plant.
Eidophelus robustus
(
Schedl, 1955b: 32
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus rugosus
(
Schedl, 1943: 34
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus rusticus
(
Wood, 1974a: 18
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus samoanus
Schedl, 1972b: 268
.
Eidophelus schedli
Johnson
nom. nov.
[
Scolytogenes
].
=
Cryphalomorphus ceylonicus
Schedl, 1959a: 477
syn. nov.
(secondary homonym).
Remarks: A replacement name for
Cryphalomorphus ceylonicus
Schedl, 1959a
is proposed because the name is occupied by
Ptilopodius ceylonicus
Schedl, 1959a
. The replacement name is a noun in the genitive case, honoring the original author.
Eidophelus semenovi
(
Kurentsov, 1941: 161
)
(
Eocryphalus
)
comb. nov.
[
Ernoporicus
].
Eidophelus separandus
(
Schedl, 1965a: 56
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus setifer
(
Wood, 1974a: 18
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus sodalis
(
Schedl, 1965a: 55
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus spessivtzevi
(
Berger, 1917: 243
)
(
Ernoporicus
)
comb. nov.
[
Ernoporicus
].
Eidophelus spirostachius
(
Schedl, 1958a: 557
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus splendens
(
Schedl, 1975a: 350
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus squamatilis
(
Schedl, 1977c: 500
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus squamosus
(
Schedl, 1942c: 175
)
(
Lepicerinus
)
comb. nov.
[
Scolytogenes
].
Eidophelus squamulosus
(
Eggers, 1936c: 633
)
(
Cylindrotomicus
)
comb. nov.
[
Scolytogenes
].
Eidophelus stephegynis
(
Hopkins, 1915: 11
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus sumatranus
Schedl, 1961a: 72
.
Eidophelus takahashii
(
Nobuchi, 1975: 43
)
(
Ernoporicus
)
comb. nov.
[
Ernoporicus
].
Eidophelus tarawai
(Beaver, 1990: 149) (
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus tonsus
(
Schedl, 1969: 49
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus tricolor
(
Lea, 1910: 141
)
(
Cryphalus
)
comb. nov.
[
Scolytogenes
].
Eidophelus trucis
(
Wood, 1974a: 19
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus uncatus
(
Schedl, 1971b: 373
)
(
Cryphalophilus
)
comb. nov.
[
Ptilopodius
].
Remarks:
Cryphalomorphus ater
Schedl, 1972a:146
, nomen nudum, was listed as a synonym of this species (
Wood and Bright, 1992
), but this is likely an error, probably misinterpreted from the genus transfer of
E. uncatus
and
Cryphalophilus ater
Schedl, 1972a:146
, by
Beaver (1991)
.
Eidophelus usagaricus
(
Eggers, 1922: 169
)
(
Neocryphalus
)
comb. nov.
[
Scolytogenes
].
Eidophelus varius
(
Schedl, 1975a: 350
)
(
Cryphalomorphus
)
comb. nov.
[
Scolytogenes
].
Eidophelus venustus
(
Schedl, 1953c: 78
)
(
Ptilopodius
)
comb. nov.
[
Ptilopodius
].
Eidophelus yinae
Johnson
nom. nov.
[
Scolytogenes
].
=
Scolytogenes venustus
Yin, 2001: 328
syn. nov.
(secondary homonym).
Remarks: A replacement name for
Scolytogenes venustus
Yin, 2001
is proposed because the name is occupied by
Ptilopodius venustus
Schedl, 1953c
. The replacement name is a noun in the genitive case, honoring the original author.
Eidophelus yunnanensis
(
Yin, 2001: 324
)
(
Scolytogenes
)
comb. nov.
[
Scolytogenes
].
Eidophelus zachvatkini
(
Krivolutskaya, 1958: 149
)
(
Eocryphalus
)
comb. nov.
[
Ernoporicus
].