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<document id="53DCBA42F7475FE949AD2C226ACF8F33" ID-DOI="10.17161/jom.v0i10.4513" ID-ISSN="2325-4467" ID-ZooBank="urn:lsid:zoobank.org:pub:2A491D5E-45DB-4D83-8137-AB1EA6B49523" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722893275264" checkinUser="felipe" docAuthor="Engel, Michael S. &amp; Packer, Laurence" docDate="2013" docId="05237E4C5906614ECC51FC0AF0E923AB" docLanguage="en" docName="JMelittology.2013.10.1-7.pdf" docOrigin="Journal of Melittology 2013 (10)" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.4:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="4" docTitle="Chiasmognathus scythicus Engel &amp; Packer 2013, new species" docType="treatment" docUuid="F5C60D49-59D0-4772-A771-E77B712E8823" docUuidSource="ZooBank" docVersion="2" lastPageNumber="7" masterDocId="F91A063459076148CD1FFFEBF1412212" masterDocTitle="A new species of Chiasmognathus from Kazakhstan (Hymenoptera: Apidae)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1725374010178" updateUser="felipe" zenodo-license-document="CC-BY-NC-ND-4.0"> <document id="F9501C90075EBDFCACF2E8C783996CAD" ID-CLB-Dataset="302770" ID-DOI="10.17161/jom.v0i10.4513" ID-GBIF-Dataset="79ffb9b6-c69e-4eb7-9178-5bbb997b3b48" ID-ISSN="2325-4467" ID-Zenodo-Dep="13737328" ID-ZooBank="urn:lsid:zoobank.org:pub:2A491D5E-45DB-4D83-8137-AB1EA6B49523" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722893275264" checkinUser="felipe" docAuthor="Engel, Michael S. &amp; Packer, Laurence" docDate="2013" docId="05237E4C5906614ECC51FC0AF0E923AB" docLanguage="en" docName="JMelittology.2013.10.1-7.pdf" docOrigin="Journal of Melittology 2013 (10)" docStyle="DocumentStyle:20C606B715423E98DFED429F4976A416.4:JMelittology.2013-.journal_article.0cover" docStyleId="20C606B715423E98DFED429F4976A416" docStyleName="JMelittology.2013-.journal_article.0cover" docStyleVersion="4" docTitle="Chiasmognathus scythicus Engel &amp; Packer 2013, new species" docType="treatment" docUuid="F5C60D49-59D0-4772-A771-E77B712E8823" docUuidSource="ZooBank" docVersion="4" lastPageNumber="7" masterDocId="F91A063459076148CD1FFFEBF1412212" masterDocTitle="A new species of Chiasmognathus from Kazakhstan (Hymenoptera: Apidae)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1725895003428" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
<mods:mods id="C614AA9CDE9A7CDD07AEB035BDF673D9" xmlns:mods="http://www.loc.gov/mods/v3"> <mods:mods id="F7881A2B0E977FF3339FA3DC481778D4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="86A88D88AE3500F7D0FEAEA905370334"> <mods:titleInfo id="122D2BB5104B53DA78872CF8A2F9FF56">
<mods:title id="0DF43699AB401AC6D9B543C63AACEB4C">A new species of Chiasmognathus from Kazakhstan (Hymenoptera: Apidae)</mods:title> <mods:title id="F18416448514259EEAE4FE3E6E84F880">A new species of Chiasmognathus from Kazakhstan (Hymenoptera: Apidae)</mods:title>
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<mods:namePart id="13ED8065FEDF1E2D6E768029D84F63FC">Engel, Michael S.</mods:namePart> <mods:namePart id="B6B13AB59D8CE33A2FD3ECD381B05DC7">Engel, Michael S.</mods:namePart>
<mods:affiliation id="DB6A701E27838345AE80C2BC177C53D5">Division of Entomology, Natural History Museum, and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu).</mods:affiliation> <mods:affiliation id="567C31A3AF0AD3591D0F702C7721DE5C">Division of Entomology, Natural History Museum, and Department of Ecology &amp; Evolutionary Biology, 1501 Crestline Drive - Suite 140, University of Kansas, Lawrence, Kansas 66045, USA (msengel @ ku. edu).</mods:affiliation>
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<mods:namePart id="85ED742A8BC5E6A07862C8E6EC66F253">Packer, Laurence</mods:namePart> <mods:namePart id="1BDAEF1DB444669F6E9679E088F9F334">Packer, Laurence</mods:namePart>
<mods:affiliation id="115907E0576B1CF83EFA11B6CB70C6A4">Department of Biology, York University, 4700 Keele Street, Toronto, Ontario M 3 J 1 P 3, Canada (xeromelissa @ mail. com).</mods:affiliation> <mods:affiliation id="111F88589C08FAD29948F5D98946DC99">Department of Biology, York University, 4700 Keele Street, Toronto, Ontario M 3 J 1 P 3, Canada (xeromelissa @ mail. com).</mods:affiliation>
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<treatment id="05237E4C5906614ECC51FC0AF0E923AB" LSID="urn:lsid:zoobank.org:act:F5C60D49-59D0-4772-A771-E77B712E8823" httpUri="http://treatment.plazi.org/id/05237E4C5906614ECC51FC0AF0E923AB" lastPageId="6" lastPageNumber="7" pageId="1" pageNumber="2"> <treatment id="05237E4C5906614ECC51FC0AF0E923AB" ID-DOI="http://doi.org/10.5281/zenodo.13737338" ID-Zenodo-Dep="13737338" LSID="urn:lsid:zoobank.org:act:F5C60D49-59D0-4772-A771-E77B712E8823" httpUri="http://treatment.plazi.org/id/05237E4C5906614ECC51FC0AF0E923AB" lastPageId="6" lastPageNumber="7" pageId="1" pageNumber="2">
<subSubSection id="C5909CD159066149CC51FC0AF571260B" pageId="1" pageNumber="2" type="nomenclature"> <subSubSection id="C5909CD159066149CC51FC0AF571260B" pageId="1" pageNumber="2" type="nomenclature">
<paragraph id="8D35CF5A59066149CC51FC0AF2A321EB" blockId="1.[256,1072,993,1081]" box="[334,994,993,1017]" pageId="1" pageNumber="2"> <paragraph id="8D35CF5A59066149CC51FC0AF2A321EB" blockId="1.[256,1072,993,1081]" box="[334,994,993,1017]" pageId="1" pageNumber="2">
<heading id="D67D783659066149CC51FC0AF2A321EB" box="[334,994,993,1017]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2"> <heading id="D67D783659066149CC51FC0AF2A321EB" box="[334,994,993,1017]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="2" reason="2">
@ -67,7 +70,7 @@ urn:lsid:zoobank.org:act:
<subSubSection id="C5909CD159066149CF4AFBCAF39A262B" box="[597,731,1057,1081]" pageId="1" pageNumber="2" type="description"> <subSubSection id="C5909CD159066149CF4AFBCAF39A262B" box="[597,731,1057,1081]" pageId="1" pageNumber="2" type="description">
<paragraph id="8D35CF5A59066149CF4AFBCAF39A262B" blockId="1.[256,1072,993,1081]" box="[597,731,1057,1081]" pageId="1" pageNumber="2"> <paragraph id="8D35CF5A59066149CF4AFBCAF39A262B" blockId="1.[256,1072,993,1081]" box="[597,731,1057,1081]" pageId="1" pageNumber="2">
( (
<figureCitation id="15B1D3DF59066149CF41FBCAF393262B" box="[606,722,1057,1081]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–8" captionStart-2="Figures 9–11" captionStartId-0="2.[160,241,1409,1431]" captionStartId-1="3.[161,242,1224,1246]" captionStartId-2="4.[160,241,1410,1432]" captionTargetBox-0="[163,1137,241,1385]" captionTargetBox-1="[194,1081,244,1200]" captionTargetBox-2="[172,1135,246,1379]" captionTargetId-0="figure-110@2.[160,1168,224,1400]" captionTargetId-1="figure-173@3.[161,1169,224,1217]" captionTargetId-2="figure-128@4.[160,1168,224,1400]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figures 1–3. Photomicrographs of male of Chiasmognathus scythicus, new species. 1. Lateral habitus. 2. Dorsal habitus. 3. Facial view." captionText-1="Figures 4–8. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." captionText-2="Figures 9–11. Photomicrographs of female of Chiasmognathus scythicus, new species. 9. Lateral habitus. 10. Dorsal habitus. 11. Facial view." pageId="1" pageNumber="2">Figs. 1–11</figureCitation> <figureCitation id="15B1D3DF59066149CF41FBCAF393262B" box="[606,722,1057,1081]" captionStart-0="Figures 1–3" captionStart-1="Figures 4–8" captionStart-2="Figures 9–11" captionStartId-0="2.[160,241,1409,1431]" captionStartId-1="3.[161,242,1224,1246]" captionStartId-2="4.[160,241,1410,1432]" captionTargetBox-0="[163,1137,241,1385]" captionTargetBox-1="[194,1081,244,1200]" captionTargetBox-2="[172,1135,246,1379]" captionTargetId-0="figure-110@2.[160,1168,224,1400]" captionTargetId-1="figure-173@3.[161,1169,224,1217]" captionTargetId-2="figure-128@4.[160,1168,224,1400]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figures 1–3. Photomicrographs of male of Chiasmognathus scythicus, new species. 1. Lateral habitus. 2. Dorsal habitus. 3. Facial view." captionText-1="Figures 4–8. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." captionText-2="Figures 9–11. Photomicrographs of female of Chiasmognathus scythicus, new species. 9. Lateral habitus. 10. Dorsal habitus. 11. Facial view." figureDoi-0="http://doi.org/10.5281/zenodo.13737330" figureDoi-1="http://doi.org/10.5281/zenodo.13737332" figureDoi-2="http://doi.org/10.5281/zenodo.13737334" httpUri-0="https://zenodo.org/record/13737330/files/figure.png" httpUri-1="https://zenodo.org/record/13737332/files/figure.png" httpUri-2="https://zenodo.org/record/13737334/files/figure.png" pageId="1" pageNumber="2">Figs. 1–11</figureCitation>
) )
</paragraph> </paragraph>
</subSubSection> </subSubSection>
@ -78,7 +81,7 @@ DIAGNOSIS: Superficially the new species resembles
<emphasis id="BFFE134859066149CE35FB89F291266B" box="[810,976,1121,1145]" italics="true" pageId="1" pageNumber="2">C. gussakovskii</emphasis> <emphasis id="BFFE134859066149CE35FB89F291266B" box="[810,976,1121,1145]" italics="true" pageId="1" pageNumber="2">C. gussakovskii</emphasis>
</taxonomicName> </taxonomicName>
but differs most notably by the shape of the hidden metasomal sterna ( but differs most notably by the shape of the hidden metasomal sterna (
<figureCitation id="15B1D3DF59066149CE3BFB6AF2CF268B" box="[804,910,1153,1177]" captionStart="Figures 4–8" captionStartId="3.[161,242,1224,1246]" captionTargetBox="[194,1081,244,1200]" captionTargetId="figure-173@3.[161,1169,224,1217]" captionTargetPageId="3" captionText="Figures 4–8. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." pageId="1" pageNumber="2">Figs. 4, 5</figureCitation> <figureCitation id="15B1D3DF59066149CE3BFB6AF2CF268B" box="[804,910,1153,1177]" captionStart="Figures 4–8" captionStartId="3.[161,242,1224,1246]" captionTargetBox="[194,1081,244,1200]" captionTargetId="figure-173@3.[161,1169,224,1217]" captionTargetPageId="3" captionText="Figures 4–8. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13737332" httpUri="https://zenodo.org/record/13737332/files/figure.png" pageId="1" pageNumber="2">Figs. 4, 5</figureCitation>
: :
<emphasis id="BFFE134859066149CE85FB6AF2ED268B" box="[922,940,1153,1177]" italics="true" pageId="1" pageNumber="2">cf</emphasis> <emphasis id="BFFE134859066149CE85FB6AF2ED268B" box="[922,940,1153,1177]" italics="true" pageId="1" pageNumber="2">cf</emphasis>
. .
@ -106,7 +109,7 @@ but differs most notably by the shape of the hidden metasomal sterna (
(approximately ¼ the length according to (approximately ¼ the length according to
<bibRefCitation id="E91BB2AB59066149C947FA69F04627AB" author="Popov, V. B." pageId="1" pageNumber="2" pagination="10 - 14" refId="ref2932" refString="Popov, V. B. 1937. Zur Kenntnis der Bienengattung Parammobatodes Popov (Hymenoptera, Apoidea). Konowia 16 (1): 10 - 14." type="journal article" year="1937">Popov, 1937</bibRefCitation> <bibRefCitation id="E91BB2AB59066149C947FA69F04627AB" author="Popov, V. B." pageId="1" pageNumber="2" pagination="10 - 14" refId="ref2932" refString="Popov, V. B. 1937. Zur Kenntnis der Bienengattung Parammobatodes Popov (Hymenoptera, Apoidea). Konowia 16 (1): 10 - 14." type="journal article" year="1937">Popov, 1937</bibRefCitation>
: fig. 2) resulting in more prominent dorsoapical lobes ( : fig. 2) resulting in more prominent dorsoapical lobes (
<figureCitation id="15B1D3DF59066149CEB5FA4AF2AE27AB" box="[938,1007,1441,1465]" captionStart="Figures 4–8" captionStartId="3.[161,242,1224,1246]" captionTargetBox="[194,1081,244,1200]" captionTargetId="figure-173@3.[161,1169,224,1217]" captionTargetPageId="3" captionText="Figures 4–8. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." pageId="1" pageNumber="2">Fig. 8</figureCitation> <figureCitation id="15B1D3DF59066149CEB5FA4AF2AE27AB" box="[938,1007,1441,1465]" captionStart="Figures 4–8" captionStartId="3.[161,242,1224,1246]" captionTargetBox="[194,1081,244,1200]" captionTargetId="figure-173@3.[161,1169,224,1217]" captionTargetPageId="3" captionText="Figures 4–8. Male terminalia of Chiasmognathus scythicus, new species. 4. Seventh metasomal sternum. 5. Eighth metasomal sternum. 6. Genital capsule, ventral view. 7. Genital capsule, lateral view. 8. Genital capsule, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.13737332" httpUri="https://zenodo.org/record/13737332/files/figure.png" pageId="1" pageNumber="2">Fig. 8</figureCitation>
). The sterna more closely resemble the shape observed in ). The sterna more closely resemble the shape observed in
<taxonomicName id="4A8AB4D959066149CFA2FA29F54C27CB" authority="(Warncke, 1983)" baseAuthorityName="Warncke" baseAuthorityYear="1983" box="[701,1037,1473,1497]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="orientanus"> <taxonomicName id="4A8AB4D959066149CFA2FA29F54C27CB" authority="(Warncke, 1983)" baseAuthorityName="Warncke" baseAuthorityYear="1983" box="[701,1037,1473,1497]" class="Insecta" family="Apidae" genus="Chiasmognathus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="orientanus">
<emphasis id="BFFE134859066149CFA2FA29F21027CB" box="[701,849,1474,1497]" italics="true" pageId="1" pageNumber="2">C. orientanus</emphasis> <emphasis id="BFFE134859066149CFA2FA29F21027CB" box="[701,849,1474,1497]" italics="true" pageId="1" pageNumber="2">C. orientanus</emphasis>
@ -161,7 +164,7 @@ DESCRIPTION:
<quantity id="4A7262BF5905614ACE02F937F28524E6" box="[797,964,1756,1780]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="6.0" metricValueMin="4.3" pageId="2" pageNumber="3" unit="mm" value="0.515" valueMax="0.6" valueMin="0.43">0.43–0.60 mm</quantity> <quantity id="4A7262BF5905614ACE02F937F28524E6" box="[797,964,1756,1780]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="6.0" metricValueMin="4.3" pageId="2" pageNumber="3" unit="mm" value="0.515" valueMax="0.6" valueMin="0.43">0.43–0.60 mm</quantity>
). Forewing marginal cell broadly truncate; both m-cu crossveins entering second submarginal cell. Terminalia as depicted in figures 4–8. ). Forewing marginal cell broadly truncate; both m-cu crossveins entering second submarginal cell. Terminalia as depicted in figures 4–8.
</paragraph> </paragraph>
<caption id="D9F59FD25905614ACDBFFA6AF31D27A6" pageId="2" pageNumber="3" startId="2.[160,241,1409,1431]" targetBox="[163,1137,241,1385]" targetPageId="2" targetType="figure"> <caption id="D9F59FD25905614ACDBFFA6AF31D27A6" ID-DOI="http://doi.org/10.5281/zenodo.13737330" ID-Zenodo-Dep="13737330" httpUri="https://zenodo.org/record/13737330/files/figure.png" pageId="2" pageNumber="3" startId="2.[160,241,1409,1431]" targetBox="[163,1137,241,1385]" targetPageId="2" targetType="figure">
<paragraph id="8D35CF5A5905614ACDBFFA6AF31D27A6" blockId="2.[160,1124,1409,1460]" pageId="2" pageNumber="3"> <paragraph id="8D35CF5A5905614ACDBFFA6AF31D27A6" blockId="2.[160,1124,1409,1460]" pageId="2" pageNumber="3">
<emphasis id="BFFE13485905614ACDBFFA6AF0632784" bold="true" box="[160,290,1409,1431]" pageId="2" pageNumber="3">Figures 1–3.</emphasis> <emphasis id="BFFE13485905614ACDBFFA6AF0632784" bold="true" box="[160,290,1409,1431]" pageId="2" pageNumber="3">Figures 1–3.</emphasis>
Photomicrographs of male of Photomicrographs of male of
@ -179,7 +182,7 @@ Dorsal habitus.
Facial view. Facial view.
</paragraph> </paragraph>
</caption> </caption>
<caption id="D9F59FD25904614BCDBEFB23F33B270A" pageId="3" pageNumber="4" startId="3.[161,242,1224,1246]" targetBox="[194,1081,244,1200]" targetPageId="3" targetType="figure"> <caption id="D9F59FD25904614BCDBEFB23F33B270A" ID-DOI="http://doi.org/10.5281/zenodo.13737332" ID-Zenodo-Dep="13737332" httpUri="https://zenodo.org/record/13737332/files/figure.png" pageId="3" pageNumber="4" startId="3.[161,242,1224,1246]" targetBox="[194,1081,244,1200]" targetPageId="3" targetType="figure">
<paragraph id="8D35CF5A5904614BCDBEFB23F33B270A" blockId="3.[161,1143,1224,1304]" pageId="3" pageNumber="4"> <paragraph id="8D35CF5A5904614BCDBEFB23F33B270A" blockId="3.[161,1143,1224,1304]" pageId="3" pageNumber="4">
<emphasis id="BFFE13485904614BCDBEFB23F06226CC" bold="true" box="[161,291,1224,1246]" pageId="3" pageNumber="4">Figures 4–8.</emphasis> <emphasis id="BFFE13485904614BCDBEFB23F06226CC" bold="true" box="[161,291,1224,1246]" pageId="3" pageNumber="4">Figures 4–8.</emphasis>
Male terminalia of Male terminalia of
@ -203,10 +206,10 @@ Genital capsule, dorsal view.
</caption> </caption>
<paragraph id="8D35CF5A5904614CCDCFFA66F53F24B2" blockId="3.[160,1175,1357,1797]" lastBlockId="4.[160,1168,1512,1792]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4"> <paragraph id="8D35CF5A5904614CCDCFFA66F53F24B2" blockId="3.[160,1175,1357,1797]" lastBlockId="4.[160,1168,1512,1792]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Integument generally shining ( Integument generally shining (
<figureCitation id="15B1D3DF5904614BCF56FA66F3F727B7" box="[585,694,1421,1445]" captionStart="Figures 1–3" captionStartId="2.[160,241,1409,1431]" captionTargetBox="[163,1137,241,1385]" captionTargetId="figure-110@2.[160,1168,224,1400]" captionTargetPageId="2" captionText="Figures 1–3. Photomicrographs of male of Chiasmognathus scythicus, new species. 1. Lateral habitus. 2. Dorsal habitus. 3. Facial view." pageId="3" pageNumber="4">Figs. 1–3</figureCitation> <figureCitation id="15B1D3DF5904614BCF56FA66F3F727B7" box="[585,694,1421,1445]" captionStart="Figures 1–3" captionStartId="2.[160,241,1409,1431]" captionTargetBox="[163,1137,241,1385]" captionTargetId="figure-110@2.[160,1168,224,1400]" captionTargetPageId="2" captionText="Figures 1–3. Photomicrographs of male of Chiasmognathus scythicus, new species. 1. Lateral habitus. 2. Dorsal habitus. 3. Facial view." figureDoi="http://doi.org/10.5281/zenodo.13737330" httpUri="https://zenodo.org/record/13737330/files/figure.png" pageId="3" pageNumber="4">Figs. 1–3</figureCitation>
). Labrum with punctures irregular in size and spacing, surface smooth; clypeus with small punctures separated by 0.5–2 times a puncture width centrally, punctures closer laterally, with a few scattered larger punctures, apex impunctate; face and vertex with small punctures separated by 0.25–2 times a puncture width (in a few places where punctures are particularly close they can be nearly contiguous), integument between punctures smooth, punctures on vertex posterior to ocelli somewhat weaker; punctures on gena and postgena separated by 0.5–2 times a puncture width, integument otherwise smooth and shining. Mesoscutum sparsely punctate on central disc, punctures small and separated by 1–4.5 times a puncture width, punctures closer on margins, integument otherwise smooth and shining; mesoscutellum with punctures slightly larger and deeper than those of mesoscutum, separated by 0.5–2 times a puncture width; metanotum with punctures separated by a puncture width; pleura with small punctures separated by 0.5–2.5 times a puncture width, integument otherwise smooth; propodeum with short and narrow basal area coarsely imbricate and impunctate, otherwise integument with punctures separated by less than or equal to a puncture width except medially around pit on posterior surface smooth. Metasomal terga and sterna faintly and finely imbricate, with fine, scattered punctures (T1 and T2 almost devoid of sculpture in some specimens). ). Labrum with punctures irregular in size and spacing, surface smooth; clypeus with small punctures separated by 0.5–2 times a puncture width centrally, punctures closer laterally, with a few scattered larger punctures, apex impunctate; face and vertex with small punctures separated by 0.25–2 times a puncture width (in a few places where punctures are particularly close they can be nearly contiguous), integument between punctures smooth, punctures on vertex posterior to ocelli somewhat weaker; punctures on gena and postgena separated by 0.5–2 times a puncture width, integument otherwise smooth and shining. Mesoscutum sparsely punctate on central disc, punctures small and separated by 1–4.5 times a puncture width, punctures closer on margins, integument otherwise smooth and shining; mesoscutellum with punctures slightly larger and deeper than those of mesoscutum, separated by 0.5–2 times a puncture width; metanotum with punctures separated by a puncture width; pleura with small punctures separated by 0.5–2.5 times a puncture width, integument otherwise smooth; propodeum with short and narrow basal area coarsely imbricate and impunctate, otherwise integument with punctures separated by less than or equal to a puncture width except medially around pit on posterior surface smooth. Metasomal terga and sterna faintly and finely imbricate, with fine, scattered punctures (T1 and T2 almost devoid of sculpture in some specimens).
</paragraph> </paragraph>
<caption id="D9F59FD25903614CCDBFFA69F33527A7" pageId="4" pageNumber="5" startId="4.[160,241,1410,1432]" targetBox="[172,1135,246,1379]" targetPageId="4" targetType="figure"> <caption id="D9F59FD25903614CCDBFFA69F33527A7" ID-DOI="http://doi.org/10.5281/zenodo.13737334" ID-Zenodo-Dep="13737334" httpUri="https://zenodo.org/record/13737334/files/figure.png" pageId="4" pageNumber="5" startId="4.[160,241,1410,1432]" targetBox="[172,1135,246,1379]" targetPageId="4" targetType="figure">
<paragraph id="8D35CF5A5903614CCDBFFA69F33527A7" blockId="4.[160,1156,1410,1461]" pageId="4" pageNumber="5"> <paragraph id="8D35CF5A5903614CCDBFFA69F33527A7" blockId="4.[160,1156,1410,1461]" pageId="4" pageNumber="5">
<emphasis id="BFFE13485903614CCDBFFA69F06F2785" bold="true" box="[160,302,1410,1432]" pageId="4" pageNumber="5">Figures 9–11.</emphasis> <emphasis id="BFFE13485903614CCDBFFA69F06F2785" bold="true" box="[160,302,1410,1432]" pageId="4" pageNumber="5">Figures 9–11.</emphasis>
Photomicrographs of female of Photomicrographs of female of
@ -245,7 +248,7 @@ Facial view.
<paragraph id="8D35CF5A5902614DCDCFFC8AF350218B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">Sculpturing as in male except punctures of mesoscutellum closer, separated by 0.5–1.5 times a puncture width.</paragraph> <paragraph id="8D35CF5A5902614DCDCFFC8AF350218B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">Sculpturing as in male except punctures of mesoscutellum closer, separated by 0.5–1.5 times a puncture width.</paragraph>
<paragraph id="8D35CF5A5902614DCDCFFC4AF305260B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6"> <paragraph id="8D35CF5A5902614DCDCFFC4AF305260B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">
Coloration as in male except sometimes lighter brown throughout on those areas where male is black, especially face below antennae ( Coloration as in male except sometimes lighter brown throughout on those areas where male is black, especially face below antennae (
<figureCitation id="15B1D3DF5902614DCE5FFC2AF2F921CB" box="[832,952,961,985]" captionStart="Figures 9–11" captionStartId="4.[160,241,1410,1432]" captionTargetBox="[172,1135,246,1379]" captionTargetId="figure-128@4.[160,1168,224,1400]" captionTargetPageId="4" captionText="Figures 9–11. Photomicrographs of female of Chiasmognathus scythicus, new species. 9. Lateral habitus. 10. Dorsal habitus. 11. Facial view." pageId="5" pageNumber="6">Figs. 9–11</figureCitation> <figureCitation id="15B1D3DF5902614DCE5FFC2AF2F921CB" box="[832,952,961,985]" captionStart="Figures 9–11" captionStartId="4.[160,241,1410,1432]" captionTargetBox="[172,1135,246,1379]" captionTargetId="figure-128@4.[160,1168,224,1400]" captionTargetPageId="4" captionText="Figures 9–11. Photomicrographs of female of Chiasmognathus scythicus, new species. 9. Lateral habitus. 10. Dorsal habitus. 11. Facial view." figureDoi="http://doi.org/10.5281/zenodo.13737334" httpUri="https://zenodo.org/record/13737334/files/figure.png" pageId="5" pageNumber="6">Figs. 9–11</figureCitation>
), and nearly black around ocellar triangle regardless of background color of integument (lighter colors likely due to fading of integument). ), and nearly black around ocellar triangle regardless of background color of integument (lighter colors likely due to fading of integument).
</paragraph> </paragraph>
<paragraph id="8D35CF5A5902614DCDCFFBCAF2F0262B" blockId="5.[160,1169,225,1786]" box="[208,945,1057,1081]" pageId="5" pageNumber="6">Pubescence more dense on clypeus and lower paraocular area.</paragraph> <paragraph id="8D35CF5A5902614DCDCFFBCAF2F0262B" blockId="5.[160,1169,225,1786]" box="[208,945,1057,1081]" pageId="5" pageNumber="6">Pubescence more dense on clypeus and lower paraocular area.</paragraph>
@ -288,15 +291,11 @@ Coloration as in male except sometimes lighter brown throughout on those areas w
<collectionCode id="EB9B579F5902614DCE79FB8AF2F4266B" box="[870,949,1121,1145]" pageId="5" pageNumber="6">KASC</collectionCode> <collectionCode id="EB9B579F5902614DCE79FB8AF2F4266B" box="[870,949,1121,1145]" pageId="5" pageNumber="6">KASC</collectionCode>
). ).
<location id="885599815902614DCED3FB8AF2B8266B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DCED3FB8AF2B8266B" box="[972,1017,1121,1145]" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="The" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">The</location> <location id="885599815902614DCED3FB8AF2B8266B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DCED3FB8AF2B8266B" box="[972,1017,1121,1145]" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="The" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">The</location>
type locality is in southeastern type locality is in southeastern Kazakhstan east of the
<collectingCountry id="F59D8FCA5902614DCC67FB6AF342268B" box="[376,515,1153,1177]" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
east of the
<location id="885599815902614DCF94FB6AF2C6268B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DCF94FB6AF2C6268B" box="[651,903,1153,1177]" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="Kapshagay Reservoir" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">Kapshagay Reservoir</location> <location id="885599815902614DCF94FB6AF2C6268B" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DCF94FB6AF2C6268B" box="[651,903,1153,1177]" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="Kapshagay Reservoir" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">Kapshagay Reservoir</location>
and around the and around the
<location id="885599815902614DC953FB6AF0CA26AB" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DC953FB6AF0CA26AB" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="Altyn Emel National Park" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">Altyn Emel National Park</location> <location id="885599815902614DC953FB6AF0CA26AB" LSID="urn:lsid:plazi:treatment:05237E4C5906614ECC51FC0AF0E923AB:885599815902614DC953FB6AF0CA26AB" country="Kazakhstan" county="Tauqum Sands" latitude="44.0343" longLatPrecision="6" longitude="79.5184" municipality="Aydarly" name="Altyn Emel National Park" pageId="5" pageNumber="6" stateProvince="Almaty Oblysy">Altyn Emel National Park</location>
, and generally part of the xeric to mountainous ecoregion of , and generally part of the xeric to mountainous ecoregion of Kazakhstan.
<collectingCountry id="F59D8FCA5902614DC979FB49F04C26CB" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
.
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<paragraph id="8D35CF5A5902614DCDCFFB0AF2B5240B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6"> <paragraph id="8D35CF5A5902614DCDCFFB0AF2B5240B" blockId="5.[160,1169,225,1786]" pageId="5" pageNumber="6">