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<mods:title id="E95E2FB9385E3069F4ADFEEAB6CAC0F1">Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<treatment LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D" httpUri="http://treatment.plazi.org/id/C246C6390F96596CA5F63331DCEABC5D" lastPageNumber="79" pageId="0" pageNumber="79">
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<taxonomicName ID-CoL="78R62" LSID="C246C639-0F96-596C-A5F6-3331DCEABC5D" authority="(DeLong &amp; Freytag)" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" baseAuthorityName="Domahovski &amp; Cavichioli" baseAuthorityYear="2023" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Coarctana rana" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="rana" status="comb. nov.">Coarctana rana (DeLong &amp; Freytag)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="79">comb. nov.</taxonomicNameLabel>
<treatment id="C246C6390F96596CA5F63331DCEABC5D" ID-GBIF-Taxon="209013701" LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D" httpUri="http://treatment.plazi.org/id/C246C6390F96596CA5F63331DCEABC5D" lastPageNumber="79" pageId="0" pageNumber="79" scope_family="Cicadellidae" scope_order="Hemiptera" scope_tribe="Gyponini">
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<taxonomicName id="977E5E92F3C5EDA65417CC04F2114488" ID-CoL="78R62" LSID="C246C639-0F96-596C-A5F6-3331DCEABC5D" authority="(DeLong &amp; Freytag)" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" baseAuthorityName="Domahovski &amp; Cavichioli" baseAuthorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana" status="comb. nov.">Coarctana rana (DeLong &amp; Freytag)</taxonomicName>
<taxonomicNameLabel id="0F37E1802F0BC202770DDEA0D41947B9" pageId="0" pageNumber="79">comb. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">Figs 17</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">, 32C, D</figureCitation>
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<paragraph id="E1CFE74645F8A419FCCEA1444AE935EB" pageId="0" pageNumber="79">
<figureCitation id="D1FCDE7056CEFA2CC45D3683E96782B1" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">Figs 17</figureCitation>
<figureCitation id="A3DC9E8C6243A19B7897A21CE4A2A46A" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">, 32C, D</figureCitation>
</paragraph>
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<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" baseAuthorityName="Domahovski &amp; Cavichioli" baseAuthorityYear="2023" class="Insecta" family="Cicadellidae" genus="Acuera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuera rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana" status="comb. nov.">Acuera rana</taxonomicName>
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<paragraph id="CBF1BDEE56571E622D0EF49E99DE8C0C" pageId="0" pageNumber="79">
<taxonomicName id="20F9F886844D7B05D132A8E712154B4D" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" baseAuthorityName="Domahovski &amp; Cavichioli" baseAuthorityYear="2023" class="Insecta" family="Cicadellidae" genus="Acuera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acuera rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana" status="comb. nov.">Acuera rana</taxonomicName>
DeLong &amp; Freytag, 1982: 309.
</paragraph>
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<paragraph pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="79">
<subSubSection id="C4F77B02DFA908AEFBED8FB4655FC5A2" pageId="0" pageNumber="79" type="diagnosis">
<paragraph id="99C832041FA33239E3E73837BCE1B067" pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="165B88A030ED5585FD04D6F71022DF52" pageId="0" pageNumber="79">
Pygofer, in dorsal view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
<figureCitation id="295D3F4B074F9B3A4DF80978DB066C3F" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
), with a small subapical process directed inwards. Subgenital plate (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
<figureCitation id="635331B45FC0E9BD74F1AB967A11A869" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
) with long filiform setae. Style (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17K</figureCitation>
<figureCitation id="AEE4566992C7C3007E1FB8ED72E03FD9" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17K</figureCitation>
) with apex expanded forming rounded lobe ventrally and small dorsal process, curved anterad. Aedeagus (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L, M</figureCitation>
<figureCitation id="36DE60EB1EBDC7D0C67E0DD4BB358879" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L, M</figureCitation>
) with apodemal process very narrow, strongly curved dorsally; shaft slender; apex slightly inflated, with two pairs of short processes: a subapical pair on posterior surface, directed dorsally and a apical pair directed ventrally, about twice as long as the subapical pair.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="79">Total length: males 7.6-8.1 mm.</paragraph>
<subSubSection id="E7A90DB9827D02C632D7AF1E4E5DD7A5" pageId="0" pageNumber="79" type="description">
<paragraph id="3032B945C14F78555D072D17E4CE883C" pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph id="648E63CD7130FC8CBBFC0B7E6E43D274" pageId="0" pageNumber="79">Total length: males 7.6-8.1 mm.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="description">
<paragraph pageId="0" pageNumber="79">Description.</paragraph>
<paragraph pageId="0" pageNumber="79">
<subSubSection id="6ADDAD8F84B9FA8B350CD57302B6B610" pageId="0" pageNumber="79" type="description">
<paragraph id="2920A7E41A3DACC1F774A9965AF7FEA6" pageId="0" pageNumber="79">Description.</paragraph>
<paragraph id="8117911B79EF4067FD3997C985CDD91D" pageId="0" pageNumber="79">
Head, in frontal view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17B</figureCitation>
<figureCitation id="8557B237152D81E80EBF1F587ACE8892" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17B</figureCitation>
), frons 1.1
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<normalizedToken id="4F8761CDA2595D3650BB40EB747A6D1F" originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately the width of scape; clypeus 1.3
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="576E48121BB8055990B56B3A943815CD" originalValue="×">x</normalizedToken>
wider than long; lateral margins straight and parallel; apex straight. Head, in lateral view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
<figureCitation id="F92D9EDEC926FB0CB0BC797D606A883C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
), crown-face transition moderately thick, with 5-6 transverse carinae; clypeus not inflated. Profemur AV row with 5-6 setae; PV row with 3-4 setae. Protibia PD row with 4 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 22-24, 12 and 13-15 macrosetae respectively. Metatarsomere I 4.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="3C75A597209DA6E5B6B2241F892FA184" originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 7-8 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="coloration">
<paragraph pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph pageId="0" pageNumber="79">
<subSubSection id="1AF8091DDF94DF2CF6692177D6D9CF1F" pageId="0" pageNumber="79" type="coloration">
<paragraph id="2F1833D84834BCBBCC1138B95DD0F985" pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph id="3F9ACFFB4E2EAEE9F064FADAF744D397" pageId="0" pageNumber="79">
Head and thorax (Figs
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A-D</figureCitation>
<figureCitation id="0B36B9D29CABB5357D2A72507DA01F9C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A-D</figureCitation>
,
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32C, D</figureCitation>
<figureCitation id="2C539D11BE39EA2992400425C02479A3" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32C, D</figureCitation>
) yellowish-brown. Head (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
<figureCitation id="1723FF7562BD8D966FFA36275C5BE9AA" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocelli; coronal suture black; ocellus surrounded by black maculae in more melanic specimens. Face (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17B</figureCitation>
<figureCitation id="AC039313A01F3A15104C484D77DF0C85" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17B</figureCitation>
) yellowish, without maculae. Pronotum (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
<figureCitation id="C1BAFAAA11E3289B1867D73EF1DDBB11" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
) with black punctures and irregular black spots near anterior margin; proepimeron (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
<figureCitation id="39465B9C345E185ECE98C34B6006BE81" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
), with black band below pronotal carina. Mesonotum (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
<figureCitation id="E33D325C98464C679AA104432EE36FCB" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17A</figureCitation>
) with dark-brown lateral angles, pair of small median black spots near scutoscutellar suture. Forewing (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17D</figureCitation>
<figureCitation id="BD0EDF7693B4138F78DCE7DD89BE177D" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17D</figureCitation>
) strongly marked by dark-brown mottling; veins yellowish-brown, outlined by dark-brown. Metatibia (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
<figureCitation id="D2964C83E5EF7AB2B2D775C242928289" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="male terminalia">
<paragraph pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph pageId="0" pageNumber="79">
<subSubSection id="5ED7D9AA5B8E1C6222435959CEAB87D1" pageId="0" pageNumber="79" type="male terminalia">
<paragraph id="9CC87D84E0BA7C713D59BEE49101297A" pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph id="178BA27797BE00FE2E1C0699175A9450" pageId="0" pageNumber="79">
Sternite VIII (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17E</figureCitation>
<figureCitation id="2F80D5F07A0B65F440150DBFA3CB7ECC" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17E</figureCitation>
) 1.5
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="F49391D40D40B6D5F3CAF41E8D5DB33A" originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; posterior margin produced medially forming short, rounded lobe. Valve (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17F</figureCitation>
<figureCitation id="F800CF0BC2747B29026B12F85833507D" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17F</figureCitation>
) narrow, 1.1
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<normalizedToken id="BF3A472CC6EEDFA0440697A4D6F41A22" originalValue="×">x</normalizedToken>
wider than long; posterior margin tapered and rounded. Pygofer, in lateral view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17G</figureCitation>
<figureCitation id="87C3AB87DF481873967DE8439DA309E3" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17G</figureCitation>
), 1.4
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<normalizedToken id="DCD5905B097BE4E778770E88EA91A87D" originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process present but reduced; anteroventral and posterodorsal margins straight; posteroventral margin broadly rounded; apex broad, slightly excavated; in dorsal view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
<figureCitation id="5421831DF37B7678C55F3A65EDF21C03" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
), with small subapical process directed inwards. Subgenital plate, in lateral view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17G</figureCitation>
<figureCitation id="35CD9A9AFE1B8611A0D332B4F9E63E40" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17G</figureCitation>
), surpassing pygofer apex; in ventral view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
<figureCitation id="43B4915197B67369A6B09D88429FD84B" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
), elongated, 3.8
<normalizedToken originalValue="×">x</normalizedToken>
<normalizedToken id="BB461566CC6CD82FB9F0892678CE0AAB" originalValue="×">x</normalizedToken>
longer than wide, maximum width medially; outer margin with long filiform setae; inner margin slightly rounded; outer margin straight; apex weakly tapered and rounded. Connective (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17I</figureCitation>
<figureCitation id="C70ADCBFD5703D5397D00CB8A13FC51C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17I</figureCitation>
) wider than long; anterior margin excavated; dorsal keel and stem reduced. Style, in dorsal view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17J</figureCitation>
<figureCitation id="29CCC77B49EA2C7075DE22FAE4070B5E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17J</figureCitation>
), with outer lobe small and rounded; strongly sinuate between outer lobe and blade; in lateral view (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17K</figureCitation>
<figureCitation id="E733F1D34C40011165BD51FD9C40A6BA" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17K</figureCitation>
), with blade slightly wider near base and tapered toward apex; dorsal and ventral margins slightly sinuate; ventral margin with apical half serrated; apex expanded forming rounded lobe ventrally and small dorsal process, curved anterad. Aedeagus (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L, M</figureCitation>
<figureCitation id="33DA03A8A5C1E2078650DA9774C5A4E6" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L, M</figureCitation>
) preatrium weakly developed; dorsal apodeme with dorsal margin slightly excavated, lateral margins produced ventrolaterally; apodemal process very narrow, strongly curved dorsally, apex tapered and acute; shaft slender, cylindrical, curved dorsally near base; apex slightly inflated, with two pairs of short processes: a subapical pair on posterior surface, directed dorsally and a apical pair directed ventrally, about twice as long as the subapical pair. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="female">
<paragraph pageId="0" pageNumber="79">Female.</paragraph>
<paragraph pageId="0" pageNumber="79">Unknown.</paragraph>
<subSubSection id="499AD85D871871CB25E0CB6FED07A86A" pageId="0" pageNumber="79" type="female">
<paragraph id="0163EF0B2E1BF29B22610FDD20C35A73" pageId="0" pageNumber="79">Female.</paragraph>
<paragraph id="AB8A5235F56B0D85AB5C5E0658E87441" pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="materials_examined">
<paragraph pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="79">
<materialsCitation collectingDate="2002-06-10" collectingDateMax="2002-07-03" collectingDateMin="2002-06-10" country="Bolivia" latitude="-16.915277" location="Vila Tunari" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" specimenCount="3" specimenCount-male="3" stateProvince="Cochabamba">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingCountry name="Bolivia">Bolivia</collectingCountry>
<subSubSection id="E5587BB09F9CA3820E44F8607E45CFEA" pageId="0" pageNumber="79" type="materials_examined">
<paragraph id="80D6F28A8F9F0B0BB068C4242B155F8A" pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph id="7217820B48833FD9DBDB2B5A0475012B" pageId="0" pageNumber="79">
<materialsCitation id="3B331FB7C51B087B9D1E30C3AC653906" ID-GBIF-Occurrence="4017211407" collectingDate="2002-06-10" collectingDateMax="2002-07-03" collectingDateMin="2002-06-10" country="Bolivia" latitude="-16.915277" location="Vila Tunari" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" specimenCount="3" specimenCount-male="3" stateProvince="Cochabamba">
<emphasis id="1B8BF238FEEBD5AD8EF79E8A3952E4DF" bold="true" pageId="0" pageNumber="79">
<collectingCountry id="C34D82C80899ABA593C97C4765A7A61F" name="Bolivia">Bolivia</collectingCountry>
</emphasis>
:
<specimenCount type="male">3♂</specimenCount>
<specimenCount id="C86E07DA527363237F8629C33BA967C4" type="male">3♂</specimenCount>
,
<normalizedToken originalValue="“Bolívia">
<normalizedToken id="9F2E219A824FA2DB06328C57E1047B40" originalValue="“Bolívia">
&quot;
<collectingMunicipality>Bolivia</collectingMunicipality>
<collectingMunicipality id="4B9E6B7535E61E75DE5BC69B6D676E22">Bolivia</collectingMunicipality>
</normalizedToken>
,
<collectingRegion country="Bolivia" name="Cochabamba">Cochabamba</collectingRegion>
<collectingRegion id="1ECA4429952B75D3A2CAAED2A69EB1DD" country="Bolivia" name="Cochabamba">Cochabamba</collectingRegion>
,\
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:FB3CF4D60D8497A4C450613F7C623833" country="Bolivia" latitude="-16.915277" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" name="Vila Tunari" stateProvince="Cochabamba">Vila Tunari</location>
<location id="FB3CF4D60D8497A4C450613F7C623833" LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:FB3CF4D60D8497A4C450613F7C623833" country="Bolivia" latitude="-16.915277" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" name="Vila Tunari" stateProvince="Cochabamba">Vila Tunari</location>
\
<geoCoordinate degrees="16" direction="south" minutes="54" orientation="latitude" precision="15" seconds="55" value="-16.915277">16°5455″S</geoCoordinate>
<geoCoordinate degrees="65" direction="west" minutes="22" orientation="longitude" precision="15" seconds="06" value="-65.36833">65°2206″W</geoCoordinate>
<geoCoordinate id="F5A28B27367C280964D68D969F4EF706" degrees="16" direction="south" minutes="54" orientation="latitude" precision="15" seconds="55" value="-16.915277">16°5455″S</geoCoordinate>
<geoCoordinate id="04462A92CB1FBB7052FB26874CC27F9D" degrees="65" direction="west" minutes="22" orientation="longitude" precision="15" seconds="06" value="-65.36833">65°2206″W</geoCoordinate>
\
<collectingDate value="2002-06-10" valueMax="2002-07-03" valueMin="2002-06-10">10.VI-03.VII.2002</collectingDate>
<collectingDate id="AD334AC48F2570B077A89A83C1AF97AC" value="2002-06-10" valueMax="2002-07-03" valueMin="2002-06-10">10.VI-03.VII.2002</collectingDate>
\ malaise Helmut Heidas&quot; (DZRJ)
</materialsCitation>
;
<materialsCitation collectingDate="2002-09-07" collectingDateMax="2002-09-20" collectingDateMin="2002-09-07" country="Bolivia" latitude="-16.915277" location="Vila Tunari" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" specimenCount="1" specimenCount-male="1" stateProvince="Cochabamba">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="21E94EC10206A91717D8A37842A8E882" ID-GBIF-Occurrence="4017211388" collectingDate="2002-09-07" collectingDateMax="2002-09-20" collectingDateMin="2002-09-07" country="Bolivia" latitude="-16.915277" location="Vila Tunari" longLatPrecision="21" longitude="-65.36833" municipality="Bolivia" specimenCount="1" specimenCount-male="1" stateProvince="Cochabamba">
<specimenCount id="A36DCBF84736C0DE3393CC4A90BE42F7" type="male">1♂</specimenCount>
, same data as preceding except
<collectingDate value="2002-09-07" valueMax="2002-09-20" valueMin="2002-09-07">07-20.IX.2002</collectingDate>
<collectingDate id="F2E2AE71FF058BC23167B1C4EE276A83" value="2002-09-07" valueMax="2002-09-20" valueMin="2002-09-07">07-20.IX.2002</collectingDate>
(DZUP)
</materialsCitation>
. -
<materialsCitation collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" location="Rondonia" specimenCount="1" stateProvince="Rondonia">
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingCountry name="Brazil">Brazil</collectingCountry>
<materialsCitation id="56D64473C1A688D507D79899BAE1B499" ID-GBIF-Occurrence="4017211421" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Brazil" location="Rondonia" specimenCount="1" stateProvince="Rondonia">
<emphasis id="5FC10776A7EC2E6EA3B4E9DFB3A32702" bold="true" pageId="0" pageNumber="79">
<collectingCountry id="92E07F755878806213076348A5980F8F" name="Brazil">Brazil</collectingCountry>
:
<normalizedToken originalValue="Rondônia">
<collectingRegion country="Brazil" name="Rondonia">Rondonia</collectingRegion>
<normalizedToken id="0AC8787D4B4BEA1E2ABC06D2B68F9447" originalValue="Rondônia">
<collectingRegion id="FE33779BE69A2518EAFFB8D386BB6786" country="Brazil" name="Rondonia">Rondonia</collectingRegion>
</normalizedToken>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="1987-10-31" collectorName="C. Elias" country="Peru" location="Ouro Preto" specimenCount="1" specimenCount-male="1">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="A7D9C2F632F055C2FB6FDA28D2798618" ID-GBIF-Occurrence="4017211317" collectingDate="1987-10-31" collectorName="C. Elias" country="Peru" location="Ouro Preto" specimenCount="1" specimenCount-male="1">
<specimenCount id="0B6EC560CE7387C90F216739F537DA59" type="male">1♂</specimenCount>
, &quot;
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:0A8AC486653B468A2623A5D820078CF8" country="Peru" name="Ouro Preto">Ouro Preto</location>
<location id="0A8AC486653B468A2623A5D820078CF8" LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:0A8AC486653B468A2623A5D820078CF8" country="Peru" name="Ouro Preto">Ouro Preto</location>
\ dOeste, RO.\
<collectingDate value="1987-10-31">31.X.1987</collectingDate>
<collectingDate id="19191676879D749E9B99455AC191AFA9" value="1987-10-31">31.X.1987</collectingDate>
\
<collectorName>C. Elias</collectorName>
<collectorName id="65C5BE4CE7DD8AEBA8251A3F23FE82E1">C. Elias</collectorName>
, leg&quot; (DZUP). -
<emphasis bold="true" pageId="0" pageNumber="79">
<collectingCountry name="Peru">Peru</collectingCountry>
<emphasis id="26F895D38D4DFF265D10AA7D31FA68CA" bold="true" pageId="0" pageNumber="79">
<collectingCountry id="78D399C629BB4EE29C14292A83C7311E" name="Peru">Peru</collectingCountry>
</emphasis>
</materialsCitation>
:
<materialsCitation collectingDate="2005-03-06" collectingDateMax="2005-03-15" collectingDateMin="2005-03-06" country="Peru" county="Peru" elevation="335" latitude="-7.081333" location="Mun. Zona Barreal" longLatPrecision="13" longitude="-79.314835" municipality="Cons." specimenCount="2" specimenCount-male="2" stateProvince="San Martin">
<specimenCount type="male">2♂</specimenCount>
<materialsCitation id="024DA925F5B62892D65B3618C99EE7EA" ID-GBIF-Occurrence="4017211400" collectingDate="2005-03-06" collectingDateMax="2005-03-15" collectingDateMin="2005-03-06" country="Peru" county="Peru" elevation="335" latitude="-7.081333" location="Mun. Zona Barreal" longLatPrecision="13" longitude="-79.314835" municipality="Cons." specimenCount="2" specimenCount-male="2" stateProvince="San Martin">
<specimenCount id="7DC5D12E76AE053475646784EDCC84D8" type="male">2♂</specimenCount>
, &quot;
<collectingCounty>Peru</collectingCounty>
<collectingCounty id="7EC131AB0F15A698D544B6BB4B290F7A">Peru</collectingCounty>
:
<collectingRegion country="Peru" name="San Martin">San Martin</collectingRegion>
<collectingRegion id="DAEFAC2A008A50E88804016E74EF079D" country="Peru" name="San Martin">San Martin</collectingRegion>
,
<collectingMunicipality>Cons.</collectingMunicipality>
<collectingMunicipality id="9B67B72BA8FBF2E5FA5384DF3969DEC8">Cons.</collectingMunicipality>
\
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:AAC657CE158BADB295BC5DF75C9F13C4" country="Peru" county="Peru" latitude="-7.081333" longLatPrecision="13" longitude="-79.314835" municipality="Cons." name="Mun. Zona Barreal" stateProvince="San Martin">Mun. Zona Barreal</location>
<location id="AAC657CE158BADB295BC5DF75C9F13C4" LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:AAC657CE158BADB295BC5DF75C9F13C4" country="Peru" county="Peru" latitude="-7.081333" longLatPrecision="13" longitude="-79.314835" municipality="Cons." name="Mun. Zona Barreal" stateProvince="San Martin">Mun. Zona Barreal</location>
23Km S\
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:04E941E52304E5D8EA2661A92C923242" country="Peru" county="Peru" latitude="-7.081333" longLatPrecision="13" longitude="-79.314835" municipality="Cons." name="Picota" stateProvince="San Martin">Picota</location>
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.35" unit="m" value="335.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.35" unit="m" value="335.0">335m</elevation>
<location id="04E941E52304E5D8EA2661A92C923242" LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:04E941E52304E5D8EA2661A92C923242" country="Peru" county="Peru" latitude="-7.081333" longLatPrecision="13" longitude="-79.314835" municipality="Cons." name="Picota" stateProvince="San Martin">Picota</location>
<quantity id="C1480C6C9712123163120357405697A4" metricMagnitude="2" metricUnit="m" metricValue="3.35" unit="m" value="335.0">
<elevation id="39D4233658EC3AE0DEA5FFA317EEE2D4" metricMagnitude="2" metricUnit="m" metricValue="3.35" unit="m" value="335.0">335m</elevation>
</quantity>
,
<geoCoordinate degrees="7" direction="south" minutes="04.88" orientation="latitude" precision="9" value="-7.081333">7°04.88S</geoCoordinate>
<geoCoordinate id="1DBC611E506A790CCC32B946EB6A6B19" degrees="7" direction="south" minutes="04.88" orientation="latitude" precision="9" value="-7.081333">7°04.88S</geoCoordinate>
,
<geoCoordinate degrees="79" direction="west" minutes="18.89" orientation="longitude" precision="9" value="-79.314835">79°18.89W</geoCoordinate>
<collectingDate value="2005-03-06" valueMax="2005-03-15" valueMin="2005-03-06">6-15Mar2005</collectingDate>
<geoCoordinate id="B5560B8E93E6306C5AB211A43678552D" degrees="79" direction="west" minutes="18.89" orientation="longitude" precision="9" value="-79.314835">79°18.89W</geoCoordinate>
<collectingDate id="906188C6D09C5A6A4D7846814279B9D7" value="2005-03-06" valueMax="2005-03-15" valueMin="2005-03-06">6-15Mar2005</collectingDate>
,\ ME Irwin, JD Vasquez,\ Malaise, dry Florest PE 11-02&quot; (DZRJ)
</materialsCitation>
;
<materialsCitation collectingDate="2009-01-23" collectingDateMax="2009-01-30" collectingDateMin="2009-01-23" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="B5FAB056056E88175F2A5BC6DCE00D28" ID-GBIF-Occurrence="4017211423" collectingDate="2009-01-23" collectingDateMax="2009-01-30" collectingDateMin="2009-01-23" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount id="81D887D46495DEDAFED84A011C91D001" type="male">1♂</specimenCount>
, &quot;
<collectingCounty>Peru</collectingCounty>
<collectingCounty id="F0E30366092F7B76DC4266A8E5AE8C4E">Peru</collectingCounty>
:
<collectingMunicipality>Dept</collectingMunicipality>
<collectingRegion country="Peru" name="Amazonas">Amazonas</collectingRegion>
<collectingMunicipality id="64FD29F5E6C72AB1200D7DF15A5153AE">Dept</collectingMunicipality>
<collectingRegion id="94CBCE7D95C9B80C29299ACAE0F94BF2" country="Peru" name="Amazonas">Amazonas</collectingRegion>
,\
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:145EFE81248CB68D1BB83221C6EC6CB8" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Distr. Aguas Verdes" stateProvince="Amazonas">
<location id="145EFE81248CB68D1BB83221C6EC6CB8" LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:145EFE81248CB68D1BB83221C6EC6CB8" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Distr. Aguas Verdes" stateProvince="Amazonas">
Distr.
<normalizedToken originalValue="Águas">Aguas</normalizedToken>
<normalizedToken id="CCEC4C11997857EBC09EBD1C428D5734" originalValue="Águas">Aguas</normalizedToken>
Verdes
</location>
,\
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:4580DFDF3751AC418563BAA9F00DA064" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Bagua Taparoto Rd" stateProvince="Amazonas">Bagua Taparoto Rd</location>
<location id="4580DFDF3751AC418563BAA9F00DA064" LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:4580DFDF3751AC418563BAA9F00DA064" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Bagua Taparoto Rd" stateProvince="Amazonas">Bagua Taparoto Rd</location>
(5N)\ at
<location LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:254D4982CD4A163BD955FAB78A57133D" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Km" stateProvince="Amazonas">Km</location>
<location id="254D4982CD4A163BD955FAB78A57133D" LSID="urn:lsid:plazi:treatment:C246C6390F96596CA5F63331DCEABC5D:254D4982CD4A163BD955FAB78A57133D" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Dept" name="Km" stateProvince="Amazonas">Km</location>
403;
<collectingDate value="2009-01-23" valueMax="2009-01-30" valueMin="2009-01-23">23-30.I.2009</collectingDate>
<collectingDate id="440BAD07029EC3645C7430FDCA30956F" value="2009-01-23" valueMax="2009-01-30" valueMin="2009-01-23">23-30.I.2009</collectingDate>
\ ME Irwin, GA Amaya,\
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">1125m</elevation>
<quantity id="AB8EE94D7BE91D684132ADA63F2E9E60" metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">
<elevation id="7BF43786C5AF24F3737B4F743D40EDE8" metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">1125m</elevation>
</quantity>
;
<geoCoordinate degrees="05" direction="south" minutes="41" orientation="latitude" precision="15" seconds="23" value="-5.689722">05°4123″S</geoCoordinate>
<geoCoordinate degrees="77" direction="west" minutes="38" orientation="longitude" precision="15" seconds="13" value="-77.63694">77°3813″W</geoCoordinate>
<geoCoordinate id="B8C2C53BC092CCF0A0439FA869076214" degrees="05" direction="south" minutes="41" orientation="latitude" precision="15" seconds="23" value="-5.689722">05°4123″S</geoCoordinate>
<geoCoordinate id="8E8C0BC4D3CC2C47248E5E4477CA484E" degrees="77" direction="west" minutes="38" orientation="longitude" precision="15" seconds="13" value="-77.63694">77°3813″W</geoCoordinate>
&quot; (DZRJ)
</materialsCitation>
;
<materialsCitation collectingDate="2009-01-30" collectingDateMax="2009-02-06" collectingDateMin="2009-01-30" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="D7F9D8517683F5AEE9A2D3AC0D73E041" ID-GBIF-Occurrence="4017211305" collectingDate="2009-01-30" collectingDateMax="2009-02-06" collectingDateMin="2009-01-30" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount id="A52668933348EC5FECDA902C294862CB" type="male">1♂</specimenCount>
, same data as preceding except
<collectingDate value="2009-01-30" valueMax="2009-02-06" valueMin="2009-01-30">30.I-06.II.2009</collectingDate>
<collectingDate id="04321F62E676189A70F7745C9422F787" value="2009-01-30" valueMax="2009-02-06" valueMin="2009-01-30">30.I-06.II.2009</collectingDate>
(DZRJ)
</materialsCitation>
;
<materialsCitation collectingDate="2009-05-15" collectingDateMax="2009-05-22" collectingDateMin="2009-05-15" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount type="male">1♂</specimenCount>
<materialsCitation id="F618F4E1FF92D69B94AE204B34A203B3" ID-GBIF-Occurrence="4017211310" collectingDate="2009-05-15" collectingDateMax="2009-05-22" collectingDateMin="2009-05-15" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Distr. Aguas Verdes" longLatPrecision="21" longitude="-77.63694" municipality="Dept" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas">
<specimenCount id="0A0ADBBC4CC044EC5723A774252ADD3B" type="male">1♂</specimenCount>
, same data as preceding except
<collectingDate value="2009-05-15" valueMax="2009-05-22" valueMin="2009-05-15">15-22.V.2009</collectingDate>
<collectingDate id="BFD9BFA9FFE10C1916EFD9DF55E6E90C" value="2009-05-15" valueMax="2009-05-22" valueMin="2009-05-15">15-22.V.2009</collectingDate>
(DZUP)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="79" type="remarks">
<paragraph pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="79">
<subSubSection id="234A0B7D5D81F01A32FFBAAA633CEB62" pageId="0" pageNumber="79" type="remarks">
<paragraph id="AD74863103AC197EFF36820F39356A81" pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph id="55792104D67EE28B953E6ACD97647336" pageId="0" pageNumber="79">
Our analysis recovered
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. rana" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="rana">
<emphasis italics="true" pageId="0" pageNumber="79">C. rana</emphasis>
<taxonomicName id="6386B039BA900EAFFBD8DC5171E9D597" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. rana" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="rana">
<emphasis id="7B97A4EFDCB919B46CA6704769928561" italics="true" pageId="0" pageNumber="79">C. rana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
<emphasis id="0F2D5EEDE8FF64FE69054D25EA4CBC3A" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
as sister to
<taxonomicName authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" family="Cicadellidae" kingdom="Animalia" lsidName="C. serpenta" order="Hemiptera" pageId="0" pageNumber="79" rank="species" species="serpenta">
<emphasis italics="true" pageId="0" pageNumber="79">C. serpenta</emphasis>
<taxonomicName id="06E2974EA4045FC02E2DAD1E1B1FA2BB" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. serpenta" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="serpenta">
<emphasis id="EDF39EA05F08CBF302839E15CB6AC42C" italics="true" pageId="0" pageNumber="79">C. serpenta</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
<emphasis id="E8C1C61E8AAB6276875BDF6A6863319B" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
with low branch support (SR &lt;50) (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
<figureCitation id="61203A1E2B6D82E4BD534B9B82AA6404" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
). These species are similar in the overall coloration (Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32C-F</figureCitation>
<figureCitation id="C7EF7C60DFB43FB580AAEC8E5DE424A0" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. punctata comb. nov., holotype female, NHRS GULI 000069783. C, D C. rana comb. nov., holotype male, OSU. E, F C. serpenta comb. nov., holotype male, AMNH. G, H C. striata sp. nov., holotype male, DZUP. I, J C. taurina sp. nov., holotype male, DZUP. K, L C. vilavelha sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure32" httpUri="https://binary.pensoft.net/fig/798185" pageId="0" pageNumber="79">32C-F</figureCitation>
); in having a small digitiform processes at pygofer apex (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
<figureCitation id="837B408E45C27BA5FD9F9085BE7FC29A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17g</figureCitation>
); the subgenital plates (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
<figureCitation id="BF0B40AE981D98413535DBB128910794" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17H</figureCitation>
) with long filiform setae; and in the shape of the apodemal processes of aedeagus, slender and strongly curved (Fig.
<figureCitation captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L</figureCitation>
<figureCitation id="9795A35B24EE2E5CAA58E578998CAD02" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Coarctana rana comb. nov., male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view; g apical portion of pygofer, dorsal view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure17" httpUri="https://binary.pensoft.net/fig/798170" pageId="0" pageNumber="79">17L</figureCitation>
).
</paragraph>
</subSubSection>