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data/03/99/87/039987A13873FFAD29F4F96CFAEBFC9D.xml
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<mods:title id="E0A300BBD300B92B9692C4E5F0E33B35">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
<document id="C9656DA26FA78BF521BF5B6B1C4825D3" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-ISSN="2118-9773" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.bibliography_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A13873FFAD29F4F96CFAEBFC9D" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium Motschulsky 1855" docType="key" docVersion="13" lastPageNumber="100" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="98" updateTime="1740685105051" updateUser="guilherme" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B301427DDB8067F66B819129E271B70C">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
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<treatment id="039987A13873FFAD29F4F96CFAEBFC9D" ID-DOI="http://doi.org/10.5281/zenodo.3795654" ID-GBIF-Taxon="163877766" ID-Zenodo-Dep="3795654" LSID="urn:lsid:plazi:treatment:039987A13873FFAD29F4F96CFAEBFC9D" httpUri="http://treatment.plazi.org/id/039987A13873FFAD29F4F96CFAEBFC9D" lastPageId="99" lastPageNumber="100" pageId="97" pageNumber="98">
<treatment id="039987A13873FFAD29F4F96CFAEBFC9D" ID-DOI="http://doi.org/10.5281/zenodo.3795654" ID-GBIF-Taxon="163877766" ID-Zenodo-Dep="3795654" LSID="urn:lsid:plazi:treatment:039987A13873FFAD29F4F96CFAEBFC9D" httpUri="http://treatment.plazi.org/id/039987A13873FFAD29F4F96CFAEBFC9D" lastPageId="99" lastPageNumber="100" pageId="97" pageNumber="98" scope_family="Ptiliidae" scope_order="Coleoptera" scope_subFamily="Ptiliinae" scope_tribe="Discheramocephalini">
<subSubSection id="C32A653C3873FFAF29F4F96CFD15F904" box="[189,649,1666,1693]" pageId="97" pageNumber="98" type="nomenclature">
<paragraph id="8B8F36B73873FFAF29F4F96CFD15F904" blockId="97.[189,649,1666,1693]" box="[189,649,1666,1693]" pageId="97" pageNumber="98">
<heading id="D0C781DB3873FFAF29F4F96CFD15F904" bold="true" box="[189,649,1666,1693]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="B944EAA53873FFAF29F4F96CFD15F904" bold="true" box="[189,649,1666,1693]" pageId="97" pageNumber="98">
Key to
<taxonomicName id="4C304D343873FFAF285EF96CFE0CF905" ID-CoL="8X77B" box="[279,400,1666,1693]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<taxonomicName id="4C304D343873FFAF285EF96CFE0CF905" ID-CoL="8X77B" authorityName="Motschulsky" authorityYear="1855" box="[279,400,1666,1693]" class="Insecta" family="Ptiliidae" genus="Cissidium" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAA53873FFAF285EF96CFE0CF905" bold="true" box="[279,400,1666,1693]" italics="true" pageId="97" pageNumber="98">Cissidium</emphasis>
</taxonomicName>
species in group 4A
@ -51,123 +54,160 @@ species in group 4A
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<key id="FFA161863873FFAD29F4F95DFAEBFC9D" lastPageId="99" lastPageNumber="100" pageId="97" pageNumber="98">
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<keyStep id="30C42E123873FFAF29F4F95DFAEBF96B" pageId="97" pageNumber="98">
<keyLead id="30C195823873FFAF29F4F95DFAEBF96B" pageId="97" pageNumber="98">1. Species with longitudinal linear depressions on the pronotum ......................................................... 2 – Species without linear depressions on the pronotum......................................................................... 4</keyLead>
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<keyLead id="30C195823873FFAF29F4F95DFAEBF955" box="[189,1399,1715,1742]" pageId="97" pageNumber="98">1. Species with longitudinal linear depressions on the pronotum ......................................................... 2</keyLead>
</paragraph>
<paragraph id="8B8F36B73873FFAF29F4F8F2FAEBF8C4" blockId="97.[189,1399,1820,1884]" pageId="97" pageNumber="98">
<paragraph id="8B8F36B73873FFAF2988F937FAEBF96B" blockId="97.[189,1399,1715,1779]" box="[193,1399,1752,1779]" pageId="97" pageNumber="98">
<keyLead id="30C195823873FFAF2988F937FAEBF96B" box="[193,1399,1752,1779]" pageId="97" pageNumber="98">– Species without linear depressions on the pronotum......................................................................... 4</keyLead>
</paragraph>
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<keyLead id="30C195823873FFAF29F4F8F2FAEBF8C4" pageId="97" pageNumber="98">
<paragraph id="8B8F36B73873FFAF29F4F8F2FAEBF8AF" blockId="97.[189,1399,1820,1884]" box="[189,1399,1820,1847]" pageId="97" pageNumber="98">
<keyLead id="30C195823873FFAF29F4F8F2FAEBF8AF" box="[189,1399,1820,1847]" pageId="97" pageNumber="98">
2. Linear depressions very faint (best seen with side lighting)..................................
<taxonomicName id="4C304D343873FFAF2DD0F8F2FA89F8AF" ID-CoL="8XXRX" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1177,1301,1820,1847]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="trangse" status="sp. nov.">
<emphasis id="B944EAA53873FFAF2DD0F8F2FA89F8AF" bold="true" box="[1177,1301,1820,1847]" italics="true" pageId="97" pageNumber="98">C. trangse</emphasis>
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– Linear depressions well marked ........................................................................................................ 3
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<paragraph id="8B8F36B73873FFAF29F4F868FAEBF873" blockId="97.[189,1399,1926,2027]" pageId="97" pageNumber="98">
<paragraph id="8B8F36B73873FFAF2988F8ACFAEBF8C4" blockId="97.[189,1399,1820,1884]" box="[193,1399,1858,1884]" pageId="97" pageNumber="98">
<keyLead id="30C195823873FFAF2988F8ACFAEBF8C4" box="[193,1399,1858,1884]" pageId="97" pageNumber="98">– Linear depressions well marked ........................................................................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123873FFAF29F4F868FAEBF873" pageId="97" pageNumber="98">
<keyLead id="30C195823873FFAF29F4F868FAEBF873" pageId="97" pageNumber="98">
<paragraph id="8B8F36B73873FFAF29F4F868FAEBF85E" blockId="97.[189,1399,1926,2027]" pageId="97" pageNumber="98">
<keyLead id="30C195823873FFAF29F4F868FAEBF85E" pageId="97" pageNumber="98">
3. Linear depressions long almost reaching mid-point of pronotum, pronotum more densely pubescent ..........................................................................................................
<taxonomicName id="4C304D343873FFAF2D39F845FA89F85E" ID-CoL="8XXPH" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1136,1301,1963,1990]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="bilineatum" status="sp. nov.">
<emphasis id="B944EAA53873FFAF2D39F845FA89F85E" bold="true" box="[1136,1301,1963,1990]" italics="true" pageId="97" pageNumber="98">C. bilineatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3873FFAF2C55F842FAEBF85E" box="[1308,1399,1964,1990]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B8F36B73873FFAF2988F83FFAEBF873" blockId="97.[189,1399,1926,2027]" box="[193,1399,2001,2027]" pageId="97" pageNumber="98">
<keyLead id="30C195823873FFAF2988F83FFAEBF873" box="[193,1399,2001,2027]" pageId="97" pageNumber="98">
– Linear depressions shorter, pronotum more sparsely pubescent .......................
<taxonomicName id="4C304D343873FFAF2D34F83FFA89F873" ID-CoL="8XXPC" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1149,1301,2001,2027]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="arcuatum" status="sp. nov.">
<emphasis id="B944EAA53873FFAF2D34F83FFA89F873" bold="true" box="[1149,1301,2001,2027]" italics="true" pageId="97" pageNumber="98">C. arcuatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3873FFAF2C55F83FFAEBF873" box="[1308,1399,2001,2027]" pageId="97" pageNumber="98" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8B8F36B73870FFAC29F4FEE5FAEBFED2" blockId="98.[189,1399,267,330]" pageId="98" pageNumber="99">
</keyStep>
<keyStep id="30C42E123870FFAC29F4FEE5FAEBFED2" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FEE5FAEBFED2" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4FEE5FAEBFEBD" blockId="98.[189,1399,267,330]" box="[189,1399,267,294]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FEE5FAEBFEBD" box="[189,1399,267,294]" pageId="98" pageNumber="99">
4. Mentum triangular ...................................................................................
<taxonomicName id="4C304D343870FFAC2D7AFEE5FAEBFEBD" ID-CoL="8XXRY" authority="Darby, 2019" authorityName="Darby" authorityYear="2019" box="[1075,1399,267,294]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="triangulum">
<emphasis id="B944EAA53870FFAC2D7AFEE5FB42FEBE" bold="true" box="[1075,1246,267,294]" italics="true" pageId="98" pageNumber="99">C. triangulum</emphasis>
<bibRefCitation id="EFA14B463870FFAC2DAFFEE5FAEBFEBD" author="Darby M." box="[1254,1399,267,293]" pageId="98" pageNumber="99" pagination="1 - 50" refId="ref59424" refString="Darby M. 2019. New Ptiliidae (Coleoptera) from Sarawak in the spirit collection of the Natural History Museum, London. European Journal of Taxonomy 512: 1 - 50. https: // doi. org / 10.5852 / ejt. 2019.512" type="journal article" year="2019">Darby, 2019</bibRefCitation>
</taxonomicName>
– Mentum not triangular ....................................................................................................................... 5
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FE9EFAEBFE4A" blockId="98.[189,1399,368,466]" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC2988FEC1FAEBFED2" blockId="98.[189,1399,267,330]" box="[193,1399,303,330]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FEC1FAEBFED2" box="[193,1399,303,330]" pageId="98" pageNumber="99">– Mentum not triangular ....................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123870FFAC29F4FE9EFAEBFE4A" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FE9EFAEBFE4A" pageId="98" pageNumber="99">5. Head small, pronotum domed, sides almost parallel before hind angles, base of pronotum wider than elytra ................................................................................................................................................. 6 – Pronotum not so domed, not wider than elytra, head larger in proportion ........................................ 7</keyLead>
</keyStep>
<paragraph id="8B8F36B73870FFAC29F4FE9EFAEBFE36" blockId="98.[189,1399,368,466]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FE9EFAEBFE36" pageId="98" pageNumber="99">5. Head small, pronotum domed, sides almost parallel before hind angles, base of pronotum wider than elytra ................................................................................................................................................. 6</keyLead>
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<paragraph id="8B8F36B73870FFAC29F4FE17FAEBFDAF" blockId="98.[189,1399,504,567]" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC2988FE56FAEBFE4A" blockId="98.[189,1399,368,466]" box="[193,1399,439,466]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FE56FAEBFE4A" box="[193,1399,439,466]" pageId="98" pageNumber="99">– Pronotum not so domed, not wider than elytra, head larger in proportion ........................................ 7</keyLead>
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<keyStep id="30C42E123870FFAC29F4FE17FAEBFDAF" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FE17FAEBFDAF" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4FE17FAEBFD8B" blockId="98.[189,1399,504,567]" box="[189,1399,504,531]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FE17FAEBFD8B" box="[189,1399,504,531]" pageId="98" pageNumber="99">
6. Pronotal hind angles acute, projected rearwards ..............................................
<taxonomicName id="4C304D343870FFAC2D3BFE17FA89FD8B" ID-CoL="8XXRN" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1138,1301,504,531]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="seditiosum" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2D3BFE17FA89FD8B" bold="true" box="[1138,1301,504,531]" italics="true" pageId="98" pageNumber="99">C. seditiosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55FE17FAEBFD8B" box="[1308,1399,505,531]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC2988FDF3FAEBFDAF" blockId="98.[189,1399,504,567]" box="[193,1399,540,567]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FDF3FAEBFDAF" box="[193,1399,540,567]" pageId="98" pageNumber="99">
– Pronotal hind angles rounded, not projected rearward .........................................
<taxonomicName id="4C304D343870FFAC2DC6FDF2FA88FDAF" ID-CoL="8XXQD" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1167,1300,540,567]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="insulare" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2DC6FDF2FA88FDAF" bold="true" box="[1167,1300,540,567]" italics="true" pageId="98" pageNumber="99">C. insulare</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55FDF3FAEBFDAF" box="[1308,1399,541,567]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FDB0FAEBFD27" blockId="98.[189,1399,606,703]" pageId="98" pageNumber="99">
</keyStep>
<keyStep id="30C42E123870FFAC29F4FDB0FAEBFD27" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FDB0FAEBFD27" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4FDB0FAEBFD04" blockId="98.[189,1399,606,703]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FDB0FAEBFD04" pageId="98" pageNumber="99">
7. Pronotum cordate, posterior margin same width as distance across the eyes, elytra almost parallelsided, very small species.....................................................................................
<taxonomicName id="4C304D343870FFAC2DC8FD6FFA89FD04" ID-CoL="8XXPF" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1153,1301,641,668]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="besucheti" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2DC8FD6FFA89FD04" bold="true" box="[1153,1301,641,668]" italics="true" pageId="98" pageNumber="99">C. besucheti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55FD6CFAEBFD04" box="[1308,1399,642,668]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
– Larger species, pronotal posterior margin clearly wider than distance across eyes .......................... 8
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FD08FAEAFCBC" blockId="98.[189,1399,742,804]" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC2988FD4BFAEBFD27" blockId="98.[189,1399,606,703]" box="[193,1399,677,703]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FD4BFAEBFD27" box="[193,1399,677,703]" pageId="98" pageNumber="99">– Larger species, pronotal posterior margin clearly wider than distance across eyes .......................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123870FFAC29F4FD08FAEAFCBC" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FD08FAEAFCBC" pageId="98" pageNumber="99">8. Posterior margins of pronotum almost parallel-sided ....................................................................... 9 – Posterior margins of pronotum clearly concave before hind angles................................................ 10</keyLead>
</keyStep>
<paragraph id="8B8F36B73870FFAC29F4FD08FAEBFC98" blockId="98.[189,1399,742,804]" box="[189,1399,742,769]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FD08FAEBFC98" box="[189,1399,742,769]" pageId="98" pageNumber="99">8. Posterior margins of pronotum almost parallel-sided ....................................................................... 9</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FCA2FAEBFC12" blockId="98.[189,1399,843,906]" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC2988FCE4FAEAFCBC" blockId="98.[189,1399,742,804]" box="[193,1398,778,804]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FCE4FAEAFCBC" box="[193,1398,778,804]" pageId="98" pageNumber="99">– Posterior margins of pronotum clearly concave before hind angles................................................ 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123870FFAC29F4FCA2FAEBFC12" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FCA2FAEBFC12" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4FCA2FAEBFCFE" blockId="98.[189,1399,843,906]" box="[189,1399,843,870]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FCA2FAEBFCFE" box="[189,1399,843,870]" pageId="98" pageNumber="99">
9. Mesoventral lateral margins serrate..........................................................................
<taxonomicName id="4C304D343870FFAC2DF9FCA5FA89FCFE" ID-CoL="8XXQJ" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1200,1301,843,870]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="loebli" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2DF9FCA5FA89FCFE" bold="true" box="[1200,1301,843,870]" italics="true" pageId="98" pageNumber="99">C. loebli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55FCA2FAEBFCFE" box="[1308,1399,844,870]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC2988FC81FAEBFC12" blockId="98.[189,1399,843,906]" box="[193,1399,878,906]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FC81FAEBFC12" box="[193,1399,878,906]" pageId="98" pageNumber="99">
– Mesoventral lateral margins without serrations..........................................
<taxonomicName id="4C304D343870FFAC2D0EFC81FAEBFC12" ID-CoL="5Z8SB" authority="Darby, 2015" authorityName="Darby" authorityYear="2015" box="[1095,1399,878,906]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="globosum">
<emphasis id="B944EAA53870FFAC2D0EFC81FB42FC11" bold="true" box="[1095,1246,878,905]" italics="true" pageId="98" pageNumber="99">C. globosum</emphasis>
<bibRefCitation id="EFA14B463870FFAC2DAFFC81FAEBFC12" author="Darby M." box="[1254,1399,879,906]" pageId="98" pageNumber="99" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby, 2015</bibRefCitation>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FC5FFAEAFC76" blockId="98.[189,1398,944,1006]" pageId="98" pageNumber="99">
</keyStep>
<keyStep id="30C42E123870FFAC29F4FC5FFAEAFC76" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FC5FFAEAFC76" pageId="98" pageNumber="99">10. Mid-keel rounded/tapering posteriorly, without projecting hind angles...........................................11 – Mid-keel with clearly projecting posterior angles........................................................................... 14</keyLead>
</keyStep>
<paragraph id="8B8F36B73870FFAC29F4FC5FFAE9FC53" blockId="98.[189,1398,944,1006]" box="[189,1397,944,971]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FC5FFAE9FC53" box="[189,1397,944,971]" pageId="98" pageNumber="99">10. Mid-keel rounded/tapering posteriorly, without projecting hind angles...........................................11</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FBF8FAEAFBCB" blockId="98.[189,1398,1045,1107]" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC2988FC3AFAEAFC76" blockId="98.[189,1398,944,1006]" box="[193,1398,980,1006]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FC3AFAEAFC76" box="[193,1398,980,1006]" pageId="98" pageNumber="99">– Mid-keel with clearly projecting posterior angles........................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123870FFAC29F4FBF8FAEAFBCB" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FBF8FAEAFBCB" pageId="98" pageNumber="99">11. Mid-keel very narrow parallel-sided then tapering to keel.............................................................. 12 – Mid-keel wider, posterior angles rounded ....................................................................................... 13</keyLead>
</keyStep>
<paragraph id="8B8F36B73870FFAC29F4FBF8FAEAFBA8" blockId="98.[189,1398,1045,1107]" box="[189,1398,1045,1072]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FBF8FAEAFBA8" box="[189,1398,1045,1072]" pageId="98" pageNumber="99">11. Mid-keel very narrow parallel-sided then tapering to keel.............................................................. 12</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FB95FAEBFB67" blockId="98.[189,1399,1146,1279]" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC2988FBD7FAEAFBCB" blockId="98.[189,1398,1045,1107]" box="[193,1398,1081,1107]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FBD7FAEAFBCB" box="[193,1398,1081,1107]" pageId="98" pageNumber="99">– Mid-keel wider, posterior angles rounded ....................................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123870FFAC29F4FB95FAEBFB67" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FB95FAEBFB67" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4FB95FAEBFB20" blockId="98.[189,1399,1146,1279]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FB95FAEBFB20" pageId="98" pageNumber="99">
12. Mid-keel tapering to keel from end of the very narrow medial process of the collar ......................... ............................................................................................................................
<taxonomicName id="4C304D343870FFAC2D36FB70FA89FB21" ID-CoL="8XXS2" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1151,1301,1182,1209]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="visendum" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2D36FB70FA89FB21" bold="true" box="[1151,1301,1182,1209]" italics="true" pageId="98" pageNumber="99">C. visendum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55FB70FAEBFB20" box="[1308,1399,1182,1208]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC2988FB2CFAEBFB67" blockId="98.[189,1399,1146,1279]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FB2CFAEBFB67" pageId="98" pageNumber="99">
<emphasis id="B944EAA53870FFAC2988FB2CFF4CFB44" bold="true" box="[193,208,1218,1244]" italics="true" pageId="98" pageNumber="99">–</emphasis>
Mid-keel parallel-sided for most of its length before tapering to join keel ......................................... ........................................................................................................................
<taxonomicName id="4C304D343870FFAC2D26FB0BFA89FB67" ID-CoL="8XXRG" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1135,1301,1252,1279]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="plaumanni" status="sp. nov.">
@ -175,69 +215,84 @@ Mid-keel parallel-sided for most of its length before tapering to join keel ....
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55FB0BFAEBFB67" box="[1308,1399,1253,1279]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FAC9FAEBFA10" blockId="98.[189,1399,1318,1416]" pageId="98" pageNumber="99">
</keyStep>
<keyStep id="30C42E123870FFAC29F4FAC9FAEBFA10" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FAC9FAEBFA10" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4FAC9FAEBFAFC" blockId="98.[189,1399,1318,1416]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FAC9FAEBFAFC" pageId="98" pageNumber="99">
13. Mesoventral collar with two short posteriorly directed processes either side of the medial process ..................................................................................................................
<taxonomicName id="4C304D343870FFAC2DDAFAA4FA89FAFC" ID-CoL="8XXR8" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1171,1301,1354,1380]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="opacum" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2DDAFAA4FA89FAFC" bold="true" box="[1171,1301,1354,1380]" italics="true" pageId="98" pageNumber="99">C. opacum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55FAA4FAEBFAFC" box="[1308,1399,1354,1380]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC2988FA80FAEBFA10" blockId="98.[189,1399,1318,1416]" box="[193,1399,1389,1416]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FA80FAEBFA10" box="[193,1399,1389,1416]" pageId="98" pageNumber="99">
– Collar without processes either side of the medial process .......................................
<taxonomicName id="4C304D343870FFAC2DFCFA83FA89FA10" ID-CoL="8XXRR" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1205,1301,1389,1416]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="steeli" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2DFCFA83FA89FA10" bold="true" box="[1205,1301,1389,1416]" italics="true" pageId="98" pageNumber="99">C. steeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55FA80FAEBFA10" box="[1308,1399,1390,1416]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4FA41FAEAFA74" blockId="98.[189,1398,1455,1517]" pageId="98" pageNumber="99">
</keyStep>
<keyStep id="30C42E123870FFAC29F4FA41FAEAFA74" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FA41FAEAFA74" pageId="98" pageNumber="99">14. Mesoventral lateral borders effaced in posterior half ..................................................................... 15. – Mesoventral lateral borders distinct to mesocoxae......................................................................... 16.</keyLead>
</keyStep>
<paragraph id="8B8F36B73870FFAC29F4FA41FAEAFA52" blockId="98.[189,1398,1455,1517]" box="[189,1398,1455,1482]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4FA41FAEAFA52" box="[189,1398,1455,1482]" pageId="98" pageNumber="99">14. Mesoventral lateral borders effaced in posterior half ..................................................................... 15.</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4F9FAFAEBF9CA" blockId="98.[189,1399,1555,1618]" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC2988FA3DFAEAFA74" blockId="98.[189,1398,1455,1517]" box="[193,1398,1490,1517]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988FA3DFAEAFA74" box="[193,1398,1490,1517]" pageId="98" pageNumber="99">– Mesoventral lateral borders distinct to mesocoxae......................................................................... 16.</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123870FFAC29F4F9FAFAEBF9CA" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4F9FAFAEBF9CA" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4F9FAFAEBF9B6" blockId="98.[189,1399,1555,1618]" box="[189,1399,1555,1583]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4F9FAFAEBF9B6" box="[189,1399,1555,1583]" pageId="98" pageNumber="99">
15. Pronotum widest in basal half, pubescence sparser..........................................
<taxonomicName id="4C304D343870FFAC2D3DF9FAFA89F9B6" ID-CoL="8XXQP" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1140,1301,1555,1582]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="luctuosum" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2D3DF9FAFA89F9B6" bold="true" box="[1140,1301,1555,1582]" italics="true" pageId="98" pageNumber="99">C. luctuosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55F9FAFAEBF9B6" box="[1308,1399,1556,1582]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B73870FFAC2988F9D6FAEBF9CA" blockId="98.[189,1399,1555,1618]" box="[193,1399,1591,1618]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988F9D6FAEBF9CA" box="[193,1399,1591,1618]" pageId="98" pageNumber="99">
– Pronotum widest medially pubescence denser ....................................................
<taxonomicName id="4C304D343870FFAC2DC6F9D9FA88F9C9" ID-CoL="8XXPZ" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1167,1300,1591,1617]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="faustum" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2DC6F9D9FA88F9C9" bold="true" box="[1167,1300,1591,1617]" italics="true" pageId="98" pageNumber="99">C. faustum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55F9D6FAEBF9CA" box="[1308,1399,1592,1618]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4F997FAEAF92E" blockId="98.[189,1399,1657,1719]" pageId="98" pageNumber="99">
</keyStep>
<keyStep id="30C42E123870FFAC29F4F997FAEAF92E" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4F997FAEAF92E" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4F997FAEBF90B" blockId="98.[189,1399,1657,1719]" box="[189,1399,1657,1684]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4F997FAEBF90B" box="[189,1399,1657,1684]" pageId="98" pageNumber="99">
16. Sides of mid-keel straight to posterior angles....................................................
<taxonomicName id="4C304D343870FFAC2D33F997FA89F90C" ID-CoL="8XXR6" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1146,1301,1657,1684]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="obscurum" status="sp. nov.">
<emphasis id="B944EAA53870FFAC2D33F997FA89F90C" bold="true" box="[1146,1301,1657,1684]" italics="true" pageId="98" pageNumber="99">C. obscurum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3870FFAC2C55F997FAEBF90B" box="[1308,1399,1657,1683]" pageId="98" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
– Sides of mid-keel not straight concave or slightly convex .............................................................. 17
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC29F4F930FAEAF8A7" blockId="98.[189,1399,1758,1855]" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC2988F973FAEAF92E" blockId="98.[189,1399,1657,1719]" box="[193,1398,1692,1719]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988F973FAEAF92E" box="[193,1398,1692,1719]" pageId="98" pageNumber="99">– Sides of mid-keel not straight concave or slightly convex .............................................................. 17</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123870FFAC29F4F930FAEAF8A7" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4F930FAEAF8A7" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4F930FAEBF884" blockId="98.[189,1399,1758,1855]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4F930FAEBF884" pageId="98" pageNumber="99">
17. Sides of mid keel slightly concave same width as the median extension of the collar anteriorly and strongly widening posteriorly .................................................................
<taxonomicName id="4C304D343870FFAC2D61F8EFFAEBF884" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1064,1399,1793,1820]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="elongatum">
<emphasis id="B944EAA53870FFAC2D61F8EFFB57F884" bold="true" box="[1064,1227,1793,1820]" italics="true" pageId="98" pageNumber="99">C. elongatum</emphasis>
<bibRefCitation id="EFA14B463870FFAC2D9BF8EFFAEBF884" author="Sawada Y." box="[1234,1399,1793,1820]" pageId="98" pageNumber="99" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
– Mid-keel sides concave.................................................................................................................... 18
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73870FFAC2988F8CBFAEAF8A7" blockId="98.[189,1399,1758,1855]" box="[193,1398,1829,1855]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC2988F8CBFAEAF8A7" box="[193,1398,1829,1855]" pageId="98" pageNumber="99">– Mid-keel sides concave.................................................................................................................... 18</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123870FFAC29F4F889FAEAF873" pageId="98" pageNumber="99">
<paragraph id="8B8F36B73870FFAC29F4F889FAEBF83D" blockId="98.[189,1399,1894,2027]" pageId="98" pageNumber="99">
<keyLead id="30C195823870FFAC29F4F889FAEBF83D" pageId="98" pageNumber="99">
@ -252,66 +307,82 @@ Mid-keel parallel-sided for most of its length before tapering to join keel ....
<keyLead id="30C195823870FFAC29F4F840FAEAF873" pageId="98" pageNumber="99">– Mid keel sides less pronouncedly concave wider at termination of the medial extension of the collar ................................................................................................................................................ 19</keyLead>
</paragraph>
</keyStep>
<paragraph id="8B8F36B73871FFAD29F4FEE5FAEBFEF5" blockId="99.[189,1399,267,365]" pageId="99" pageNumber="100">
<keyStep id="30C42E123871FFAD29F4FEE5FAEBFEF5" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FEE5FAEBFEF5" pageId="99" pageNumber="100">
19. Posterior border of pronotum well-marked to emargination opposite the scutellum ...................... 20 – Posterior border of pronotum very short reaching only half way to emargination opposite the scutellum..................................................................................................................
<paragraph id="8B8F36B73871FFAD29F4FEE5FAEAFEBD" blockId="99.[189,1399,267,365]" box="[189,1398,267,294]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FEE5FAEAFEBD" box="[189,1398,267,294]" pageId="99" pageNumber="100">19. Posterior border of pronotum well-marked to emargination opposite the scutellum ...................... 20</keyLead>
</paragraph>
<paragraph id="8B8F36B73871FFAD2988FEDEFAEBFEF5" blockId="99.[189,1399,267,365]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD2988FEDEFAEBFEF5" pageId="99" pageNumber="100">
– Posterior border of pronotum very short reaching only half way to emargination opposite the scutellum..................................................................................................................
<taxonomicName id="4C304D343871FFAD2DE0FEBDFA89FEF5" ID-CoL="8XXPW" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1193,1301,339,365]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="eciton" status="sp. nov.">
<emphasis id="B944EAA53871FFAD2DE0FEBDFA89FEF5" bold="true" box="[1193,1301,339,365]" italics="true" pageId="99" pageNumber="100">C. eciton</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3871FFAD2C55FEBDFAEBFEF5" box="[1308,1399,339,365]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73871FFAD29F4FE7BFAEAFE4B" blockId="99.[189,1398,405,467]" pageId="99" pageNumber="100">
</keyStep>
<keyStep id="30C42E123871FFAD29F4FE7BFAEAFE4B" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FE7BFAEAFE4B" pageId="99" pageNumber="100">20. Tip of keel sharply pointed .............................................................................................................. 21 – Tip of keel blunt/rounded................................................................................................................. 22</keyLead>
</keyStep>
<paragraph id="8B8F36B73871FFAD29F4FE7BFAEAFE37" blockId="99.[189,1398,405,467]" box="[189,1398,405,432]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FE7BFAEAFE37" box="[189,1398,405,432]" pageId="99" pageNumber="100">20. Tip of keel sharply pointed .............................................................................................................. 21</keyLead>
</paragraph>
<paragraph id="8B8F36B73871FFAD29F4FE15FAEBFDA1" blockId="99.[189,1399,507,569]" pageId="99" pageNumber="100">
<paragraph id="8B8F36B73871FFAD2988FE57FAEAFE4B" blockId="99.[189,1398,405,467]" box="[193,1398,441,467]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD2988FE57FAEAFE4B" box="[193,1398,441,467]" pageId="99" pageNumber="100">– Tip of keel blunt/rounded................................................................................................................. 22</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123871FFAD29F4FE15FAEBFDA1" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FE15FAEBFDA1" pageId="99" pageNumber="100">
<paragraph id="8B8F36B73871FFAD29F4FE15FAEBFD8D" blockId="99.[189,1399,507,569]" box="[189,1399,507,533]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FE15FAEBFD8D" box="[189,1399,507,533]" pageId="99" pageNumber="100">
21. Pronotal posterior border very narrow......................................................................
<taxonomicName id="4C304D343871FFAD2DE7FE15FA88FD8D" ID-CoL="8XXRW" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1198,1300,507,533]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="sueae" status="sp. nov.">
<emphasis id="B944EAA53871FFAD2DE7FE15FA88FD8D" bold="true" box="[1198,1300,507,533]" italics="true" pageId="99" pageNumber="100">C. sueae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3871FFAD2C55FE15FAEBFD8D" box="[1308,1399,507,533]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B73871FFAD2988FDF1FAEBFDA1" blockId="99.[189,1399,507,569]" box="[193,1399,542,569]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD2988FDF1FAEBFDA1" box="[193,1399,542,569]" pageId="99" pageNumber="100">
– Pronotal posterior border wider, clearly distinct.............................................
<taxonomicName id="4C304D343871FFAD2D2CFDF1FA89FDA1" ID-CoL="8XXRT" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1125,1301,542,569]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="subfaustum" status="sp. nov.">
<emphasis id="B944EAA53871FFAD2D2CFDF1FA89FDA1" bold="true" box="[1125,1301,542,569]" italics="true" pageId="99" pageNumber="100">C. subfaustum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3871FFAD2C55FDF1FAEBFDA1" box="[1308,1399,543,569]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73871FFAD29F4FD8FFAEAFD07" blockId="99.[189,1399,608,671]" pageId="99" pageNumber="100">
</keyStep>
<keyStep id="30C42E123871FFAD29F4FD8FFAEAFD07" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FD8FFAEAFD07" pageId="99" pageNumber="100">
<paragraph id="8B8F36B73871FFAD29F4FD8FFAEBFDE3" blockId="99.[189,1399,608,671]" box="[189,1399,608,635]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FD8FFAEBFDE3" box="[189,1399,608,635]" pageId="99" pageNumber="100">
22. Medial extension of collar tapering to a blunt point...............................................
<taxonomicName id="4C304D343871FFAD2DE9FD8FFA89FDE3" ID-CoL="8XXQ2" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1184,1301,608,635]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="flavum" status="sp. nov.">
<emphasis id="B944EAA53871FFAD2DE9FD8FFA89FDE3" bold="true" box="[1184,1301,608,635]" italics="true" pageId="99" pageNumber="100">C. flavum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3871FFAD2C55FD8FFAEBFDE3" box="[1308,1399,609,635]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
– Medial extension of collar parallel-sided ........................................................................................ 23
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8B8F36B73871FFAD29F4FD29FAEBFC9D" blockId="99.[189,1399,710,773]" pageId="99" pageNumber="100">
<paragraph id="8B8F36B73871FFAD2988FD6BFAEAFD07" blockId="99.[189,1399,608,671]" box="[193,1398,644,671]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD2988FD6BFAEAFD07" box="[193,1398,644,671]" pageId="99" pageNumber="100">– Medial extension of collar parallel-sided ........................................................................................ 23</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E123871FFAD29F4FD29FAEBFC9D" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FD29FAEBFC9D" pageId="99" pageNumber="100">
<paragraph id="8B8F36B73871FFAD29F4FD29FAEBFD79" blockId="99.[189,1399,710,773]" box="[189,1399,710,737]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD29F4FD29FAEBFD79" box="[189,1399,710,737]" pageId="99" pageNumber="100">
23. Pronotum wider, lateral margins more sharply rounded...............................
<taxonomicName id="4C304D343871FFAD2D1AFD29FA89FD79" ID-CoL="8XXPX" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1107,1301,710,737]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="ecuadoriense" status="sp. nov.">
<emphasis id="B944EAA53871FFAD2D1AFD29FA89FD79" bold="true" box="[1107,1301,710,737]" italics="true" pageId="99" pageNumber="100">C. ecuadoriense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3871FFAD2C55FD29FAEBFD79" box="[1308,1399,711,737]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B73871FFAD2988FD05FAEBFC9D" blockId="99.[189,1399,710,773]" box="[193,1399,746,773]" pageId="99" pageNumber="100">
<keyLead id="30C195823871FFAD2988FD05FAEBFC9D" box="[193,1399,746,773]" pageId="99" pageNumber="100">
– Pronotum narrower, sides more smoothly rounded ................................................
<taxonomicName id="4C304D343871FFAD2DE8FD04FA89FC9C" ID-CoL="8XXP9" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1185,1301,746,772]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="99" pageNumber="100" phylum="Arthropoda" rank="species" species="angusi" status="sp. nov.">
<emphasis id="B944EAA53871FFAD2DE8FD04FA89FC9C" bold="true" box="[1185,1301,746,772]" italics="true" pageId="99" pageNumber="100">C. angusi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE3871FFAD2C55FD05FAEBFC9D" box="[1308,1399,747,773]" pageId="99" pageNumber="100" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>

267
data/03/99/87/039987A1389AFF4729F4FEAEFAEBFC17.xml
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<treatment id="039987A1389AFF4729F4FEAEFAEBFC17" ID-DOI="http://doi.org/10.5281/zenodo.3795704" ID-GBIF-Taxon="163877765" ID-Zenodo-Dep="3795704" LSID="urn:lsid:plazi:treatment:039987A1389AFF4729F4FEAEFAEBFC17" httpUri="http://treatment.plazi.org/id/039987A1389AFF4729F4FEAEFAEBFC17" lastPageId="137" lastPageNumber="138" pageId="136" pageNumber="137">
<treatment id="039987A1389AFF4729F4FEAEFAEBFC17" ID-DOI="http://doi.org/10.5281/zenodo.3795704" ID-GBIF-Taxon="163877765" ID-Zenodo-Dep="3795704" LSID="urn:lsid:plazi:treatment:039987A1389AFF4729F4FEAEFAEBFC17" httpUri="http://treatment.plazi.org/id/039987A1389AFF4729F4FEAEFAEBFC17" lastPageId="137" lastPageNumber="138" pageId="136" pageNumber="137" scope_family="Ptiliidae" scope_genus="Cissidium" scope_order="Coleoptera" scope_subFamily="Ptiliinae" scope_tribe="Discheramocephalini">
<subSubSection id="C32A653C389AFF4629F4FEAEFE75FEC1" box="[189,489,319,346]" pageId="136" pageNumber="137" type="nomenclature">
<paragraph id="8B8F36B7389AFF4629F4FEAEFE75FEC1" blockId="136.[189,489,319,346]" box="[189,489,319,346]" pageId="136" pageNumber="137">
<heading id="D0C781DB389AFF4629F4FEAEFE75FEC1" bold="true" box="[189,489,319,346]" fontSize="11" level="3" pageId="136" pageNumber="137" reason="3">
<emphasis id="B944EAA5389AFF4629F4FEAEFE75FEC1" bold="true" box="[189,489,319,346]" pageId="136" pageNumber="137">
Key to
<taxonomicName id="4C304D34389AFF46285EFED1FEF1FEC1" ID-CoL="8X77B" box="[279,365,319,345]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">species</taxonomicName>
<taxonomicName id="4C304D34389AFF46285EFED1FEF1FEC1" ID-CoL="8X77B" authorityName="Motschulsky" authorityYear="1855" box="[279,365,319,345]" class="Insecta" family="Ptiliidae" genus="Cissidium" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="genus">species</taxonomicName>
group 4B
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C389AFF4729F4FE80FAEBFC17" lastPageId="137" lastPageNumber="138" pageId="136" pageNumber="137" type="key">
<paragraph id="8B8F36B7389AFF4629F4FE80FAEBFE34" blockId="136.[189,1399,366,428]" pageId="136" pageNumber="137">1. Mesoventral lateral margins serrate................................................................................................... 2 – Mesoventral lateral margins without serrations................................................................................. 6</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FE3AFAEBFD8A" blockId="136.[189,1399,468,530]" pageId="136" pageNumber="137">2. Pronotum lateral margins angulate .................................................................................................... 3 – Pronotum lateral margins rounded..................................................................................................... 4</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FDD7FAEBFD26" blockId="136.[189,1399,569,703]" pageId="136" pageNumber="137">
<key id="FFA16186389AFF4729F4FE80FAEBFC17" lastPageId="137" lastPageNumber="138" pageId="136" pageNumber="137">
<keyStep id="30C42E12389AFF4629F4FE80FAEBFE34" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FE80FAEBFE10" blockId="136.[189,1399,366,428]" box="[189,1399,366,393]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FE80FAEBFE10" box="[189,1399,366,393]" pageId="136" pageNumber="137">1. Mesoventral lateral margins serrate................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988FE7CFAEBFE34" blockId="136.[189,1399,366,428]" box="[193,1399,402,428]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FE7CFAEBFE34" box="[193,1399,402,428]" pageId="136" pageNumber="137">– Mesoventral lateral margins without serrations................................................................................. 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389AFF4629F4FE3AFAEBFD8A" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FE3AFAEBFE76" blockId="136.[189,1399,468,530]" box="[189,1399,468,494]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FE3AFAEBFE76" box="[189,1399,468,494]" pageId="136" pageNumber="137">2. Pronotum lateral margins angulate .................................................................................................... 3</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988FE16FAEBFD8A" blockId="136.[189,1399,468,530]" box="[193,1399,503,530]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FE16FAEBFD8A" box="[193,1399,503,530]" pageId="136" pageNumber="137">– Pronotum lateral margins rounded..................................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389AFF4629F4FDD7FAEBFD26" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FDD7FAEBFDEF" blockId="136.[189,1399,569,703]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FDD7FAEBFDEF" pageId="136" pageNumber="137">
3. Pronotum lateral margins less strongly angled, almost parallel-sided before hind angles, wider than the elytral base .........................................................................................
<taxonomicName id="4C304D34389AFF462D67FDB3FA89FDEF" ID-CoL="8XXPN" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1070,1301,604,631]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="cryptophagoides" status="sp. nov.">
<emphasis id="B944EAA5389AFF462D67FDB3FA89FDEF" bold="true" box="[1070,1301,604,631]" italics="true" pageId="136" pageNumber="137">C. cryptophagoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55FDB3FAEBFDEF" box="[1308,1399,605,631]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988FD6FFAEBFD26" blockId="136.[189,1399,569,703]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FD6FFAEBFD26" pageId="136" pageNumber="137">
– Pronotal base narrower with two linear depressions, sides strongly angled ....................................... ....................................................................................................................
<taxonomicName id="4C304D34389AFF462D01FD4AFAEBFD26" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1096,1399,676,703]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="shibatai">
<emphasis id="B944EAA5389AFF462D01FD4AFB56FD27" bold="true" box="[1096,1226,676,703]" italics="true" pageId="136" pageNumber="137">C. shibatai</emphasis>
<bibRefCitation id="EFA14B46389AFF462D9BFD4AFAEBFD26" author="Sawada Y." box="[1234,1399,676,702]" pageId="136" pageNumber="137" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FD08FAEBFCD0" blockId="136.[189,1399,742,840]" pageId="136" pageNumber="137">
</keyStep>
<keyStep id="30C42E12389AFF4629F4FD08FAEBFCD0" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FD08FAEBFCBC" blockId="136.[189,1399,742,840]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FD08FAEBFCBC" pageId="136" pageNumber="137">
4. Elytra narrow, pronotum lateral margins almost parallel-sided base without linear depressions........ ........................................................................................................................
<taxonomicName id="4C304D34389AFF462D28FCE4FAEBFCBC" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1121,1399,777,804]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="sakaii">
<emphasis id="B944EAA5389AFF462D28FCE4FB57FCBC" bold="true" box="[1121,1227,777,804]" italics="true" pageId="136" pageNumber="137">C. sakaii</emphasis>
<bibRefCitation id="EFA14B46389AFF462D9BFCE4FAEBFCBC" author="Sawada Y." box="[1234,1399,778,804]" pageId="136" pageNumber="137" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
– Elytra rounded ................................................................................................................................... 5
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FC81FAEBFC35" blockId="136.[189,1399,879,941]" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF462988FCC0FAEBFCD0" blockId="136.[189,1399,742,840]" box="[193,1399,813,840]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FCC0FAEBFCD0" box="[193,1399,813,840]" pageId="136" pageNumber="137">– Elytra rounded ................................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389AFF4629F4FC81FAEBFC35" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FC81FAEBFC11" blockId="136.[189,1399,879,941]" box="[189,1399,879,906]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FC81FAEBFC11" box="[189,1399,879,906]" pageId="136" pageNumber="137">
5. Mid-keel wider anteriorly ......................................................................
<taxonomicName id="4C304D34389AFF462D6DFC81FAEBFC11" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1060,1399,879,906]" class="Insecta" family="Ptiliidae" genus="Camptodium" higherTaxonomySource="CoL" kingdom="Plantae" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="nishikawai">
<emphasis id="B944EAA5389AFF462D6DFC81FB56FC12" bold="true" box="[1060,1226,879,906]" italics="true" pageId="136" pageNumber="137">C. nishikawai</emphasis>
<bibRefCitation id="EFA14B46389AFF462D9BFC81FAEBFC11" author="Sawada Y." box="[1234,1399,879,905]" pageId="136" pageNumber="137" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988FC7DFAEBFC35" blockId="136.[189,1399,879,941]" box="[193,1399,914,941]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FC7DFAEBFC35" box="[193,1399,914,941]" pageId="136" pageNumber="137">
– Mid-keel wider posteriorly ....................................................................
<taxonomicName id="4C304D34389AFF462D61FC7DFAEBFC35" ID-CoL="VK88" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[1064,1399,914,941]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="elongatum">
<emphasis id="B944EAA5389AFF462D61FC7DFB57FC35" bold="true" box="[1064,1227,914,941]" italics="true" pageId="136" pageNumber="137">C. elongatum</emphasis>
<bibRefCitation id="EFA14B46389AFF462D9BFC7DFAEBFC35" author="Sawada Y." box="[1234,1399,915,941]" pageId="136" pageNumber="137" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FC3AFAEBFB8B" blockId="136.[189,1399,980,1043]" pageId="136" pageNumber="137">6. Mesoventral lateral margins partly or completely effaced .............................................................. 15 – Mesoventral lateral margins distinct to mesocoxal borders............................................................... 7</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FBD5FAEBFBE0" blockId="136.[189,1399,1082,1144]" pageId="136" pageNumber="137">
</keyStep>
<keyStep id="30C42E12389AFF4629F4FC3AFAEBFB8B" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FC3AFAEAFC77" blockId="136.[189,1399,980,1043]" box="[189,1398,980,1007]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FC3AFAEAFC77" box="[189,1398,980,1007]" pageId="136" pageNumber="137">6. Mesoventral lateral margins partly or completely effaced .............................................................. 15</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988FC17FAEBFB8B" blockId="136.[189,1399,980,1043]" box="[193,1399,1016,1043]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FC17FAEBFB8B" box="[193,1399,1016,1043]" pageId="136" pageNumber="137">– Mesoventral lateral margins distinct to mesocoxal borders............................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389AFF4629F4FBD5FAEBFBE0" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FBD5FAEBFBCD" blockId="136.[189,1399,1082,1144]" box="[189,1399,1082,1109]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FBD5FAEBFBCD" box="[189,1399,1082,1109]" pageId="136" pageNumber="137">
7. Mesoventral collar without a median extension ................................................
<taxonomicName id="4C304D34389AFF462D3EFBD4FA89FBCD" ID-CoL="8XXR7" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1143,1301,1082,1109]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="obsoletum" status="sp. nov.">
<emphasis id="B944EAA5389AFF462D3EFBD4FA89FBCD" bold="true" box="[1143,1301,1082,1109]" italics="true" pageId="136" pageNumber="137">C. obsoletum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55FBD5FAEBFBCD" box="[1308,1399,1083,1109]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
– Mesoventral collar with a median extension ..................................................................................... 8
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FB4EFAEAFB45" blockId="136.[189,1399,1183,1246]" pageId="136" pageNumber="137">8. Mesoventrite lateral margins with a distinct concave bulge at the posterior angle ........................... 9 – Mesoventral lateral margins without a concave bulge..................................................................... 14</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FAEBFAE9FAFE" blockId="136.[189,1398,1285,1383]" pageId="136" pageNumber="137">9. Mid-keel not fusing with the posterior border of the mesoventral collar the anterior angles not interrupting the line of the posterior margin................................................................................... 10 – Mid keel fusing with the posterior border of the mesoventral collar ...............................................11</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FA60FAEBFA54" blockId="136.[189,1399,1422,1484]" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF462988FBB0FAEBFBE0" blockId="136.[189,1399,1082,1144]" box="[193,1399,1118,1144]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FBB0FAEBFBE0" box="[193,1399,1118,1144]" pageId="136" pageNumber="137">– Mesoventral collar with a median extension ..................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389AFF4629F4FB4EFAEAFB45" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FB4EFAEBFB22" blockId="136.[189,1399,1183,1246]" box="[189,1399,1183,1210]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FB4EFAEBFB22" box="[189,1399,1183,1210]" pageId="136" pageNumber="137">8. Mesoventrite lateral margins with a distinct concave bulge at the posterior angle ........................... 9</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988FB2AFAEAFB45" blockId="136.[189,1399,1183,1246]" box="[193,1398,1219,1246]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FB2AFAEAFB45" box="[193,1398,1219,1246]" pageId="136" pageNumber="137">– Mesoventral lateral margins without a concave bulge..................................................................... 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389AFF4629F4FAEBFAE9FAFE" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FAEBFAEAFADB" blockId="136.[189,1398,1285,1383]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FAEBFAEAFADB" pageId="136" pageNumber="137">9. Mid-keel not fusing with the posterior border of the mesoventral collar the anterior angles not interrupting the line of the posterior margin................................................................................... 10</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988FAA3FAE9FAFE" blockId="136.[189,1398,1285,1383]" box="[193,1397,1356,1383]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FAA3FAE9FAFE" box="[193,1397,1356,1383]" pageId="136" pageNumber="137">– Mid keel fusing with the posterior border of the mesoventral collar ...............................................11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389AFF4629F4FA60FAEBFA54" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FA60FAEBFA31" blockId="136.[189,1399,1422,1484]" box="[189,1399,1422,1449]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FA60FAEBFA31" box="[189,1399,1422,1449]" pageId="136" pageNumber="137">
10. Raised posterior margin before keel broad and strongly sculpted.....................
<taxonomicName id="4C304D34389AFF462D3EFA60FA89FA31" ID-CoL="8XXQ9" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1143,1301,1422,1449]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="ibicarense" status="sp. nov.">
<emphasis id="B944EAA5389AFF462D3EFA60FA89FA31" bold="true" box="[1143,1301,1422,1449]" italics="true" pageId="136" pageNumber="137">C. ibicarense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55FA61FAEBFA31" box="[1308,1399,1423,1449]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988FA5CFAEBFA54" blockId="136.[189,1399,1422,1484]" box="[193,1399,1458,1484]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988FA5CFAEBFA54" box="[193,1399,1458,1484]" pageId="136" pageNumber="137">
– Raised posterior margin before keel narrower not sculpted .............................
<taxonomicName id="4C304D34389AFF462D39FA5CFA89FA54" ID-CoL="8XXRC" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1136,1301,1458,1484]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="peruviense" status="sp. nov.">
<emphasis id="B944EAA5389AFF462D39FA5CFA89FA54" bold="true" box="[1136,1301,1458,1484]" italics="true" pageId="136" pageNumber="137">C. peruviense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55FA5CFAEBFA54" box="[1308,1399,1458,1484]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4629F4FA1AFAEAF9A9" blockId="136.[189,1399,1523,1586]" pageId="136" pageNumber="137">
</keyStep>
<keyStep id="30C42E12389AFF4629F4FA1AFAEAF9A9" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4FA1AFAEBF996" blockId="136.[189,1399,1523,1586]" box="[189,1399,1523,1550]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4FA1AFAEBF996" box="[189,1399,1523,1550]" pageId="136" pageNumber="137">
11. Larger species ±
<quantity id="4CC89B52389AFF4628FAFA1AFD87F996" box="[435,539,1524,1550]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="136" pageNumber="137" unit="mm" value="0.9">0.90 mm</quantity>
.........................................................................................
@ -113,19 +190,33 @@ group 4B
<emphasis id="B944EAA5389AFF462DE2FA1AFA89F996" bold="true" box="[1195,1301,1524,1550]" italics="true" pageId="136" pageNumber="137">C. orami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55FA1AFAEBF996" box="[1308,1399,1524,1550]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988F9F6FAEAF9A9" blockId="136.[189,1399,1523,1586]" box="[193,1398,1559,1586]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988F9F6FAEAF9A9" box="[193,1398,1559,1586]" pageId="136" pageNumber="137">
– Smaller species ±
<quantity id="4CC89B52389AFF462889F9F9FDF7F9AA" box="[448,619,1559,1586]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="136" pageNumber="137" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">0.65–0.70 mm</quantity>
..................................................................................................... 12
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4629F4F9B7FAEAF922" blockId="136.[189,1399,1625,1723]" pageId="136" pageNumber="137">
</keyStep>
<keyStep id="30C42E12389AFF4629F4F9B7FAEAF922" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4F9B7FAEBF90F" blockId="136.[189,1399,1625,1723]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4F9B7FAEBF90F" pageId="136" pageNumber="137">
12 Mesoventral lateral margin hind angle sharp the concave extrusion very small................................. .................................................................................................................................
<taxonomicName id="4C304D34389AFF462DEFF993FA89F90F" ID-CoL="8XXPV" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1190,1301,1660,1687]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="dybasi" status="sp. nov.">
<emphasis id="B944EAA5389AFF462DEFF993FA89F90F" bold="true" box="[1190,1301,1660,1687]" italics="true" pageId="136" pageNumber="137">C. dybasi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55F993FAEBF90F" box="[1308,1399,1661,1687]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
– Mesoventral lateral margin hind angle more rounded..................................................................... 13
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4629F4F90CFAEBF8B8" blockId="136.[189,1399,1762,1824]" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF462988F94FFAEAF922" blockId="136.[189,1399,1625,1723]" box="[193,1398,1696,1723]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988F94FFAEAF922" box="[193,1398,1696,1723]" pageId="136" pageNumber="137">– Mesoventral lateral margin hind angle more rounded..................................................................... 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389AFF4629F4F90CFAEBF8B8" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4F90CFAEAF964" blockId="136.[189,1399,1762,1824]" box="[189,1398,1762,1789]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4F90CFAEAF964" box="[189,1398,1762,1789]" pageId="136" pageNumber="137">
13. Mid-keel broad, short, mesoventral collar extension short ....................
<taxonomicName id="4C304D34389AFF462D62F90CFAEAF964" ID-CoL="367FT" authority="(Matthews, 1889)" baseAuthorityName="Matthews" baseAuthorityYear="1889" box="[1067,1398,1762,1788]" class="Insecta" family="Ptiliidae" genus="Dimorphella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="B944EAA5389AFF462D62F90CFB02F964" bold="true" box="[1067,1182,1762,1788]" italics="true" pageId="136" pageNumber="137">C. reitteri</emphasis>
@ -133,79 +224,143 @@ group 4B
<bibRefCitation id="EFA14B46389AFF462DE7F90CFAF2F964" author="Matthews A." box="[1198,1390,1762,1788]" pageId="136" pageNumber="137" pagination="188 - 195" refId="ref59760" refString="Matthews A. 1889. New genera and species of Trichopterygidae. Annals and Magazine of Natural History (6) III: 188 - 195. Available from https: // biodiversitylibrary. org / page / 15621981 [accessed 4 Feb. 2020]." type="journal article" year="1889">Matthews, 1889</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988F8E8FAEBF8B8" blockId="136.[189,1399,1762,1824]" box="[193,1399,1797,1824]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988F8E8FAEBF8B8" box="[193,1399,1797,1824]" pageId="136" pageNumber="137">
– Mid keel narrower, longer, mesoventral collar extension longer .......................
<taxonomicName id="4C304D34389AFF462DCBF8E8FA88F8B8" ID-CoL="8XXPJ" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1154,1300,1797,1824]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="bomjesus" status="sp. nov.">
<emphasis id="B944EAA5389AFF462DCBF8E8FA88F8B8" bold="true" box="[1154,1300,1797,1824]" italics="true" pageId="136" pageNumber="137">C. bomjesus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55F8E8FAEBF8B8" box="[1308,1399,1798,1824]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4629F4F8A6FAEBF81E" blockId="136.[189,1399,1863,1926]" pageId="136" pageNumber="137">
</keyStep>
<keyStep id="30C42E12389AFF4629F4F8A6FAEBF81E" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4F8A6FAEBF8FA" blockId="136.[189,1399,1863,1926]" box="[189,1399,1863,1890]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4F8A6FAEBF8FA" box="[189,1399,1863,1890]" pageId="136" pageNumber="137">
14. Mid-keel very much reduced almost absent, keel strongly sculpted ...................
<taxonomicName id="4C304D34389AFF462DC0F8A6FA89F8FA" ID-CoL="8XXQM" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1161,1301,1863,1890]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="losbanos" status="sp. nov.">
<emphasis id="B944EAA5389AFF462DC0F8A6FA89F8FA" bold="true" box="[1161,1301,1863,1890]" italics="true" pageId="136" pageNumber="137">C. losbanos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55F8A6FAEBF8FA" box="[1308,1399,1864,1890]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF462988F882FAEBF81E" blockId="136.[189,1399,1863,1926]" box="[193,1399,1899,1926]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988F882FAEBF81E" box="[193,1399,1899,1926]" pageId="136" pageNumber="137">
– Keel narrow before joining keel, less strongly sculpted................................
<taxonomicName id="4C304D34389AFF462D13F885FAEBF81E" ID-CoL="VK8R" authority="Darby, 2015" authorityName="Darby" authorityYear="2015" box="[1114,1399,1899,1926]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="sawadai">
<emphasis id="B944EAA5389AFF462D13F885FB43F81E" bold="true" box="[1114,1247,1899,1926]" italics="true" pageId="136" pageNumber="137">C. sawadai</emphasis>
<bibRefCitation id="EFA14B46389AFF462DAFF885FAEBF81E" author="Darby M." box="[1254,1399,1899,1926]" pageId="136" pageNumber="137" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby, 2015</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389AFF4729F4F843FAEAFED1" blockId="136.[189,1399,1965,2027]" lastBlockId="137.[189,1398,267,330]" lastPageId="137" lastPageNumber="138" pageId="136" pageNumber="137">
</keyStep>
<keyStep id="30C42E12389AFF4629F4F843FAEAF873" pageId="136" pageNumber="137">
<paragraph id="8B8F36B7389AFF4629F4F843FAEBF850" blockId="136.[189,1399,1965,2027]" box="[189,1399,1965,1992]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF4629F4F843FAEBF850" box="[189,1399,1965,1992]" pageId="136" pageNumber="137">
15. Keel very broad, flat, with rounded tip ..............................................................
<taxonomicName id="4C304D34389AFF462D33F843FA89F850" ID-CoL="8XXRQ" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1146,1301,1965,1992]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="136" pageNumber="137" phylum="Arthropoda" rank="species" species="similatum" status="sp. nov.">
<emphasis id="B944EAA5389AFF462D33F843FA89F850" bold="true" box="[1146,1301,1965,1992]" italics="true" pageId="136" pageNumber="137">C. similatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389AFF462C55F840FAEBF850" box="[1308,1399,1966,1992]" pageId="136" pageNumber="137" rank="species">sp. nov.</taxonomicNameLabel>
– Keel narrower .................................................................................................................................. 16 16. Mid-keel sides strongly widened anteriorly .................................................................................... 17 – Mid-keel sides not so widened anteriorly, sides concave ................................................................ 19
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389BFF4729F4FE9FFAEBFE4C" blockId="137.[189,1399,369,468]" pageId="137" pageNumber="138">
17. Posterior margins of humeri very long ............................................................................................ 18 – Posterior margins of humeri shorter, mid-keel with concave sides .................................................... ...............................................................................................................
<paragraph id="8B8F36B7389AFF462988F83FFAEAF873" blockId="136.[189,1399,1965,2027]" box="[193,1398,2001,2027]" pageId="136" pageNumber="137">
<keyLead id="30C19582389AFF462988F83FFAEAF873" box="[193,1398,2001,2027]" pageId="136" pageNumber="137">– Keel narrower .................................................................................................................................. 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389BFF4729F4FEE5FAEAFED1" pageId="137" pageNumber="138">
<paragraph id="8B8F36B7389BFF4729F4FEE5FAEAFEBD" blockId="137.[189,1398,267,330]" box="[189,1398,267,294]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF4729F4FEE5FAEAFEBD" box="[189,1398,267,294]" pageId="137" pageNumber="138">16. Mid-keel sides strongly widened anteriorly .................................................................................... 17</keyLead>
</paragraph>
<paragraph id="8B8F36B7389BFF472988FEDEFAEAFED1" blockId="137.[189,1398,267,330]" box="[193,1398,303,330]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF472988FEDEFAEAFED1" box="[193,1398,303,330]" pageId="137" pageNumber="138">– Mid-keel sides not so widened anteriorly, sides concave ................................................................ 19</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389BFF4729F4FE9FFAEBFE4C" pageId="137" pageNumber="138">
<paragraph id="8B8F36B7389BFF4729F4FE9FFAEAFE13" blockId="137.[189,1399,369,468]" box="[189,1398,369,396]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF4729F4FE9FFAEAFE13" box="[189,1398,369,396]" pageId="137" pageNumber="138">17. Posterior margins of humeri very long ............................................................................................ 18</keyLead>
</paragraph>
<paragraph id="8B8F36B7389BFF472988FE78FAEBFE4C" blockId="137.[189,1399,369,468]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF472988FE78FAEBFE4C" pageId="137" pageNumber="138">
– Posterior margins of humeri shorter, mid-keel with concave sides .................................................... ...............................................................................................................
<taxonomicName id="4C304D34389BFF472D68FE57FAEBFE4C" ID-CoL="VK86" authority="Motschulsky, 1855" authorityName=", Motschulsky" authorityYear="1855" box="[1057,1399,440,468]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="basale">
<emphasis id="B944EAA5389BFF472D68FE57FB0DFE4B" bold="true" box="[1057,1169,440,467]" italics="true" pageId="137" pageNumber="138">C. basale</emphasis>
<bibRefCitation id="EFA14B46389BFF472DDEFE57FAEBFE4C" author="Motschulsky V. von" box="[1175,1399,441,468]" pageId="137" pageNumber="138" pagination="8 - 25" refId="ref59847" refString="Motschulsky V. von. 1855. Voyages. Lettre de M. de Motschulsky a M. Menetries. No 2, au bord du bateau a vapeur United-States, 20 Mars 1854. Etudes entomologiques 4: 8 - 25." type="journal article" year="1855">Motschulsky, 1855</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389BFF4729F4FE15FAEBFDA1" blockId="137.[189,1399,506,569]" pageId="137" pageNumber="138">
</keyStep>
<keyStep id="30C42E12389BFF4729F4FE15FAEBFDA1" pageId="137" pageNumber="138">
<paragraph id="8B8F36B7389BFF4729F4FE15FAEBFD8E" blockId="137.[189,1399,506,569]" box="[189,1399,506,534]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF4729F4FE15FAEBFD8E" box="[189,1399,506,534]" pageId="137" pageNumber="138">
18. Longer species, mid-keel wider...........................................................................
<taxonomicName id="4C304D34389BFF472DCCFE15FAEBFD8E" ID-CoL="VK8F" authority="Darby, 2015" authorityName="Darby" authorityYear="2015" box="[1157,1399,506,534]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="lisae">
<emphasis id="B944EAA5389BFF472DCCFE15FB42FD8D" bold="true" box="[1157,1246,506,533]" italics="true" pageId="137" pageNumber="138">C. lisae</emphasis>
<bibRefCitation id="EFA14B46389BFF472DAFFE15FAEBFD8E" author="Darby M." box="[1254,1399,507,534]" pageId="137" pageNumber="138" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby, 2015</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389BFF472988FDF1FAEBFDA1" blockId="137.[189,1399,506,569]" box="[193,1399,542,569]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF472988FDF1FAEBFDA1" box="[193,1399,542,569]" pageId="137" pageNumber="138">
– Shorter species, mid keel shorter, keel much narrower ...................................
<taxonomicName id="4C304D34389BFF472D22FDF1FAEBFDA1" ID-CoL="5YWSC" authority="Darby, 2015" authorityName="Darby" authorityYear="2015" box="[1131,1399,542,569]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="tigrum">
<emphasis id="B944EAA5389BFF472D22FDF1FB43FDA1" bold="true" box="[1131,1247,543,569]" italics="true" pageId="137" pageNumber="138">C. tigrum</emphasis>
<bibRefCitation id="EFA14B46389BFF472DAFFDF1FAEBFDA1" author="Darby M." box="[1254,1399,542,569]" pageId="137" pageNumber="138" pagination="179 - 190" refId="ref59379" refString="Darby M. 2015. Studies of Bolivian Ptiliidae (Coleoptera) 1: The genus Cissidium Motschulsky with six new species. Zootaxa 3937 (1): 179 - 190. https: // doi. org / 10.11646 / zootaxa. 3937.1.9" type="journal article" year="2015">Darby, 2015</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389BFF4729F4FD8FFAEAFD07" blockId="137.[189,1399,608,671]" pageId="137" pageNumber="138">
</keyStep>
<keyStep id="30C42E12389BFF4729F4FD8FFAEAFD07" pageId="137" pageNumber="138">
<paragraph id="8B8F36B7389BFF4729F4FD8FFAEBFDE3" blockId="137.[189,1399,608,671]" box="[189,1399,608,635]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF4729F4FD8FFAEBFDE3" box="[189,1399,608,635]" pageId="137" pageNumber="138">
19. Mid-keel hind angles rounded ...........................................................................
<taxonomicName id="4C304D34389BFF472D31FD8FFA89FDE3" ID-CoL="8XXR5" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1144,1301,608,635]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="obscenum" status="sp. nov.">
<emphasis id="B944EAA5389BFF472D31FD8FFA89FDE3" bold="true" box="[1144,1301,608,635]" italics="true" pageId="137" pageNumber="138">C. obscenum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389BFF472C55FD8FFAEBFDE3" box="[1308,1399,609,635]" pageId="137" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
– Mid-keel hind angles not rounded, extending towards mesocoxal posterior border....................... 20
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389BFF4729F4FD29FAEAFC9C" blockId="137.[189,1399,710,773]" pageId="137" pageNumber="138">
<paragraph id="8B8F36B7389BFF472988FD6BFAEAFD07" blockId="137.[189,1399,608,671]" box="[193,1398,644,671]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF472988FD6BFAEAFD07" box="[193,1398,644,671]" pageId="137" pageNumber="138">– Mid-keel hind angles not rounded, extending towards mesocoxal posterior border....................... 20</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389BFF4729F4FD29FAEAFC9C" pageId="137" pageNumber="138">
<paragraph id="8B8F36B7389BFF4729F4FD29FAEBFD79" blockId="137.[189,1399,710,773]" box="[189,1399,710,737]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF4729F4FD29FAEBFD79" box="[189,1399,710,737]" pageId="137" pageNumber="138">
20. Mesoventral collar without a medial extension ..................................................
<taxonomicName id="4C304D34389BFF472DC8FD29FA89FD79" ID-CoL="8XXPT" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1153,1301,710,737]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="dividuum" status="sp. nov.">
<emphasis id="B944EAA5389BFF472DC8FD29FA89FD79" bold="true" box="[1153,1301,710,737]" italics="true" pageId="137" pageNumber="138">C. dividuum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757DE389BFF472C55FD29FAEBFD79" box="[1308,1399,711,737]" pageId="137" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
– Mesoventral collar with a medial extension .................................................................................... 21
</keyLead>
</paragraph>
<paragraph id="8B8F36B7389BFF4729F4FCC3FAEBFC17" blockId="137.[189,1399,812,911]" pageId="137" pageNumber="138">
<paragraph id="8B8F36B7389BFF472988FD05FAEAFC9C" blockId="137.[189,1399,710,773]" box="[193,1398,746,773]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF472988FD05FAEAFC9C" box="[193,1398,746,773]" pageId="137" pageNumber="138">– Mesoventral collar with a medial extension .................................................................................... 21</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E12389BFF4729F4FCC3FAEBFC17" pageId="137" pageNumber="138">
<paragraph id="8B8F36B7389BFF4729F4FCC3FAEBFCF3" blockId="137.[189,1399,812,911]" pageId="137" pageNumber="138">
<keyLead id="30C19582389BFF4729F4FCC3FAEBFCF3" pageId="137" pageNumber="138">
21. Mesoventral median extension long, with a splayed end, keel very long and narrow ........................................................................................................
<taxonomicName id="4C304D34389BFF472D0DFCBEFA88FCF3" ID-CoL="8XXRS" authority="Darby, 2020" authorityName="Darby" authorityYear="2020" box="[1092,1300,848,875]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="137" pageNumber="138" phylum="Arthropoda" rank="species" species="subangulatum" status="sp. nov.">
<emphasis id="B944EAA5389BFF472D0DFCBEFA88FCF3" bold="true" box="[1092,1300,848,875]" italics="true" pageId="137" pageNumber="138">C. subangulatum</emphasis>
</taxonomicName>
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– Shorter species, mid-keel shorter, keel shorter and wider at base ...................
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, pl. XIV, fig. 35.
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–
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<treatmentCitation id="4C19731AD82CFFCFFAAFF96EF721F8EF" author="Verdcourt B." box="[1291,1394,1774,1800]" page="49" pageId="8" pageNumber="9" year="2006">
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.? “
.
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? “
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,
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, dry forest on dark, sandy soil on coral rag on Tondooni peninsula within reserve, leg.
, Zanzibar, dry forest on dark, sandy soil on coral rag on Tondooni peninsula within reserve, leg.
<collectorName id="604D30DDD837FFD4FAA2F80AF77DF843" box="[1286,1326,1930,1956]" pageId="19" pageNumber="20">BR</collectorName>
,
<collectorName id="604D30DDD837FFD4FA9EF80AF340F82F" pageId="19" pageNumber="20">B. H. Warren</collectorName>
@ -164,14 +159,11 @@ material examined
<date id="B90673CBD837FFD4FEDEF82EF056F820" box="[378,517,1965,1992]" pageId="19" pageNumber="20" value="2009-02-07">
<collectingDate id="A9428A23D837FFD4FEDEF82EF056F820" box="[378,517,1965,1992]" pageId="19" pageNumber="20" value="2009-02-07">7 Feb. 2009</collectingDate>
</date>
; fig.
<quantity id="0A40F8EED837FFD4FD9FF82DF026F820" box="[571,629,1965,1991]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="19" pageNumber="20" unit="in" value="60.0">60 in</quantity>
; fig. 60 in
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<emphasis id="FFCC8919D837FFD4FD44F82EF14BF820" box="[736,792,1965,1991]" italics="true" pageId="19" pageNumber="20">et al.</emphasis>
(
<date id="B90673CBD837FFD4FC82F82DF135F82F" box="[806,870,1965,1992]" pageId="19" pageNumber="20" value="2010">2010</date>
b)
(2010b)
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.

82
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, 27-28, pl. 3, fig. 2.
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@ -98,8 +98,10 @@ sp. nov.
Verdcourt
<date id="B90673CBD837FFD4FD83FE33F035FE29" box="[551,614,435,462]" pageId="19" pageNumber="20" value="1958">1958</date>
: 94, 100, fig. 10.
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–
@ -186,8 +188,8 @@ Muratov
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<mods:title id="9CC8E4F15A3B6772C297ABD2BA64E7E8">Revision of Dadagulella gen. nov., the “ Gulella radius group ” (Gastropoda: Streptaxidae) of the eastern Afrotropics, including six new species and three new subspecies</mods:title>
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<emphasis bold="true" box="[517,933,267,294]" italics="true" pageId="28" pageNumber="29">Dadagulella minuscula mahorana</emphasis>
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<emphasis id="FFCC8919D838FFDBFDA1FE8BF1F6FEC1" bold="true" box="[517,933,267,294]" italics="true" pageId="28" pageNumber="29">Dadagulella minuscula mahorana</emphasis>
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<figureCitation box="[655,745,303,329]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="28" pageNumber="29">Figs 40</figureCitation>
<subSubSection id="85A20680D838FFDBFD2BFEAFF1F7FEAE" box="[655,932,302,329]" pageId="28" pageNumber="29" type="description">
<paragraph id="CD07550BD838FFDBFD2BFEAFF1F7FEAE" blockId="28.[517,1071,267,329]" box="[655,932,302,329]" pageId="28" pageNumber="29">
<figureCitation id="5583498ED838FFDBFD2BFEAFF0BAFEAE" box="[655,745,303,329]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="28" pageNumber="29">Figs 40</figureCitation>
,
<figureCitation box="[757,789,302,328]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="28" pageNumber="29">64</figureCitation>
<figureCitation id="5583498ED838FFDBFD51FEAEF146FEAF" box="[757,789,302,328]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="28" pageNumber="29">64</figureCitation>
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<figureCitation box="[801,834,303,329]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="28" pageNumber="29">84</figureCitation>
<figureCitation id="5583498ED838FFDBFC85FEAFF111FEAE" box="[801,834,303,329]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="28" pageNumber="29">84</figureCitation>
;
<tableCitation box="[845,932,303,329]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="28" pageNumber="29" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
<tableCitation id="803A60B0D838FFDBFCE9FEAFF1F7FEAE" box="[845,932,303,329]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetBox="[202,1925,710,1733]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="28" pageNumber="29" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
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?
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<emphasis box="[209,284,369,394]" italics="true" pageId="28" pageNumber="29">Ennea</emphasis>
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<emphasis id="FFCC8919D838FFDBFF75FEF1F34FFE6D" box="[209,284,369,394]" italics="true" pageId="28" pageNumber="29">Ennea</emphasis>
(
<emphasis box="[301,438,368,394]" italics="true" pageId="28" pageNumber="29">Uniplicaria</emphasis>
<emphasis id="FFCC8919D838FFDBFE89FEF0F3E5FE6D" box="[301,438,368,394]" italics="true" pageId="28" pageNumber="29">Uniplicaria</emphasis>
)
<emphasis box="[455,585,368,394]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
<emphasis id="FFCC8919D838FFDBFE63FEF0F01AFE6D" box="[455,585,368,394]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
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“grössere Form auf
<collectingCountry box="[827,928,368,394]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
<collectingCountry id="B5AF159BD838FFDBFC9FFEF0F1F3FE6D" box="[827,928,368,394]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
” –
<treatmentCitation author="Kobelt W." box="[967,1175,368,395]" page="166" pageId="28" pageNumber="29" year="1905">
<bibRefCitation author="Kobelt W." box="[967,1175,368,395]" pageId="28" pageNumber="29" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt 1905: 166</bibRefCitation>
<treatmentCitation id="4C19731AD838FFDBFC63FEF0F6C4FE6D" author="Kobelt W." box="[967,1175,368,395]" page="166" pageId="28" pageNumber="29" year="1905">
<bibRefCitation id="A92928FAD838FFDBFC63FEF0F6C4FE6D" author="Kobelt W." box="[967,1175,368,395]" pageId="28" pageNumber="29" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt 1905: 166</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="etymology">
<paragraph blockId="28.[189,324,434,460]" box="[189,324,434,460]" pageId="28" pageNumber="29">
<heading bold="true" box="[189,324,434,460]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis bold="true" box="[189,324,434,460]" pageId="28" pageNumber="29">Etymology</emphasis>
<subSubSection id="85A20680D838FFDBFF19FE32F1BFFDF9" pageId="28" pageNumber="29" type="etymology">
<paragraph id="CD07550BD838FFDBFF19FE32F317FE2B" blockId="28.[189,324,434,460]" box="[189,324,434,460]" pageId="28" pageNumber="29">
<heading id="964FE267D838FFDBFF19FE32F317FE2B" bold="true" box="[189,324,434,460]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="FFCC8919D838FFDBFF19FE32F317FE2B" bold="true" box="[189,324,434,460]" pageId="28" pageNumber="29">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[189,1399,480,542]" pageId="28" pageNumber="29">
<paragraph id="CD07550BD838FFDBFF19FE61F1BFFDF9" blockId="28.[189,1399,480,542]" pageId="28" pageNumber="29">
From English noun or adjective ‘Mahoran’, denoting a person or thing from
<collectingCountry box="[1093,1195,481,507]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
<collectingCountry id="B5AF159BD838FFDBFBE1FE61F6F8FE1C" box="[1093,1195,481,507]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
; Latinized by the addition of the feminine ending ‘-
<emphasis box="[586,601,517,542]" italics="true" pageId="28" pageNumber="29">a</emphasis>
<emphasis id="FFCC8919D838FFDBFDEEFD85F00AFDF9" box="[586,601,517,542]" italics="true" pageId="28" pageNumber="29">a</emphasis>
’ and used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="materials_examined">
<paragraph blockId="28.[189,492,581,607]" box="[189,492,581,607]" pageId="28" pageNumber="29">
<heading bold="true" box="[189,492,581,607]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis bold="true" box="[189,492,581,607]" pageId="28" pageNumber="29">
<typeStatus box="[189,250,581,607]" pageId="28" pageNumber="29">Type</typeStatus>
<subSubSection id="85A20680D838FFDBFF19FDC5F0BAFD56" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="CD07550BD838FFDBFF19FDC5F3BFFDB8" blockId="28.[189,492,581,607]" box="[189,492,581,607]" pageId="28" pageNumber="29">
<heading id="964FE267D838FFDBFF19FDC5F3BFFDB8" bold="true" box="[189,492,581,607]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="FFCC8919D838FFDBFF19FDC5F3BFFDB8" bold="true" box="[189,492,581,607]" pageId="28" pageNumber="29">
<typeStatus id="1203EBA9D838FFDBFF19FDC5F2A9FDB8" box="[189,250,581,607]" pageId="28" pageNumber="29">Type</typeStatus>
material examined
</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[189,1398,628,690]" pageId="28" pageNumber="29">
<materialsCitation ID-GBIF-Occurrence="2610647310" box="[189,1272,628,655]" collectionCode="NMW" country="Mayotte" location="MAYOTTE" pageId="28" pageNumber="29" specimenCode="NMW.1955.158.25051" specimenCount="1" specimenCount-adult="1" typeStatus="holotype">
<collectingCountry box="[189,335,628,654]" name="Mayotte" pageId="28" pageNumber="29">MAYOTTE</collectingCountry>
<paragraph id="CD07550BD838FFDBFF19FDF4F0BAFD56" blockId="28.[189,1398,628,690]" pageId="28" pageNumber="29">
<materialsCitation id="7DD05F56D838FFDBFF19FDF4F6ABFD69" ID-GBIF-Occurrence="2610647310" box="[189,1272,628,655]" collectionCode="NMW" country="Mayotte" location="Mayotte I." pageId="28" pageNumber="29" specimenCode="NMW.1955.158.25051" specimenCount="1" specimenCount-adult="1" typeStatus="holotype">
<collectingCountry id="B5AF159BD838FFDBFF19FDF4F31CFD69" box="[189,335,628,654]" name="Mayotte" pageId="28" pageNumber="29">MAYOTTE</collectingCountry>
:
<typeStatus box="[345,448,628,654]" pageId="28" pageNumber="29">holotype</typeStatus>
<specimenCode box="[455,730,628,655]" pageId="28" pageNumber="29">NMW.1955.158.25051</specimenCode>
<typeStatus id="1203EBA9D838FFDBFEFDFDF4F393FD69" box="[345,448,628,654]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
<specimenCode id="9D1EFD70D838FFDBFE63FDF4F089FD68" box="[455,730,628,655]" pageId="28" pageNumber="29">NMW.1955.158.25051</specimenCode>
:
<specimenCount box="[741,800,628,654]" pageId="28" pageNumber="29" type="adult">1 ad.</specimenCount>
, “Mayotte I.”, standing as “
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[1126,1256,628,654]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[1126,1256,628,654]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
<specimenCount id="DBBE9E82D838FFDBFD41FDF4F173FD69" box="[741,800,628,654]" count="1" pageId="28" pageNumber="29" type="adult">1 ad.</specimenCount>
, “
<location id="C86703D0D838FFDBFC92FDF4F1E7FD69" LSID="urn:lsid:plazi:treatment:4511E41DD838FFDAFDA1FE8BF394FBA8:C86703D0D838FFDBFC92FDF4F1E7FD69" box="[822,948,628,654]" country="Mayotte" name="Mayotte I." pageId="28" pageNumber="29">Mayotte I.</location>
”, standing as “
<taxonomicName id="0AB82E88D838FFDBFBC2FDF4F6BBFD69" authorityName="Morelet" authorityYear="1877" box="[1126,1256,628,654]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D838FFDBFBC2FDF4F6BBFD69" box="[1126,1256,628,654]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
</taxonomicName>
”
</materialsCitation>
.
<typeStatus box="[1284,1398,628,654]" pageId="28" pageNumber="29">Paratypes</typeStatus>
<collectionCode box="[189,267,663,690]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="29">NMW</collectionCode>
.1955.158.25065:
<specimenCount box="[475,534,663,689]" pageId="28" pageNumber="29" type="generic">2 ad.</specimenCount>
, data as
<typeStatus box="[634,741,663,689]" pageId="28" pageNumber="29">holotype</typeStatus>
.
<materialsCitation id="7DD05F56D838FFDBFAA0FDF4F0BAFD56" ID-GBIF-Occurrence="2610647310" collectionCode="NMW" country="Mayotte" location="Mayotte I." pageId="28" pageNumber="29" specimenCode="1955.158.25065" specimenCount="2" specimenCount-adult="2" typeStatus="paratype">
<typeStatus id="1203EBA9D838FFDBFAA0FDF4F725FD69" box="[1284,1398,628,654]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
<collectionCode id="ABA9CDCED838FFDBFF19FD17F358FD55" box="[189,267,663,690]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="28" pageNumber="29">NMW</collectionCode>
.
<specimenCode id="9D1EFD70D838FFDBFEAAFD17F383FD55" box="[270,464,663,690]" pageId="28" pageNumber="29">1955.158.25065</specimenCode>
:
<specimenCount id="DBBE9E82D838FFDBFE7FFD17F045FD56" box="[475,534,663,689]" count="2" pageId="28" pageNumber="29" type="adult">2 ad.</specimenCount>
, data as holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="distribution">
<paragraph blockId="28.[189,349,728,755]" box="[189,349,728,755]" pageId="28" pageNumber="29">
<heading bold="true" box="[189,349,728,755]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis bold="true" box="[189,349,728,755]" pageId="28" pageNumber="29">
<typeStatus box="[189,250,729,755]" pageId="28" pageNumber="29">Type</typeStatus>
<subSubSection id="85A20680D838FFDBFF19FD59F12BFCC6" pageId="28" pageNumber="29" type="distribution">
<paragraph id="CD07550BD838FFDBFF19FD59F30EFD15" blockId="28.[189,349,728,755]" box="[189,349,728,755]" pageId="28" pageNumber="29">
<heading id="964FE267D838FFDBFF19FD59F30EFD15" bold="true" box="[189,349,728,755]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="FFCC8919D838FFDBFF19FD59F30EFD15" bold="true" box="[189,349,728,755]" pageId="28" pageNumber="29">
<typeStatus id="1203EBA9D838FFDBFF19FD59F2A9FD14" box="[189,250,729,755]" pageId="28" pageNumber="29">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[189,888,775,802]" box="[189,888,775,802]" pageId="28" pageNumber="29">
<collectingCountry box="[189,287,776,802]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
<paragraph id="CD07550BD838FFDBFF19FC88F12BFCC6" blockId="28.[189,888,775,802]" box="[189,888,775,802]" pageId="28" pageNumber="29">
<collectingCountry id="B5AF159BD838FFDBFF19FC88F34CFCC5" box="[189,287,776,802]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
(approx.
<geoCoordinate box="[400,496,775,802]" degrees="12.85" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="-12.85">12.85°S</geoCoordinate>
<geoCoordinate id="A88C33CCD838FFDBFE34FC87F3A3FCC5" box="[400,496,775,802]" degrees="12.85" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="-12.85">12.85°S</geoCoordinate>
,
<geoCoordinate box="[509,608,775,802]" degrees="45.15" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="45.15">45.15°E</geoCoordinate>
<geoCoordinate id="A88C33CCD838FFDBFE59FC87F033FCC5" box="[509,608,775,802]" degrees="45.15" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="45.15">45.15°E</geoCoordinate>
),
<collectingCountry box="[629,737,775,801]" name="Comoros" pageId="28" pageNumber="29">Comoros</collectingCountry>
<collectingCountry id="B5AF159BD838FFDBFDD1FC87F0B2FCC6" box="[629,737,775,801]" name="Comoros" pageId="28" pageNumber="29">Comoros</collectingCountry>
archipelago.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="materials_examined">
<paragraph blockId="28.[189,505,841,867]" box="[189,505,841,867]" pageId="28" pageNumber="29">
<heading bold="true" box="[189,505,841,867]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis bold="true" box="[189,505,841,867]" pageId="28" pageNumber="29">Other material examined</emphasis>
<subSubSection id="85A20680D838FFDBFF19FCC9F0EEFC3F" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="CD07550BD838FFDBFF19FCC9F3AAFC84" blockId="28.[189,505,841,867]" box="[189,505,841,867]" pageId="28" pageNumber="29">
<heading id="964FE267D838FFDBFF19FCC9F3AAFC84" bold="true" box="[189,505,841,867]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="FFCC8919D838FFDBFF19FCC9F3AAFC84" bold="true" box="[189,505,841,867]" pageId="28" pageNumber="29">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[189,1399,887,985]" pageId="28" pageNumber="29">
<collectingCountry box="[189,335,888,914]" name="Mayotte" pageId="28" pageNumber="29">MAYOTTE</collectingCountry>
<paragraph id="CD07550BD838FFDBFF19FCF8F0EEFC3F" blockId="28.[189,1399,887,985]" pageId="28" pageNumber="29">
<collectingCountry id="B5AF159BD838FFDBFF19FCF8F31CFC75" box="[189,335,888,914]" name="Mayotte" pageId="28" pageNumber="29">MAYOTTE</collectingCountry>
:
<collectionCode box="[355,472,887,914]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="28" pageNumber="29" type="Museum">NHMUK</collectionCode>
<collectionCode id="ABA9CDCED838FFDBFEC7FCF7F38BFC75" box="[355,472,887,914]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="28" pageNumber="29" type="Museum">NHMUK</collectionCode>
.1882.5.27.6-7:
<specimenCount box="[662,731,887,914]" pageId="28" pageNumber="29" type="generic">2 ad.</specimenCount>
<specimenCount id="DBBE9E82D838FFDBFD32FCF8F088FC76" box="[662,731,887,914]" pageId="28" pageNumber="29" type="generic">2 ad.</specimenCount>
, “Mayotte” (approx.
<geoCoordinate box="[1009,1106,887,914]" degrees="12.85" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="-12.85">12.85°S</geoCoordinate>
<geoCoordinate id="A88C33CCD838FFDBFC55FCF7F601FC75" box="[1009,1106,887,914]" degrees="12.85" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="555" value="-12.85">12.85°S</geoCoordinate>
,
<geoCoordinate box="[1128,1227,887,914]" degrees="45.15" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="45.15">45.15°E</geoCoordinate>
<geoCoordinate id="A88C33CCD838FFDBFBCCFCF7F698FC75" box="[1128,1227,887,914]" degrees="45.15" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="555" value="45.15">45.15°E</geoCoordinate>
), standing as “
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[202,332,923,949]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[202,332,923,949]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
<taxonomicName id="0AB82E88D838FFDBFF6EFC1BF31FFC52" authorityName="Morelet" authorityYear="1877" box="[202,332,923,949]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D838FFDBFF6EFC1BF31FFC52" box="[202,332,923,949]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
</taxonomicName>
”.
<collectionCode box="[361,477,923,949]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="28" pageNumber="29" type="Museum">NHMUK</collectionCode>
<collectionCode id="ABA9CDCED838FFDBFECDFC1BF38EFC52" box="[361,477,923,949]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="28" pageNumber="29" type="Museum">NHMUK</collectionCode>
.20110171:
<specimenCount box="[611,671,923,949]" pageId="28" pageNumber="29" type="generic">2 ad.</specimenCount>
<specimenCount id="DBBE9E82D838FFDBFDC7FC1BF0CCFC52" box="[611,671,923,949]" pageId="28" pageNumber="29" type="generic">2 ad.</specimenCount>
, “Mayotte”, standing as “
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[976,1106,923,949]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[976,1106,923,949]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
<taxonomicName id="0AB82E88D838FFDBFC74FC1BF601FC52" authorityName="Morelet" authorityYear="1877" box="[976,1106,923,949]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D838FFDBFC74FC1BF601FC52" box="[976,1106,923,949]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
</taxonomicName>
”.
<collectionCode box="[1134,1227,923,949]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29" type="Museum">RMNH</collectionCode>
<collectionCode id="ABA9CDCED838FFDBFBCAFC1BF698FC52" box="[1134,1227,923,949]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="29" type="Museum">RMNH</collectionCode>
.MOL.273928:
<specimenCount box="[189,248,958,984]" pageId="28" pageNumber="29" type="generic">1 ad.</specimenCount>
<specimenCount id="DBBE9E82D838FFDBFF19FC3EF2ABFC3F" box="[189,248,958,984]" pageId="28" pageNumber="29" type="generic">1 ad.</specimenCount>
, “Mayotte”, standing as “
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[551,681,958,984]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[551,681,958,984]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
<taxonomicName id="0AB82E88D838FFDBFD83FC3EF0FAFC3F" authorityName="Morelet" authorityYear="1877" box="[551,681,958,984]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D838FFDBFD83FC3EF0FAFC3F" box="[551,681,958,984]" italics="true" pageId="28" pageNumber="29">fischeriana</emphasis>
</taxonomicName>
”.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="description">
<paragraph blockId="28.[189,334,1023,1049]" box="[189,334,1023,1049]" pageId="28" pageNumber="29">
<heading bold="true" box="[189,334,1023,1049]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis bold="true" box="[189,334,1023,1049]" pageId="28" pageNumber="29">Description</emphasis>
<subSubSection id="85A20680D838FFDBFF19FC7FF361FA61" pageId="28" pageNumber="29" type="description">
<paragraph id="CD07550BD838FFDBFF19FC7FF31DFBFE" blockId="28.[189,334,1023,1049]" box="[189,334,1023,1049]" pageId="28" pageNumber="29">
<heading id="964FE267D838FFDBFF19FC7FF31DFBFE" bold="true" box="[189,334,1023,1049]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="FFCC8919D838FFDBFF19FC7FF31DFBFE" bold="true" box="[189,334,1023,1049]" pageId="28" pageNumber="29">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[189,1399,1070,1414]" pageId="28" pageNumber="29">
<paragraph id="CD07550BD838FFDBFF19FBAEF361FA61" blockId="28.[189,1399,1070,1414]" pageId="28" pageNumber="29">
SHELL (
<figureCitation box="[277,369,1070,1096]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="28" pageNumber="29">Figs 40</figureCitation>
<figureCitation id="5583498ED838FFDBFEB1FBAEF322FBAF" box="[277,369,1070,1096]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="28" pageNumber="29">Figs 40</figureCitation>
,
<figureCitation box="[383,416,1070,1097]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="28" pageNumber="29">64</figureCitation>
<figureCitation id="5583498ED838FFDBFEDBFBAEF3F3FBAE" box="[383,416,1070,1097]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="28" pageNumber="29">64</figureCitation>
). Variable in size, shape and dentition, large (
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4.30
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<emphasis id="FFCC8919D838FFDBFC5FFBAFF656FBAF" box="[1019,1029,1071,1096]" italics="true" pageId="28" pageNumber="29">-</emphasis>
4.60 mm
</quantity>
high x
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<quantity id="0A40F8EED838FFDBFB6BFBAEF724FBAE" box="[1231,1399,1070,1097]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="28" pageNumber="29" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">
1.90
<emphasis box="[1282,1292,1071,1096]" italics="true" pageId="28" pageNumber="29">-</emphasis>
<emphasis id="FFCC8919D838FFDBFAA6FBAFF75FFBAF" box="[1282,1292,1071,1096]" italics="true" pageId="28" pageNumber="29">-</emphasis>
2.20 mm
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wide), of 6.5
<emphasis box="[343,353,1106,1131]" italics="true" pageId="28" pageNumber="29">-</emphasis>
<emphasis id="FFCC8919D838FFDBFEF3FBD2F332FB8C" box="[343,353,1106,1131]" italics="true" pageId="28" pageNumber="29">-</emphasis>
7.5 whorls. Ovate-acuminate, spire narrowly acuminate to coeloconoid (spire angle 50
<emphasis box="[1389,1399,1106,1131]" italics="true" pageId="28" pageNumber="29">-</emphasis>
<emphasis id="FFCC8919D838FFDBFAC9FBD2F724FB8C" box="[1389,1399,1106,1131]" italics="true" pageId="28" pageNumber="29">-</emphasis>
57°). Apex pointed. Embryonic whorls smoothly granulate. Subsequent whorls with finer, much more numerous ribs than the
<typeStatus box="[479,587,1176,1202]" pageId="28" pageNumber="29">lectotype</typeStatus>
<typeStatus id="1203EBA9D838FFDBFE7BFB18F018FB55" box="[479,587,1176,1202]" pageId="28" pageNumber="29" type="lectotype">lectotype</typeStatus>
of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[633,774,1176,1202]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis box="[633,774,1176,1202]" italics="true" pageId="28" pageNumber="29">D. r. radius</emphasis>
<taxonomicName id="0AB82E88D838FFDBFDDDFB19F155FB55" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[633,774,1176,1202]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis id="FFCC8919D838FFDBFDDDFB19F155FB55" box="[633,774,1176,1202]" italics="true" pageId="28" pageNumber="29">D. r. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[785,918,1176,1203]" pageId="28" pageNumber="29" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D838FFDBFCB5FB18F1C5FB54" box="[785,918,1176,1203]" pageId="28" pageNumber="29" rank="species">comb. nov.</taxonomicNameLabel>
(15
<emphasis box="[968,978,1177,1202]" italics="true" pageId="28" pageNumber="29">-</emphasis>
<emphasis id="FFCC8919D838FFDBFC6CFB19F181FB55" box="[968,978,1177,1202]" italics="true" pageId="28" pageNumber="29">-</emphasis>
23 per mm on penultimate whorl). Sutures relatively deep. Umbilicus closed or nearly so. Peristome complete, or incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 6-fold, consisting of: one lamella-like parietal tooth, sometimes slightly bifid; one slab-like palatal tooth, often bifid, but with parieto-palatal sinus barely present; one basal denticle; two shallow columellar denticles; and one deep-set columellar baffle, folded and sub-trifid, always visible. Shells and anatomy of juveniles unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="28" pageNumber="29" type="multiple">
<paragraph blockId="28.[189,421,1453,1480]" box="[189,421,1453,1480]" pageId="28" pageNumber="29">
<heading bold="true" box="[189,421,1453,1480]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis bold="true" box="[189,421,1453,1480]" pageId="28" pageNumber="29">Range and habitat</emphasis>
<subSubSection id="85A20680D838FFDBFF19FA2EF3A9FA11" pageId="28" pageNumber="29" type="biology_ecology">
<paragraph id="CD07550BD838FFDBFF19FA2EF3F6FA20" blockId="28.[189,421,1453,1480]" box="[189,421,1453,1480]" pageId="28" pageNumber="29">
<heading id="964FE267D838FFDBFF19FA2EF3F6FA20" bold="true" box="[189,421,1453,1480]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="FFCC8919D838FFDBFF19FA2EF3F6FA20" bold="true" box="[189,421,1453,1480]" pageId="28" pageNumber="29">Range and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[189,506,1500,1527]" box="[189,506,1500,1527]" pageId="28" pageNumber="29">
<collectingCountry box="[189,291,1501,1527]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
<paragraph id="CD07550BD838FFDBFF19FA5DF3A9FA11" blockId="28.[189,506,1500,1527]" box="[189,506,1500,1527]" pageId="28" pageNumber="29">
<collectingCountry id="B5AF159BD838FFDBFF19FA5DF370FA10" box="[189,291,1501,1527]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
; habitat unknown.
</paragraph>
</subSubSection>
<subSubSection lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="discussion">
<paragraph blockId="28.[189,303,1566,1592]" box="[189,303,1566,1592]" pageId="28" pageNumber="29">
<heading bold="true" box="[189,303,1566,1592]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis bold="true" box="[189,303,1566,1592]" pageId="28" pageNumber="29">Remarks</emphasis>
<subSubSection id="85A20680D838FFDAFF19F99EF394FBA8" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="discussion">
<paragraph id="CD07550BD838FFDBFF19F99EF37CF9DF" blockId="28.[189,303,1566,1592]" box="[189,303,1566,1592]" pageId="28" pageNumber="29">
<heading id="964FE267D838FFDBFF19F99EF37CF9DF" bold="true" box="[189,303,1566,1592]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
<emphasis id="FFCC8919D838FFDBFF19F99EF37CF9DF" bold="true" box="[189,303,1566,1592]" pageId="28" pageNumber="29">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="28.[189,1399,1612,2027]" lastBlockId="29.[189,1399,267,648]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<paragraph id="CD07550BD838FFDAFF19F9CCF052FD60" blockId="28.[189,1399,1612,2027]" lastBlockId="29.[189,1399,267,648]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
This
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<collectingCountry id="B5AF159BD838FFDBFF53F9CDF30AF980" box="[247,345,1613,1639]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
subspecies is here separated from its counterpart on Nzwani for the first time. Specimens of
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[189,336,1648,1674]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[189,336,1648,1674]" italics="true" pageId="28" pageNumber="29">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D838FFDBFF19F9F0F303F96D" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[189,336,1648,1674]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D838FFDBFF19F9F0F303F96D" box="[189,336,1648,1674]" italics="true" pageId="28" pageNumber="29">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[343,450,1648,1674]" pageId="28" pageNumber="29" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D838FFDBFEF3F9F0F391F96D" box="[343,450,1648,1674]" pageId="28" pageNumber="29" rank="genus">gen. nov.</taxonomicNameLabel>
species from
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<collectingCountry id="B5AF159BD838FFDBFDC0F9F0F095F96D" box="[612,710,1648,1674]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
were first discussed by
<bibRefCitation author="Morelet A." box="[990,1169,1648,1674]" pageId="28" pageNumber="29" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet (1881)</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFC7AF9F0F6C2F96D" author="Morelet A." box="[990,1169,1648,1674]" pageId="28" pageNumber="29" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet (1881)</bibRefCitation>
under his proposed replacement name for
<emphasis box="[465,659,1683,1709]" italics="true" pageId="28" pageNumber="29">
<emphasis id="FFCC8919D838FFDBFE75F914F0C0F94A" box="[465,659,1683,1709]" italics="true" pageId="28" pageNumber="29">
Pupa
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[538,659,1683,1709]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D838FFDBFDBEF913F0C0F94A" authorityName="Morelet" authorityYear="1877" box="[538,659,1683,1709]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
</emphasis>
(see above). He noted specimens from
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<collectingCountry id="B5AF159BD838FFDBFB25F913F6B0F94A" box="[1153,1251,1683,1709]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
were larger and had more complex dentition than those from
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<collectingRegion id="0F7C9BE9D838FFDBFCA9F936F125F937" box="[781,886,1718,1744]" country="Comoros" name="Anjouan" pageId="28" pageNumber="29">Anjouan</collectingRegion>
.
<bibRefCitation author="Tryon G. W." box="[898,1057,1718,1745]" pageId="28" pageNumber="29" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon (1885)</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFC26F937F672F936" author="Tryon G. W." box="[898,1057,1718,1745]" pageId="28" pageNumber="29" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon (1885)</bibRefCitation>
and
<bibRefCitation author="Kobelt W." box="[1118,1285,1718,1745]" pageId="28" pageNumber="29" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt (1905)</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFBFAF936F756F936" author="Kobelt W." box="[1118,1285,1718,1745]" pageId="28" pageNumber="29" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt (1905)</bibRefCitation>
followed Morelet in treating
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<emphasis box="[428,597,1754,1780]" italics="true" pageId="28" pageNumber="29">D. fischeriana</emphasis>
<taxonomicName id="0AB82E88D838FFDBFE08F95AF006F913" authorityName="Morelet" authorityYear="1877" box="[428,597,1754,1780]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D838FFDBFE08F95AF006F913" box="[428,597,1754,1780]" italics="true" pageId="28" pageNumber="29">D. fischeriana</emphasis>
</taxonomicName>
(i.e.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[666,826,1754,1780]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
<emphasis box="[666,826,1754,1780]" italics="true" pageId="28" pageNumber="29">D. minuscula</emphasis>
<taxonomicName id="0AB82E88D838FFDBFD3EF95AF169F913" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[666,826,1754,1780]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
<emphasis id="FFCC8919D838FFDBFD3EF95AF169F913" box="[666,826,1754,1780]" italics="true" pageId="28" pageNumber="29">D. minuscula</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[837,971,1753,1780]" pageId="28" pageNumber="29" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[981,1016,1754,1780]" italics="true" pageId="28" pageNumber="29">s.l.</emphasis>
<taxonomicNameLabel id="E4FF3462D838FFDBFCE1F959F198F913" box="[837,971,1753,1780]" pageId="28" pageNumber="29" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D838FFDBFC71F95AF1ABF913" box="[981,1016,1754,1780]" italics="true" pageId="28" pageNumber="29">s.l.</emphasis>
) as a variable species that was present on the “
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<collectingCountry id="B5AF159BD838FFDBFEDEF97DF3BAF8F0" box="[378,489,1789,1815]" name="Comoros" pageId="28" pageNumber="29">Comoros</collectingCountry>
Islands” (
<bibRefCitation author="Tryon G. W." box="[612,754,1789,1816]" pageId="28" pageNumber="29" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon 1885</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFDC0F97DF0A1F8FF" author="Tryon G. W." box="[612,754,1789,1816]" pageId="28" pageNumber="29" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon 1885</bibRefCitation>
) or on
<collectingRegion box="[844,948,1789,1815]" country="Comoros" name="Anjouan" pageId="28" pageNumber="29">Anjouan</collectingRegion>
<collectingRegion id="0F7C9BE9D838FFDBFCE8F97DF1E7F8F0" box="[844,948,1789,1815]" country="Comoros" name="Anjouan" pageId="28" pageNumber="29">Anjouan</collectingRegion>
, with a “grössere Form auf
<collectingCountry box="[1288,1389,1789,1815]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
<collectingCountry id="B5AF159BD838FFDBFAACF97DF73EF8F0" box="[1288,1389,1789,1815]" name="Mayotte" pageId="28" pageNumber="29">Mayotte</collectingCountry>
” (
<bibRefCitation author="Kobelt W." box="[197,349,1824,1851]" pageId="28" pageNumber="29" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt 1905</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFF61F8A0F30EF8DC" author="Kobelt W." box="[197,349,1824,1851]" pageId="28" pageNumber="29" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt 1905</bibRefCitation>
, although his dimensions of
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x
<quantity box="[800,870,1824,1851]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="28" pageNumber="29" unit="mm" value="2.0">2 mm</quantity>
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must be erroneous). Both authors appear to have copied
<bibRefCitation author="Morelet A." box="[341,541,1859,1886]" pageId="28" pageNumber="29" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet’s (1877)</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFEF1F8C3F04EF8B9" author="Morelet A." box="[341,541,1859,1886]" pageId="28" pageNumber="29" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet’s (1877)</bibRefCitation>
figure of the
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of
<emphasis box="[857,1054,1859,1885]" italics="true" pageId="28" pageNumber="29">
<emphasis id="FFCC8919D838FFDBFCFDF8C4F64DF8BA" box="[857,1054,1859,1885]" italics="true" pageId="28" pageNumber="29">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[933,1054,1859,1885]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D838FFDBFC01F8C3F64DF8BA" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[933,1054,1859,1885]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
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, unless they saw material very like it, and appear to have repeated (translated)
<bibRefCitation author="Morelet A." box="[915,1114,1895,1921]" pageId="28" pageNumber="29" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet’s (1881)</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFC37F8E7F609F866" author="Morelet A." box="[915,1114,1895,1921]" pageId="28" pageNumber="29" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet’s (1881)</bibRefCitation>
comments on dentition.
<bibRefCitation author="Tryon G. W." box="[189,345,1930,1957]" pageId="28" pageNumber="29" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon (1885)</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFF19F80BF30AF842" author="Tryon G. W." box="[189,345,1930,1957]" pageId="28" pageNumber="29" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon (1885)</bibRefCitation>
says “sometimes the parietal lamina is accompanied by a more profound very small tooth” but this may be a misinterpretation of
<bibRefCitation author="Morelet A." box="[637,836,1965,1991]" pageId="28" pageNumber="29" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet’s (1881)</bibRefCitation>
<bibRefCitation id="A92928FAD838FFDBFDD9F82DF117F820" author="Morelet A." box="[637,836,1965,1991]" pageId="28" pageNumber="29" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet’s (1881)</bibRefCitation>
“le pli pariétal peut être accompagné d’une très petite denticule plus profonde”, with “plus profonde” meaning ‘deeper’, rather than ‘profound’, i.e. ‘more obvious’ or ‘more important’. Morelet’s “denticule” is likely to refer to the bifid shape of the parietal tooth, rather than to an additional parietal denticle (like that of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[1016,1133,303,329]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
<emphasis box="[1016,1133,303,329]" italics="true" pageId="29" pageNumber="30">D. browni</emphasis>
<taxonomicName id="0AB82E88D839FFDAFC5CFEB0F63EFEAE" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="van Bruggen" baseAuthorityYear="1969" box="[1016,1133,303,329]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="browni" status="comb. nov.">
<emphasis id="FFCC8919D839FFDAFC5CFEB0F63EFEAE" box="[1016,1133,303,329]" italics="true" pageId="29" pageNumber="30">D. browni</emphasis>
</taxonomicName>
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<emphasis box="[1276,1310,303,329]" italics="true" pageId="29" pageNumber="30">s.l.</emphasis>
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<emphasis id="FFCC8919D839FFDAFB58FEAFF74DFEAE" box="[1276,1310,303,329]" italics="true" pageId="29" pageNumber="30">s.l.</emphasis>
) which is not present in any of the specimens of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[684,842,338,364]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
<emphasis box="[684,842,338,364]" italics="true" pageId="29" pageNumber="30">D. minuscula</emphasis>
<taxonomicName id="0AB82E88D839FFDAFD08FED3F119FE8B" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[684,842,338,364]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
<emphasis id="FFCC8919D839FFDAFD08FED3F119FE8B" box="[684,842,338,364]" italics="true" pageId="29" pageNumber="30">D. minuscula</emphasis>
</taxonomicName>
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<emphasis box="[991,1025,338,364]" italics="true" pageId="29" pageNumber="30">s.l.</emphasis>
<taxonomicNameLabel id="E4FF3462D839FFDAFCF7FED2F185FE8A" box="[851,982,338,365]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D839FFDAFC7BFED2F652FE8B" box="[991,1025,338,364]" italics="true" pageId="29" pageNumber="30">s.l.</emphasis>
we examined. It is not clear to which of the other teeth
<bibRefCitation author="Morelet A." box="[482,682,373,400]" pageId="29" pageNumber="30" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet’s (1881)</bibRefCitation>
<bibRefCitation id="A92928FAD839FFDAFE46FEF5F0F9FE77" author="Morelet A." box="[482,682,373,400]" pageId="29" pageNumber="30" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet’s (1881)</bibRefCitation>
“cinquième petite dent, dans la gorge de la coquille” refers, because all
<collectingCountry box="[327,425,410,436]" name="Mayotte" pageId="29" pageNumber="30">Mayotte</collectingCountry>
<collectingCountry id="B5AF159BD839FFDAFEE3FE1AF3FAFE53" box="[327,425,410,436]" name="Mayotte" pageId="29" pageNumber="30">Mayotte</collectingCountry>
specimens we examined have 6-fold dentition. It could even be the fifth tooth, i.e. the shallow columellar denticle, of the
<typeStatus box="[668,826,444,470]" pageId="29" pageNumber="30">paralectotype</typeStatus>
<typeStatus id="1203EBA9D839FFDAFD38FE3CF169FE31" box="[668,826,444,470]" pageId="29" pageNumber="30" type="paralectotype">paralectotype</typeStatus>
of
<emphasis box="[872,1072,444,470]" italics="true" pageId="29" pageNumber="30">
<emphasis id="FFCC8919D839FFDAFCCCFE3DF663FE31" box="[872,1072,444,470]" italics="true" pageId="29" pageNumber="30">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[951,1072,444,470]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D839FFDAFC13FE3CF663FE31" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[951,1072,444,470]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[1083,1219,444,471]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D839FFDAFB9FFE3CF690FE30" box="[1083,1219,444,471]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
, since
<bibRefCitation author="Morelet A." pageId="29" pageNumber="30" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet (1881)</bibRefCitation>
<bibRefCitation id="A92928FAD839FFDAFABEFE3CF358FE1D" author="Morelet A." pageId="29" pageNumber="30" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet (1881)</bibRefCitation>
is not explicit about the shell he is referring to. This raises another unfortunate possibility: that the
<typeStatus box="[236,394,515,541]" pageId="29" pageNumber="30">paralectotype</typeStatus>
<typeStatus id="1203EBA9D839FFDAFF48FD83F3D9FDFA" box="[236,394,515,541]" pageId="29" pageNumber="30" type="paralectotype">paralectotype</typeStatus>
of
<emphasis box="[439,639,515,541]" italics="true" pageId="29" pageNumber="30">
<emphasis id="FFCC8919D839FFDAFE13FD84F02CFDFA" box="[439,639,515,541]" italics="true" pageId="29" pageNumber="30">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[518,639,515,541]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D839FFDAFDA2FD83F02CFDFA" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[518,639,515,541]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[649,785,515,542]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D839FFDAFD2DFD83F142FDF9" box="[649,785,515,542]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
, apparently from Anjouan, was in fact a specimen without type status that was added later, and may even have come from
<collectingCountry box="[1038,1136,551,577]" name="Mayotte" pageId="29" pageNumber="30">Mayotte</collectingCountry>
<collectingCountry id="B5AF159BD839FFDAFBAAFDA7F623FDA6" box="[1038,1136,551,577]" name="Mayotte" pageId="29" pageNumber="30">Mayotte</collectingCountry>
(or elsewhere), which would accord with
<bibRefCitation author="Morelet A." box="[415,613,586,612]" pageId="29" pageNumber="30" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet’s (1881)</bibRefCitation>
<bibRefCitation id="A92928FAD839FFDAFE3BFDCAF036FD83" author="Morelet A." box="[415,613,586,612]" pageId="29" pageNumber="30" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet’s (1881)</bibRefCitation>
amendments to his description. However, we have no other reason to doubt the existing labels.
</paragraph>
<paragraph blockId="29.[189,1399,687,1103]" pageId="29" pageNumber="30">
<paragraph id="CD07550BD839FFDAFF19FD30F394FBA8" blockId="29.[189,1399,687,1103]" pageId="29" pageNumber="30">
We here introduce the name
<emphasis box="[535,733,687,713]" italics="true" pageId="29" pageNumber="30">D. m. mahorana</emphasis>
<emphasis id="FFCC8919D839FFDAFDB3FD30F08EFD2E" box="[535,733,687,713]" italics="true" pageId="29" pageNumber="30">D. m. mahorana</emphasis>
subsp. nov. for the larger, coeloconoid specimens from
<collectingCountry box="[189,290,723,749]" name="Mayotte" pageId="29" pageNumber="30">Mayotte</collectingCountry>
<collectingCountry id="B5AF159BD839FFDAFF19FD53F371FD0A" box="[189,290,723,749]" name="Mayotte" pageId="29" pageNumber="30">Mayotte</collectingCountry>
, the name
<taxonomicName authority="Morelet" authorityName="Morelet" authorityYear="1877" box="[419,651,723,749]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[419,549,723,749]" italics="true" pageId="29" pageNumber="30">fischeriana</emphasis>
<taxonomicName id="0AB82E88D839FFDAFE07FD53F0D8FD0A" authority="Morelet" authorityName="Morelet" authorityYear="1877" box="[419,651,723,749]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D839FFDAFE07FD53F076FD0A" box="[419,549,723,749]" italics="true" pageId="29" pageNumber="30">fischeriana</emphasis>
Morelet
</taxonomicName>
being unavailable (see above). They are indeed larger than the two specimens of
<emphasis box="[401,593,758,784]" italics="true" pageId="29" pageNumber="30">
<emphasis id="FFCC8919D839FFDAFE35FD77F002FCF7" box="[401,593,758,784]" italics="true" pageId="29" pageNumber="30">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[472,593,758,784]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D839FFDAFE7CFD76F002FCF7" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[472,593,758,784]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[600,730,758,785]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D839FFDAFDFCFD76F089FCF6" box="[600,730,758,785]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
(the size ranges do not overlap), are more elongate, have a more coeloconoid spire, finer and more numerous ribs, and 6-fold dentition including two shallow columellar denticles. These same features also distinguish them from
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1017,1129,829,855]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis box="[1017,1129,829,855]" italics="true" pageId="29" pageNumber="30">D. radius</emphasis>
<taxonomicName id="0AB82E88D839FFDAFC5DFCBEF63AFCB0" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1017,1129,829,855]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis id="FFCC8919D839FFDAFC5DFCBEF63AFCB0" box="[1017,1129,829,855]" italics="true" pageId="29" pageNumber="30">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1138,1269,829,855]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[1277,1311,829,855]" italics="true" pageId="29" pageNumber="30">s.l.</emphasis>
<taxonomicNameLabel id="E4FF3462D839FFDAFBD6FCBDF6A6FCB0" box="[1138,1269,829,855]" pageId="29" pageNumber="30" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D839FFDAFB59FCBDF74CFCB0" box="[1277,1311,829,855]" italics="true" pageId="29" pageNumber="30">s.l.</emphasis>
, which they overlap in size but only barely in the higher number of ribs per millimetre. Although as with
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<emphasis italics="true" pageId="29" pageNumber="30">D. radius</emphasis>
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<emphasis id="FFCC8919D839FFDAFAFEFCE1F35BFC79" italics="true" pageId="29" pageNumber="30">D. radius</emphasis>
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, it is difficult to distinguish intra- and interspecific variability, it is clear that the differences between the
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<emphasis id="FFCC8919D839FFDAFDC8FC28F17DFC26" box="[620,814,935,961]" italics="true" pageId="29" pageNumber="30">D. m. mahorana</emphasis>
subsp. nov. and the
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D. m.
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, each collected on different islands, are substantial and sufficient for most modern authors to consider them distinct species. Alternatively, and given the residual uncertainty about the
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D. m.
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, they may form part of a complex of subspecies. Again, further data is needed to address this.
</paragraph>
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439
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(
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<figureCitation box="[677,767,1178,1204]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="29" pageNumber="30">Figs 41</figureCitation>
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<figureCitation id="5583498ED839FFDAFD01FB1AF0ACFB53" box="[677,767,1178,1204]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="29" pageNumber="30">Figs 41</figureCitation>
,
<figureCitation box="[779,812,1178,1204]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="29" pageNumber="30">84</figureCitation>
<figureCitation id="5583498ED839FFDAFCAFFB1AF17FFB53" box="[779,812,1178,1204]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="29" pageNumber="30">84</figureCitation>
;
<tableCitation box="[823,910,1178,1204]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="29" pageNumber="30" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
<tableCitation id="803A60B0D839FFDAFC93FB1AF1DDFB53" box="[823,910,1178,1204]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetBox="[202,1925,710,1733]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="29" pageNumber="30" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="30" type="reference_group">
<paragraph blockId="29.[189,803,1243,1270]" box="[189,803,1243,1270]" pageId="29" pageNumber="30">
<treatmentCitationGroup box="[189,803,1243,1270]" pageId="29" pageNumber="30">
<treatmentCitation author="Verdcourt B." box="[189,723,1243,1270]" page="139" pageId="29" pageNumber="30" year="1994">
<taxonomicName ID-CoL="3HLWK" authority="Verdcourt, 1994: 137 - 139" authorityName="Verdcourt" authorityPageNumber="137 - 139" authorityYear="1994" box="[189,723,1243,1270]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="rondoensis">
<emphasis box="[189,412,1244,1270]" italics="true" pageId="29" pageNumber="30">Gulella rondoensis</emphasis>
<bibRefCitation author="Verdcourt B." box="[419,723,1243,1270]" pageId="29" pageNumber="30" pagination="137 - 139" refId="ref28031" refString="Verdcourt B. 1994. A new species of Gulella from SE. Tanzania (Pulmonata: Streptaxidae). Archiv fur Molluskenkunde 123 (1 / 6): 137 - 139." type="journal article" year="1994">Verdcourt, 1994: 137-139</bibRefCitation>
<subSubSection id="85A20680D839FFDAFF19FB5CF0F0FAD0" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="CD07550BD839FFDAFF19FB5CF170FB11" blockId="29.[189,803,1243,1270]" box="[189,803,1243,1270]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="EDA87225D839FFDAFF19FB5CF170FB11" box="[189,803,1243,1270]" pageId="29" pageNumber="30">
<treatmentCitation id="4C19731AD839FFDAFF19FB5CF080FB11" author="Verdcourt B." box="[189,723,1243,1270]" page="139" pageId="29" pageNumber="30" year="1994">
<taxonomicName id="0AB82E88D839FFDAFF19FB5CF080FB11" ID-CoL="3HLWK" authority="Verdcourt, 1994: 137 - 139" authorityName="Verdcourt" authorityPageNumber="137 - 139" authorityYear="1994" box="[189,723,1243,1270]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="rondoensis">
<emphasis id="FFCC8919D839FFDAFF19FB5CF3CFFB11" box="[189,412,1244,1270]" italics="true" pageId="29" pageNumber="30">Gulella rondoensis</emphasis>
<bibRefCitation id="A92928FAD839FFDAFE07FB5BF080FB11" author="Verdcourt B." box="[419,723,1243,1270]" pageId="29" pageNumber="30" pagination="137 - 139" refId="ref28031" refString="Verdcourt B. 1994. A new species of Gulella from SE. Tanzania (Pulmonata: Streptaxidae). Archiv fur Molluskenkunde 123 (1 / 6): 137 - 139." type="journal article" year="1994">Verdcourt, 1994: 137-139</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, fig. 1.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="29.[189,675,1309,1336]" box="[189,675,1309,1336]" pageId="29" pageNumber="30">
<treatmentCitationGroup box="[189,675,1309,1336]" pageId="29" pageNumber="30">
<taxonomicName ID-CoL="3HLWK" authorityName="Verdcourt" authorityYear="1994" box="[189,412,1309,1335]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="rondoensis">
<emphasis box="[189,412,1309,1335]" italics="true" pageId="29" pageNumber="30">Gulella rondoensis</emphasis>
<paragraph id="CD07550BD839FFDAFF19FA9DF0F0FAD0" blockId="29.[189,675,1309,1336]" box="[189,675,1309,1336]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="EDA87225D839FFDAFF19FA9DF0F0FAD0" box="[189,675,1309,1336]" pageId="29" pageNumber="30">
<taxonomicName id="0AB82E88D839FFDAFF19FA9DF3CFFAD0" ID-CoL="3HLWK" authorityName="Verdcourt" authorityYear="1994" box="[189,412,1309,1335]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="rondoensis">
<emphasis id="FFCC8919D839FFDAFF19FA9DF3CFFAD0" box="[189,412,1309,1335]" italics="true" pageId="29" pageNumber="30">Gulella rondoensis</emphasis>
</taxonomicName>
–
<treatmentCitation author="Verdcourt B." box="[441,671,1309,1335]" page="48" pageId="29" pageNumber="30" year="2006">
<bibRefCitation author="Verdcourt B." box="[441,671,1309,1335]" pageId="29" pageNumber="30" refId="ref28139" refString="Verdcourt B. 2006. A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi). Verdcourt B., Maidenhead, UK." type="book" year="2006">Verdcourt 2006: 48</bibRefCitation>
<treatmentCitation id="4C19731AD839FFDAFE1DFA9DF0CCFAD0" author="Verdcourt B." box="[441,671,1309,1335]" page="48" pageId="29" pageNumber="30" year="2006">
<bibRefCitation id="A92928FAD839FFDAFE1DFA9DF0CCFAD0" author="Verdcourt B." box="[441,671,1309,1335]" pageId="29" pageNumber="30" refId="ref28139" refString="Verdcourt B. 2006. A Revised List of the Non-marine Mollusca of East Africa (Kenya, Tanzania and Uganda, excluding Lake Malawi). Verdcourt B., Maidenhead, UK." type="book" year="2006">Verdcourt 2006: 48</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="29" pageNumber="30" type="materials_examined">
<paragraph blockId="29.[189,492,1375,1401]" box="[189,492,1375,1401]" pageId="29" pageNumber="30">
<heading bold="true" box="[189,492,1375,1401]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis bold="true" box="[189,492,1375,1401]" pageId="29" pageNumber="30">
<typeStatus box="[189,250,1375,1401]" pageId="29" pageNumber="30">Type</typeStatus>
<subSubSection id="85A20680D839FFDAFF19FADFF357F964" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="CD07550BD839FFDAFF19FADFF3BFFA9E" blockId="29.[189,492,1375,1401]" box="[189,492,1375,1401]" pageId="29" pageNumber="30">
<heading id="964FE267D839FFDAFF19FADFF3BFFA9E" bold="true" box="[189,492,1375,1401]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="FFCC8919D839FFDAFF19FADFF3BFFA9E" bold="true" box="[189,492,1375,1401]" pageId="29" pageNumber="30">
<typeStatus id="1203EBA9D839FFDAFF19FADFF2A9FA9E" box="[189,250,1375,1401]" pageId="29" pageNumber="30">Type</typeStatus>
material examined
</emphasis>
</heading>
</paragraph>
<paragraph blockId="29.[189,1399,1421,1555]" pageId="29" pageNumber="30">
<materialsCitation ID-GBIF-Occurrence="2610647312" collectingDate="1991-02-10" collectionCode="SMF" collectorName="Bidgood" country="Tanzania" latitude="-10.12" location="Albizia Durazz" longLatPrecision="779" longitude="39.22" municipality="Milicia Sim" pageId="29" pageNumber="30" specimenCode="SMF.310150" specimenCount="1" specimenCount-adult="1" stateProvince="Lindi District" typeStatus="holotype">
<collectingCountry box="[189,343,1422,1448]" name="Tanzania" pageId="29" pageNumber="30">TANZANIA</collectingCountry>
<paragraph id="CD07550BD839FFDAFF19FA0EF0F4F9F4" blockId="29.[189,1399,1421,1555]" pageId="29" pageNumber="30">
<materialsCitation id="7DD05F56D839FFDAFF19FA0EF0F4F9F4" ID-GBIF-Occurrence="2610647312" collectingDate="1991-02-10" collectionCode="SMF" collectorName="Bidgood &amp; Abdallah &amp; Vollesen" country="Tanzania" elevation="650" latitude="-10.12" location="Rondo Forest Reserve" longLatPrecision="779" longitude="39.22" municipality="Rondo Plateau" pageId="29" pageNumber="30" specimenCode="SMF.310150" specimenCount="1" specimenCount-adult="1" stateProvince="Lindi District" typeStatus="holotype">
<collectingCountry id="B5AF159BD839FFDAFF19FA0EF304FA4F" box="[189,343,1422,1448]" name="Tanzania" pageId="29" pageNumber="30">TANZANIA</collectingCountry>
:
<typeStatus box="[360,463,1422,1448]" pageId="29" pageNumber="30">holotype</typeStatus>
<specimenCode box="[477,633,1421,1448]" pageId="29" pageNumber="30">SMF.310150</specimenCode>
<typeStatus id="1203EBA9D839FFDAFECCFA0EF39CFA4F" box="[360,463,1422,1448]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
<specimenCode id="9D1EFD70D839FFDAFE79FA0DF02AFA4F" box="[477,633,1421,1448]" pageId="29" pageNumber="30">SMF.310150</specimenCode>
:
<specimenCount box="[651,717,1422,1448]" pageId="29" pageNumber="30" type="adult">1 ad.</specimenCount>
, Rondo Forest Reserve, Rondo Plateau,
<collectingRegion box="[1233,1398,1422,1448]" country="Tanzania" name="Lindi" pageId="29" pageNumber="30">Lindi District</collectingRegion>
<specimenCount id="DBBE9E82D839FFDAFD2FFA0EF09EFA4F" box="[651,717,1422,1448]" pageId="29" pageNumber="30" type="adult">1 ad.</specimenCount>
,
<location id="C86703D0D839FFDAFD45FA0EF1A5FA4F" LSID="urn:lsid:plazi:treatment:4511E41DD839FFD8FE7EFBF6F6ECFCF7:C86703D0D839FFDAFD45FA0EF1A5FA4F" box="[737,1014,1422,1448]" country="Tanzania" latitude="-10.12" longLatPrecision="779" longitude="39.22" municipality="Rondo Plateau" name="Rondo Forest Reserve" pageId="29" pageNumber="30" stateProvince="Lindi District">Rondo Forest Reserve</location>
,
<collectingMunicipality id="2D63CF71D839FFDAFBADFA0EF6EDFA4F" box="[1033,1214,1422,1448]" pageId="29" pageNumber="30">Rondo Plateau</collectingMunicipality>
,
<collectingRegion id="0F7C9BE9D839FFDAFB75FA0EF725FA4F" box="[1233,1398,1422,1448]" country="Tanzania" name="Lindi" pageId="29" pageNumber="30">Lindi District</collectingRegion>
(
<geoCoordinate box="[198,295,1457,1483]" degrees="10.12" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="-10.12">10.12°S</geoCoordinate>
<geoCoordinate id="A88C33CCD839FFDAFF62FA31F374FA2C" box="[198,295,1457,1483]" degrees="10.12" direction="south" orientation="latitude" pageId="29" pageNumber="30" precision="555" value="-10.12">10.12°S</geoCoordinate>
,
<geoCoordinate box="[308,408,1457,1483]" degrees="39.22" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="39.22">39.22°E</geoCoordinate>
<geoCoordinate id="A88C33CCD839FFDAFE90FA31F3CBFA2C" box="[308,408,1457,1483]" degrees="39.22" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="555" value="39.22">39.22°E</geoCoordinate>
), evergreen forest with
<taxonomicName authority="Sim" authorityName="Sim" box="[687,827,1457,1483]" class="Magnoliopsida" family="Moraceae" genus="Milicia" kingdom="Plantae" order="Rosales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<collectingMunicipality box="[687,827,1457,1483]" pageId="29" pageNumber="30">
<emphasis box="[687,772,1457,1483]" italics="true" pageId="29" pageNumber="30">Milicia</emphasis>
<taxonomicName id="0AB82E88D839FFDAFD0BFA31F168FA2C" authority="Sim" authorityName="Sim" box="[687,827,1457,1483]" class="Magnoliopsida" family="Moraceae" genus="Milicia" kingdom="Plantae" order="Rosales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="FFCC8919D839FFDAFD0BFA31F157FA2C" box="[687,772,1457,1483]" italics="true" pageId="29" pageNumber="30">Milicia</emphasis>
Sim
</collectingMunicipality>
</taxonomicName>
,
<taxonomicName authority="Durazz." authorityName="Durazz." box="[843,1026,1457,1484]" class="Magnoliopsida" family="Asteraceae" genus="Albizia" kingdom="Plantae" order="Asterales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<location LSID="urn:lsid:plazi:treatment:4511E41DD839FFD8FE7EFBF6F6ECFCF7:C86703D0D839FFDAFCEFFA31F1ADFA2B" box="[843,1022,1457,1484]" country="Tanzania" latitude="-10.12" longLatPrecision="779" longitude="39.22" municipality="Milicia Sim" name="Albizia Durazz" pageId="29" pageNumber="30" stateProvince="Lindi District">
<emphasis box="[843,926,1457,1483]" italics="true" pageId="29" pageNumber="30">Albizia</emphasis>
Durazz
</location>
.
<taxonomicName id="0AB82E88D839FFDAFCEFFA31F651FA2B" authority="Durazz." authorityName="Durazz." box="[843,1026,1457,1484]" class="Magnoliopsida" family="Asteraceae" genus="Albizia" kingdom="Plantae" order="Asterales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="FFCC8919D839FFDAFCEFFA31F1CDFA2C" box="[843,926,1457,1483]" italics="true" pageId="29" pageNumber="30">Albizia</emphasis>
Durazz.
</taxonomicName>
,
<location LSID="urn:lsid:plazi:treatment:4511E41DD839FFD8FE7EFBF6F6ECFCF7:C86703D0D839FFDAFBB5FA31F6C7FA2B" box="[1041,1172,1457,1484]" country="Tanzania" latitude="-10.12" longLatPrecision="779" longitude="39.22" municipality="Milicia Sim" name="Dialium L." pageId="29" pageNumber="30" stateProvince="Lindi District">
<taxonomicName authorityName="Linnaeus" authorityYear="1767" box="[1041,1137,1457,1483]" class="Magnoliopsida" family="Fabaceae" genus="Dialium" kingdom="Plantae" order="Fabales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis box="[1041,1137,1457,1483]" italics="true" pageId="29" pageNumber="30">Dialium</emphasis>
</taxonomicName>
<taxonomicName id="0AB82E88D839FFDAFBB5FA31F6C7FA2B" authorityName="Linnaeus" authorityYear="1767" box="[1041,1172,1457,1484]" class="Magnoliopsida" family="Fabaceae" genus="Dialium" kingdom="Plantae" order="Fabales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="FFCC8919D839FFDAFBB5FA31F622FA2C" box="[1041,1137,1457,1483]" italics="true" pageId="29" pageNumber="30">Dialium</emphasis>
L.
</location>
</taxonomicName>
, etc. in small gully and amphitheatre around a well on escarpment,
<quantity box="[752,827,1492,1519]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="29" pageNumber="30" unit="m" value="650.0">650 m</quantity>
<quantity id="0A40F8EED839FFDAFD54FA54F168FA08" box="[752,827,1492,1519]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="29" pageNumber="30" unit="m" value="650.0">
<elevation id="4695B238D839FFDAFD54FA54F168FA08" box="[752,827,1492,1519]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="29" pageNumber="30" unit="m" value="650.0">650 m</elevation>
</quantity>
alt., leg.
<collectorName box="[936,1040,1492,1518]" pageId="29" pageNumber="30">Bidgood</collectorName>
<collectorName id="604D30DDD839FFDAFC0CFA54F643FA09" box="[936,1040,1492,1518]" pageId="29" pageNumber="30">Bidgood</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:4511E41DD839FFD8FE7EFBF6F6ECFCF7:C86703D0D839FFDAFBBEFA54F740FA09" box="[1050,1299,1492,1519]" country="Tanzania" latitude="-10.12" longLatPrecision="779" longitude="39.22" municipality="Milicia Sim" name="Abdallah &amp; Vollesen" pageId="29" pageNumber="30" stateProvince="Lindi District">Abdallah &amp; Vollesen</location>
<collectorName id="604D30DDD839FFDAFBBEFA54F6D5FA09" box="[1050,1158,1492,1518]" pageId="29" pageNumber="30">Abdallah</collectorName>
&amp;
<collectorName id="604D30DDD839FFDAFB0FFA54F740FA09" box="[1195,1299,1492,1518]" pageId="29" pageNumber="30">Vollesen</collectorName>
,
<date pageId="29" pageNumber="30" value="1991-02-10">
<collectingDate pageId="29" pageNumber="30" value="1991-02-10">10 Feb. 1991</collectingDate>
<date id="B90673CBD839FFDAFABAFA55F2ABF9F5" pageId="29" pageNumber="30" value="1991-02-10">
<collectingDate id="A9428A23D839FFDAFABAFA55F2ABF9F5" pageId="29" pageNumber="30" value="1991-02-10">10 Feb. 1991</collectingDate>
</date>
(examined digital photograph only).
</materialsCitation>
</paragraph>
<paragraph blockId="29.[189,505,1593,1619]" box="[189,505,1593,1619]" pageId="29" pageNumber="30">
<heading bold="true" box="[189,505,1593,1619]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis bold="true" box="[189,505,1593,1619]" pageId="29" pageNumber="30">Other material examined</emphasis>
<paragraph id="CD07550BD839FFDAFF19F9B9F3AAF9B4" blockId="29.[189,505,1593,1619]" box="[189,505,1593,1619]" pageId="29" pageNumber="30">
<heading id="964FE267D839FFDAFF19F9B9F3AAF9B4" bold="true" box="[189,505,1593,1619]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="FFCC8919D839FFDAFF19F9B9F3AAF9B4" bold="true" box="[189,505,1593,1619]" pageId="29" pageNumber="30">Other material examined</emphasis>
</heading>
</paragraph>
<paragraph blockId="29.[189,260,1641,1667]" box="[189,260,1641,1667]" pageId="29" pageNumber="30">None.</paragraph>
<paragraph id="CD07550BD839FFDAFF19F9E9F357F964" blockId="29.[189,260,1641,1667]" box="[189,260,1641,1667]" pageId="29" pageNumber="30">None.</paragraph>
</subSubSection>
<subSubSection lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph blockId="29.[189,334,1706,1732]" box="[189,334,1706,1732]" pageId="29" pageNumber="30">
<heading bold="true" box="[189,334,1706,1732]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis bold="true" box="[189,334,1706,1732]" pageId="29" pageNumber="30">Description</emphasis>
<subSubSection id="85A20680D839FFD9FF19F92AF192FEAC" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
<paragraph id="CD07550BD839FFDAFF19F92AF31DF923" blockId="29.[189,334,1706,1732]" box="[189,334,1706,1732]" pageId="29" pageNumber="30">
<heading id="964FE267D839FFDAFF19F92AF31DF923" bold="true" box="[189,334,1706,1732]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="FFCC8919D839FFDAFF19F92AF31DF923" bold="true" box="[189,334,1706,1732]" pageId="29" pageNumber="30">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="29.[189,1399,1753,2027]" lastBlockId="30.[189,1398,267,331]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
<paragraph id="CD07550BD839FFD9FF19F959F192FEAC" blockId="29.[189,1399,1753,2027]" lastBlockId="30.[189,1398,267,331]" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
SHELL (
<figureCitation box="[276,364,1753,1779]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="29" pageNumber="30">Fig. 41</figureCitation>
<figureCitation id="5583498ED839FFDAFEB0F959F33FF914" box="[276,364,1753,1779]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="29" pageNumber="30">Fig. 41</figureCitation>
). Large (
<quantity box="[470,578,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="29" pageNumber="30" unit="mm" value="4.1">4.10 mm</quantity>
<quantity id="0A40F8EED839FFDAFE72F959F011F914" box="[470,578,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="29" pageNumber="30" unit="mm" value="4.1">4.10 mm</quantity>
high x
<quantity box="[670,776,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="mm" value="2.2">2.20 mm</quantity>
<quantity id="0A40F8EED839FFDAFD3AF959F15BF914" box="[670,776,1753,1779]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="29" pageNumber="30" unit="mm" value="2.2">2.20 mm</quantity>
wide), of 6.25 whorls. Subconical (maximum width being in bottom third of the shell, at body whorl). Spire narrowly acuminate, almost cyrtoconoid (convex) rather than coeloconoid (spire angle 52°). Apex sharply pointed. Embryonic whorls punctate or malleate, rather than merely granulate. Later whorls with relatively fine ribs (about 14 per mm on penultimate whorl). Sutures of intermediate depth. Umbilicus narrowly open. Peristome incomplete parietally. Dentition weak, 3-fold (although could be interpreted as 2-fold), consisting of: one lamellalike parietal tooth, with a swelling above it that recalls the one in
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[964,1109,1965,1991]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="conoidea">
<emphasis box="[964,1109,1965,1991]" italics="true" pageId="29" pageNumber="30">D. conoidea</emphasis>
<taxonomicName id="0AB82E88D839FFDAFC60F82EF606F820" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[964,1109,1965,1991]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="conoidea">
<emphasis id="FFCC8919D839FFDAFC60F82EF606F820" box="[964,1109,1965,1991]" italics="true" pageId="29" pageNumber="30">D. conoidea</emphasis>
</taxonomicName>
; one weak palatal tooth, not forming a parieto-palatal sinus; and one very weak, shallow columellar swelling. Further minute swellings just perceptible in the
<typeStatus box="[568,671,267,293]" pageId="30" pageNumber="31">holotype</typeStatus>
<typeStatus id="1203EBA9D83AFFD9FD9CFE8BF0CCFEC2" box="[568,671,267,293]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
(
<figureCitation box="[686,771,267,293]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="30" pageNumber="31">Fig. 41</figureCitation>
<figureCitation id="5583498ED83AFFD9FD0AFE8BF150FEC2" box="[686,771,267,293]" captionStart="Figs 41-42" captionStartId="30.[189,241,1930,1956]" captionTargetBox="[314,1273,931,1890]" captionTargetId="figure@30.[314,1273,931,1890]" captionTargetPageId="30" captionText="Figs 41-42. Adult shells of Dadagulella gen. nov. species. 41. D. rondoensis (Verdcourt, 1994) comb. nov., holotype, Rondo (photo by R. Janssen). 42. D. conoidea comb. nov., holotype, Kwamgumi (after Verdcourt 1996)." figureDoi="http://doi.org/10.5281/zenodo.3806788" httpUri="https://zenodo.org/record/3806788/files/figure.png" pageId="30" pageNumber="31">Fig. 41</figureCitation>
) were not noted by
<bibRefCitation author="Verdcourt B." box="[1005,1206,267,293]" pageId="30" pageNumber="31" pagination="137 - 139" refId="ref28031" refString="Verdcourt B. 1994. A new species of Gulella from SE. Tanzania (Pulmonata: Streptaxidae). Archiv fur Molluskenkunde 123 (1 / 6): 137 - 139." type="journal article" year="1994">Verdcourt (1994)</bibRefCitation>
<bibRefCitation id="A92928FAD83AFFD9FC49FE8BF6E5FEC2" author="Verdcourt B." box="[1005,1206,267,293]" pageId="30" pageNumber="31" pagination="137 - 139" refId="ref28031" refString="Verdcourt B. 1994. A new species of Gulella from SE. Tanzania (Pulmonata: Streptaxidae). Archiv fur Molluskenkunde 123 (1 / 6): 137 - 139." type="journal article" year="1994">Verdcourt (1994)</bibRefCitation>
who interpreted the dentition as 2-fold. Shells and anatomy of juveniles unknown.
</paragraph>
<paragraph blockId="30.[189,421,372,399]" box="[189,421,372,399]" pageId="30" pageNumber="31">
<heading bold="true" box="[189,421,372,399]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis bold="true" box="[189,421,372,399]" pageId="30" pageNumber="31">Range and habitat</emphasis>
</subSubSection>
<subSubSection id="85A20680D83AFFD9FF19FEF5F0ACFE58" pageId="30" pageNumber="31" type="biology_ecology">
<paragraph id="CD07550BD83AFFD9FF19FEF5F3F6FE69" blockId="30.[189,421,372,399]" box="[189,421,372,399]" pageId="30" pageNumber="31">
<heading id="964FE267D83AFFD9FF19FEF5F3F6FE69" bold="true" box="[189,421,372,399]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="FFCC8919D83AFFD9FF19FEF5F3F6FE69" bold="true" box="[189,421,372,399]" pageId="30" pageNumber="31">Range and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="30.[189,767,421,448]" box="[189,767,421,448]" pageId="30" pageNumber="31">
<paragraph id="CD07550BD83AFFD9FF19FE26F0ACFE58" blockId="30.[189,767,421,448]" box="[189,767,421,448]" pageId="30" pageNumber="31">
Forest at the
<typeStatus box="[341,392,422,448]" pageId="30" pageNumber="31">type</typeStatus>
<typeStatus id="1203EBA9D83AFFD9FEF1FE26F3DBFE27" box="[341,392,422,448]" pageId="30" pageNumber="31">type</typeStatus>
locality, southeastern
<collectingCountry box="[653,762,421,447]" name="Tanzania" pageId="30" pageNumber="31">Tanzania</collectingCountry>
<collectingCountry id="B5AF159BD83AFFD9FD29FE25F0A9FE58" box="[653,762,421,447]" name="Tanzania" pageId="30" pageNumber="31">Tanzania</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="discussion">
<paragraph blockId="30.[189,303,489,515]" box="[189,303,489,515]" pageId="30" pageNumber="31">
<heading bold="true" box="[189,303,489,515]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis bold="true" box="[189,303,489,515]" pageId="30" pageNumber="31">Remarks</emphasis>
<subSubSection id="85A20680D83AFFD8FF19FE69F6ECFCF7" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="discussion">
<paragraph id="CD07550BD83AFFD9FF19FE69F37CFDE4" blockId="30.[189,303,489,515]" box="[189,303,489,515]" pageId="30" pageNumber="31">
<heading id="964FE267D83AFFD9FF19FE69F37CFDE4" bold="true" box="[189,303,489,515]" fontSize="11" level="3" pageId="30" pageNumber="31" reason="3">
<emphasis id="FFCC8919D83AFFD9FF19FE69F37CFDE4" bold="true" box="[189,303,489,515]" pageId="30" pageNumber="31">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="30.[189,1399,537,826]" lastBlockId="31.[189,1399,267,785]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<paragraph id="CD07550BD83AFFD8FF19FD99F6ECFCF7" blockId="30.[189,1399,537,826]" lastBlockId="31.[189,1399,267,785]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
This species differs from
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[496,642,537,563]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis box="[496,642,537,563]" italics="true" pageId="30" pageNumber="31">D. conoidea</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FE54FD9AF0D1FDD4" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[496,642,537,563]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis id="FFCC8919D83AFFD9FE54FD9AF0D1FDD4" box="[496,642,537,563]" italics="true" pageId="30" pageNumber="31">D. conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[651,783,537,564]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83AFFD9FD2FFD99F15CFDD3" box="[651,783,537,564]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
in its weaker dentition, smaller size, and in having punctate apical whorls. Along with
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[603,746,575,601]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis box="[603,746,575,601]" italics="true" pageId="30" pageNumber="31">D. conoidea</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FDFFFDC0F0B9FDBE" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[603,746,575,601]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis id="FFCC8919D83AFFD9FDFFFDC0F0B9FDBE" box="[603,746,575,601]" italics="true" pageId="30" pageNumber="31">D. conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[753,882,575,601]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83AFFD9FD55FDBFF121FDBE" box="[753,882,575,601]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[937,1031,575,601]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="delta" status="sp. nov.">
<emphasis box="[937,1031,575,601]" italics="true" pageId="30" pageNumber="31">D. delta</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FC0DFDC0F654FDBE" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[937,1031,575,601]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="delta" status="sp. nov.">
<emphasis id="FFCC8919D83AFFD9FC0DFDC0F654FDBE" box="[937,1031,575,601]" italics="true" pageId="30" pageNumber="31">D. delta</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1038,1128,575,601]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83AFFD9FBAAFDBFF63BFDBE" box="[1038,1128,575,601]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
it has weaker dentition than other
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[314,461,612,638]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[314,461,612,638]" italics="true" pageId="30" pageNumber="31">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FE9EFDE4F39EFD99" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[314,461,612,638]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83AFFD9FE9EFDE4F39EFD99" box="[314,461,612,638]" italics="true" pageId="30" pageNumber="31">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[468,575,613,639]" pageId="30" pageNumber="31" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83AFFD9FE70FDE5F06CFD98" box="[468,575,613,639]" pageId="30" pageNumber="31" rank="genus">gen. nov.</taxonomicNameLabel>
However, both
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[762,926,612,638]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis box="[762,926,612,638]" italics="true" pageId="30" pageNumber="31">D. rondoensis</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FD5EFDE5F1CDFD99" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[762,926,612,638]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis id="FFCC8919D83AFFD9FD5EFDE5F1CDFD99" box="[762,926,612,638]" italics="true" pageId="30" pageNumber="31">D. rondoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[933,1062,612,639]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83AFFD9FC01FDE4F675FD98" box="[933,1062,612,639]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[1119,1262,612,638]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis box="[1119,1262,612,638]" italics="true" pageId="30" pageNumber="31">D. conoidea</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FBFBFDE5F6BDFD99" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[1119,1262,612,638]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis id="FFCC8919D83AFFD9FBFBFDE5F6BDFD99" box="[1119,1262,612,638]" italics="true" pageId="30" pageNumber="31">D. conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1269,1399,612,639]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83AFFD9FB51FDE4F724FD98" box="[1269,1399,612,639]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
are distinctively more conical, i.e. less ovate than other
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[828,975,650,676]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[828,975,650,676]" italics="true" pageId="30" pageNumber="31">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FC98FD0AF19CFD43" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[828,975,650,676]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83AFFD9FC98FD0AF19CFD43" box="[828,975,650,676]" italics="true" pageId="30" pageNumber="31">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[980,1086,650,676]" pageId="30" pageNumber="31" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83AFFD9FC70FD0AF66DFD43" box="[980,1086,650,676]" pageId="30" pageNumber="31" rank="genus">gen. nov.</taxonomicNameLabel>
These two are attributed to the genus somewhat speculatively, on the basis of their acuminate spire and pointed apex. These features mean they do not obviously fit into
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1856" box="[623,711,724,750]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis box="[623,711,724,750]" italics="true" pageId="30" pageNumber="31">Gulella</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FDCBFD54F094FD09" authorityName="L. Pfeiffer" authorityYear="1856" box="[623,711,724,750]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus">
<emphasis id="FFCC8919D83AFFD9FDCBFD54F094FD09" box="[623,711,724,750]" italics="true" pageId="30" pageNumber="31">Gulella</emphasis>
</taxonomicName>
or any of its named subgenera, or indeed other plausible streptaxid genera. No anatomical or juvenile shell data are available for either species, both being known from single specimens. The punctuate apex of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[821,985,799,825]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis box="[821,985,799,825]" italics="true" pageId="30" pageNumber="31">D. rondoensis</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FC91FCA0F18AFCDE" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[821,985,799,825]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis id="FFCC8919D83AFFD9FC91FCA0F18AFCDE" box="[821,985,799,825]" italics="true" pageId="30" pageNumber="31">D. rondoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[991,1123,799,826]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83AFFD9FC7BFC9FF630FCDD" box="[991,1123,799,826]" pageId="30" pageNumber="31" rank="species">comb. nov.</taxonomicNameLabel>
, unique in
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[1252,1399,799,825]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[1252,1399,799,825]" italics="true" pageId="30" pageNumber="31">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83AFFD9FB40FC9FF724FCDE" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[1252,1399,799,825]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="30" pageNumber="31" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83AFFD9FB40FC9FF724FCDE" box="[1252,1399,799,825]" italics="true" pageId="30" pageNumber="31">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[189,301,268,294]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FF19FE8CF37EFEC1" box="[189,301,268,294]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
, may not be significant in this respect (see
<bibRefCitation author="Rowson B." box="[827,1006,267,294]" pageId="31" pageNumber="32" pagination="221 - 228" refId="ref27244" refString="Rowson B. 2007 b. Gulella (Juventigulella) ngerezae spec. nov. (Gastropoda, Pulmonata, Streptaxidae), a new endemic land snail from the Ukaguru Mountains, Tanzania. Basteria 71 (4 - 6): 221 - 228." type="journal article" year="2007">Rowson 2007b</bibRefCitation>
<bibRefCitation id="A92928FAD83BFFD8FC9FFE8CF1BDFEC2" author="Rowson B." box="[827,1006,267,294]" pageId="31" pageNumber="32" pagination="221 - 228" refId="ref27244" refString="Rowson B. 2007 b. Gulella (Juventigulella) ngerezae spec. nov. (Gastropoda, Pulmonata, Streptaxidae), a new endemic land snail from the Ukaguru Mountains, Tanzania. Basteria 71 (4 - 6): 221 - 228." type="journal article" year="2007">Rowson 2007b</bibRefCitation>
for a discussion on the value of apical sculpture in distinguishing African streptaxid genera). The apex of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[1080,1227,305,331]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis box="[1080,1227,305,331]" italics="true" pageId="31" pageNumber="32">D. conoidea</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FB9CFEB2F698FEAC" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[1080,1227,305,331]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FB9CFEB2F698FEAC" box="[1080,1227,305,331]" italics="true" pageId="31" pageNumber="32">D. conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1237,1369,305,332]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FB71FEB1F70AFEAB" box="[1237,1369,305,332]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
is less obviously punctuate, although it may have been worn smooth (
<bibRefCitation author="Verdcourt B." box="[983,1172,343,370]" pageId="31" pageNumber="32" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">Verdcourt 1996</bibRefCitation>
<bibRefCitation id="A92928FAD83BFFD8FC73FED7F6C7FE95" author="Verdcourt B." box="[983,1172,343,370]" pageId="31" pageNumber="32" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">Verdcourt 1996</bibRefCitation>
). In the description of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[223,389,381,407]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis box="[223,389,381,407]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FF7BFEFDF3D6FE70" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[223,389,381,407]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FF7BFEFDF3D6FE70" box="[223,389,381,407]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[398,533,381,407]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FE2AFEFDF046FE70" box="[398,533,381,407]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
,
<bibRefCitation author="Verdcourt B." box="[546,749,381,407]" pageId="31" pageNumber="32" pagination="137 - 139" refId="ref28031" refString="Verdcourt B. 1994. A new species of Gulella from SE. Tanzania (Pulmonata: Streptaxidae). Archiv fur Molluskenkunde 123 (1 / 6): 137 - 139." type="journal article" year="1994">Verdcourt (1994)</bibRefCitation>
<bibRefCitation id="A92928FAD83BFFD8FD86FEFDF0BEFE70" author="Verdcourt B." box="[546,749,381,407]" pageId="31" pageNumber="32" pagination="137 - 139" refId="ref28031" refString="Verdcourt B. 1994. A new species of Gulella from SE. Tanzania (Pulmonata: Streptaxidae). Archiv fur Molluskenkunde 123 (1 / 6): 137 - 139." type="journal article" year="1994">Verdcourt (1994)</bibRefCitation>
discussed a resemblance only to
<taxonomicName authority="(Crosse, 1885)" baseAuthorityName="Crosse" baseAuthorityYear="1885" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="galactochila">
<emphasis box="[1154,1399,381,407]" italics="true" pageId="31" pageNumber="32">Gulella galactochila</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FB26FEFDF33DFE5A" authority="(Crosse, 1885)" baseAuthorityName="Crosse" baseAuthorityYear="1885" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="galactochila">
<emphasis id="FFCC8919D83BFFD8FB26FEFDF724FE70" box="[1154,1399,381,407]" italics="true" pageId="31" pageNumber="32">Gulella galactochila</emphasis>
(Crosse, 1885)
</taxonomicName>
, a much larger and more broadly acuminate Tanzanian species that we consider to lack the characteristic features of
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[523,670,456,482]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[523,670,456,482]" italics="true" pageId="31" pageNumber="32">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FDAFFE48F0CDFE05" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[523,670,456,482]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83BFFD8FDAFFE48F0CDFE05" box="[523,670,456,482]" italics="true" pageId="31" pageNumber="32">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[676,782,457,483]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FD00FE49F15DFE04" box="[676,782,457,483]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
Although
<taxonomicName baseAuthorityName="Crosse" baseAuthorityYear="1885" box="[903,1085,456,483]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="galactochila">
<emphasis box="[903,1085,456,483]" italics="true" pageId="31" pageNumber="32">G. galactochila</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FC23FE49F66EFE05" baseAuthorityName="Crosse" baseAuthorityYear="1885" box="[903,1085,456,483]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="galactochila">
<emphasis id="FFCC8919D83BFFD8FC23FE49F66EFE05" box="[903,1085,456,483]" italics="true" pageId="31" pageNumber="32">G. galactochila</emphasis>
</taxonomicName>
has not been dissected, the anatomy of another species very similar to it (
<taxonomicName authority="van Bruggen, 2003" authorityName="van Bruggen" authorityYear="2003" box="[719,1146,494,520]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="udzungwensis">
<emphasis box="[719,917,494,520]" italics="true" pageId="31" pageNumber="32">G. udzungwensis</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FD6BFE6EF629FDEF" authority="van Bruggen, 2003" authorityName="van Bruggen" authorityYear="2003" box="[719,1146,494,520]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="udzungwensis">
<emphasis id="FFCC8919D83BFFD8FD6BFE6EF1C6FDEF" box="[719,917,494,520]" italics="true" pageId="31" pageNumber="32">G. udzungwensis</emphasis>
van
<bibRefCitation author="Bruggen A. C. van" box="[972,1146,494,520]" pageId="31" pageNumber="32" pagination="87 - 90" refId="ref26174" refString="Bruggen A. C. van 2003. Gulella udzungwensis spec. nov. (Gastropoda, Pulmonata, Streptaxidae), a new species from the Udzungwa Mountains in Tanzania. Basteria 67 (1 - 3): 87 - 90." type="journal article" year="2003">Bruggen, 2003</bibRefCitation>
<bibRefCitation id="A92928FAD83BFFD8FC68FE6EF629FDEF" author="Bruggen A. C. van" box="[972,1146,494,520]" pageId="31" pageNumber="32" pagination="87 - 90" refId="ref26174" refString="Bruggen A. C. van 2003. Gulella udzungwensis spec. nov. (Gastropoda, Pulmonata, Streptaxidae), a new species from the Udzungwa Mountains in Tanzania. Basteria 67 (1 - 3): 87 - 90." type="journal article" year="2003">Bruggen, 2003</bibRefCitation>
</taxonomicName>
) lacks the anatomical features of
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[322,469,532,558]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[322,469,532,558]" italics="true" pageId="31" pageNumber="32">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FEE6FD94F386FDC9" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[322,469,532,558]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83BFFD8FEE6FD94F386FDC9" box="[322,469,532,558]" italics="true" pageId="31" pageNumber="32">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[476,584,532,558]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FE78FD94F01BFDC9" box="[476,584,532,558]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
(Rowson unpublished). In his discussion of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[1110,1254,532,558]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis box="[1110,1254,532,558]" italics="true" pageId="31" pageNumber="32">D. conoidea</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FBF2FD94F6B5FDC9" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[1110,1254,532,558]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FBF2FD94F6B5FDC9" box="[1110,1254,532,558]" italics="true" pageId="31" pageNumber="32">D. conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1261,1394,532,558]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FB49FD94F721FDC9" box="[1261,1394,532,558]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
,
<bibRefCitation author="Verdcourt B." box="[189,392,569,596]" pageId="31" pageNumber="32" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">Verdcourt (1996)</bibRefCitation>
<bibRefCitation id="A92928FAD83BFFD8FF19FDB9F3DBFDB3" author="Verdcourt B." box="[189,392,569,596]" pageId="31" pageNumber="32" pagination="135 - 137" refId="ref28065" refString="Verdcourt B. 1996. A new species of Gulella Pfr. (Gastropoda Pulmonata, Streptaxidae) from NE. Tanzania. Basteria 59 (4 - 6): 135 - 137." type="journal article" year="1996">Verdcourt (1996)</bibRefCitation>
made no reference to either
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[744,911,569,595]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis box="[744,911,569,595]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FD4CFDBAF1DCFDB4" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[744,911,569,595]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FD4CFDBAF1DCFDB4" box="[744,911,569,595]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[920,1053,569,596]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FC3CFDB9F64EFDB3" box="[920,1053,569,596]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
or
<taxonomicName baseAuthorityName="Crosse" baseAuthorityYear="1885" box="[1096,1282,569,596]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="galactochila">
<emphasis box="[1096,1282,569,596]" italics="true" pageId="31" pageNumber="32">G. galactochila</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FBECFDBAF751FDB4" baseAuthorityName="Crosse" baseAuthorityYear="1885" box="[1096,1282,569,596]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="galactochila">
<emphasis id="FFCC8919D83BFFD8FBECFDBAF751FDB4" box="[1096,1282,569,596]" italics="true" pageId="31" pageNumber="32">G. galactochila</emphasis>
</taxonomicName>
, but only to two species that we here treat in
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[621,768,607,633]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[621,768,607,633]" italics="true" pageId="31" pageNumber="32">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FDC9FDDFF153FD9E" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[621,768,607,633]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83BFFD8FDC9FDDFF153FD9E" box="[621,768,607,633]" italics="true" pageId="31" pageNumber="32">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[777,886,608,634]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FCADFDE0F125FD9D" box="[777,886,608,634]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
(
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[905,1044,607,633]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis box="[905,1044,607,633]" italics="true" pageId="31" pageNumber="32">D. r. radius</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FC2DFDE0F647FD9E" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[905,1044,607,633]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis id="FFCC8919D83BFFD8FC2DFDE0F647FD9E" box="[905,1044,607,633]" italics="true" pageId="31" pageNumber="32">D. r. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1053,1185,607,634]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FBB9FDDFF6F2FD9D" box="[1053,1185,607,634]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1962" box="[1246,1398,607,633]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="cuspidata" status="comb. nov.">
<emphasis box="[1246,1398,607,633]" italics="true" pageId="31" pageNumber="32">D. cuspidata</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FB7AFDE0F725FD9E" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1962" box="[1246,1398,607,633]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="cuspidata" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FB7AFDE0F725FD9E" box="[1246,1398,607,633]" italics="true" pageId="31" pageNumber="32">D. cuspidata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[189,325,645,671]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FF19FD05F316FD78" box="[189,325,645,671]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
). We contend firstly that
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[628,795,645,671]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis box="[628,795,645,671]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
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<emphasis id="FFCC8919D83BFFD8FDD0FD06F148FD78" box="[628,795,645,671]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
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and
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<emphasis box="[997,1143,645,671]" italics="true" pageId="31" pageNumber="32">D. conoidea</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FC41FD06F624FD78" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" box="[997,1143,645,671]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FC41FD06F624FD78" box="[997,1143,645,671]" italics="true" pageId="31" pageNumber="32">D. conoidea</emphasis>
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<taxonomicNameLabel box="[1152,1284,645,671]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FB24FD05F757FD78" box="[1152,1284,645,671]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
are more similar to one another than either is to
<taxonomicName baseAuthorityName="Crosse" baseAuthorityYear="1885" box="[650,834,683,709]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="galactochila">
<emphasis box="[650,834,683,709]" italics="true" pageId="31" pageNumber="32">G. galactochila</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FD2EFD2BF111FD22" baseAuthorityName="Crosse" baseAuthorityYear="1885" box="[650,834,683,709]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="galactochila">
<emphasis id="FFCC8919D83BFFD8FD2EFD2BF111FD22" box="[650,834,683,709]" italics="true" pageId="31" pageNumber="32">G. galactochila</emphasis>
</taxonomicName>
, and secondly that the resemblance between
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis italics="true" pageId="31" pageNumber="32">D. conoidea</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FAFEFD2CF37AFD0D" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1996" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conoidea" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FAFEFD2CF37AFD0D" italics="true" pageId="31" pageNumber="32">D. conoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[303,434,720,747]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FE8BFD50F3E1FD0C" box="[303,434,720,747]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
,
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[445,577,720,746]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis box="[445,577,720,746]" italics="true" pageId="31" pageNumber="32">D. r. radius</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FE19FD51F012FD0D" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[445,577,720,746]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis id="FFCC8919D83BFFD8FE19FD51F012FD0D" box="[445,577,720,746]" italics="true" pageId="31" pageNumber="32">D. r. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[583,712,720,747]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FDE3FD50F09BFD0C" box="[583,712,720,747]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
and
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1962" box="[767,916,720,746]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="cuspidata" status="comb. nov.">
<emphasis box="[767,916,720,746]" italics="true" pageId="31" pageNumber="32">D. cuspidata</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FD5BFD51F1C7FD0D" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1962" box="[767,916,720,746]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="cuspidata" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FD5BFD51F1C7FD0D" box="[767,916,720,746]" italics="true" pageId="31" pageNumber="32">D. cuspidata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[922,1051,720,747]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FC3EFD50F648FD0C" box="[922,1051,720,747]" pageId="31" pageNumber="32" rank="species">comb. nov.</taxonomicNameLabel>
extends also to
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[1236,1399,720,746]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov">
<emphasis box="[1236,1399,720,746]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FB70FD51F724FD0D" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Verdcourt" baseAuthorityYear="1994" box="[1236,1399,720,746]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="rondoensis" status="comb. nov.">
<emphasis id="FFCC8919D83BFFD8FB70FD51F724FD0D" box="[1236,1399,720,746]" italics="true" pageId="31" pageNumber="32">D. rondoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[189,314,758,785]" pageId="31" pageNumber="32" rank="species">comb. nov</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FF19FD76F369FCF6" box="[189,314,758,785]" pageId="31" pageNumber="32" rank="species">comb. nov</taxonomicNameLabel>
.. Our attribution of them to
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[643,790,758,784]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[643,790,758,784]" italics="true" pageId="31" pageNumber="32">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83BFFD8FD27FD76F145FCF7" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[643,790,758,784]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83BFFD8FD27FD76F145FCF7" box="[643,790,758,784]" italics="true" pageId="31" pageNumber="32">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[797,904,759,785]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83BFFD8FCB9FD77F1DBFCF6" box="[797,904,759,785]" pageId="31" pageNumber="32" rank="genus">gen. nov.</taxonomicNameLabel>
reflects this point of view.
</paragraph>
</subSubSection>

695
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<taxonomicName ID-CoL="5DZBW" authority="minuscula (Morelet, 1877)" authorityName="minuscula (Morelet" authorityYear="1877" box="[420,1030,739,766]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="subSpecies" species="minuscula" status="comb. nov." subSpecies="minuscula">
<emphasis bold="true" box="[420,837,739,766]" italics="true" pageId="26" pageNumber="27">Dadagulella minuscula minuscula</emphasis>
<treatment id="4511E41DD83EFFDCFE00FD63F6D1F80C" ID-DOI="http://doi.org/10.5281/zenodo.3815767" ID-GBIF-Taxon="163971584" ID-Zenodo-Dep="3815767" LSID="urn:lsid:plazi:treatment:4511E41DD83EFFDCFE00FD63F6D1F80C" httpUri="http://treatment.plazi.org/id/4511E41DD83EFFDCFE00FD63F6D1F80C" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" scope_class="Gastropoda" scope_family="Streptaxidae" scope_order="Stylommatophora" scope_superFamily="Streptaxoidea">
<subSubSection id="85A20680D83EFFDDFE00FD63F6C3FD19" box="[420,1168,739,766]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="CD07550BD83EFFDDFE00FD63F6C3FD19" blockId="26.[420,1168,739,801]" box="[420,1168,739,766]" pageId="26" pageNumber="27">
<taxonomicName id="0AB82E88D83EFFDDFE00FD63F655FD19" ID-CoL="5DZBW" authority="minuscula (Morelet, 1877)" authorityName="minuscula (Morelet" authorityYear="1877" box="[420,1030,739,766]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="subSpecies" species="minuscula" status="comb. nov." subSpecies="minuscula">
<emphasis id="FFCC8919D83EFFDDFE00FD63F116FD19" bold="true" box="[420,837,739,766]" italics="true" pageId="26" pageNumber="27">Dadagulella minuscula minuscula</emphasis>
(
<bibRefCitation author="Morelet A." box="[853,1022,739,766]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877</bibRefCitation>
<bibRefCitation id="A92928FAD83EFFDDFCF1FD63F1ADFD19" author="Morelet A." box="[853,1022,739,766]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel box="[1038,1168,739,766]" pageId="26" pageNumber="27" rank="species">comb. nov.</taxonomicNameLabel>
<figureCitation box="[616,745,775,801]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="26" pageNumber="27">Figs 38-39</figureCitation>
,
<figureCitation box="[757,829,775,801]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="26" pageNumber="27">62-63</figureCitation>
,
<figureCitation box="[840,873,775,801]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="26" pageNumber="27">84</figureCitation>
;
<tableCitation box="[885,972,775,801]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="26" pageNumber="27" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
<taxonomicNameLabel id="E4FF3462D83EFFDDFBAAFD63F6C3FD19" box="[1038,1168,739,766]" pageId="26" pageNumber="27" rank="species">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="reference_group">
<paragraph blockId="26.[189,988,841,904]" box="[189,619,841,867]" pageId="26" pageNumber="27">
<treatmentCitationGroup box="[189,619,841,867]" pageId="26" pageNumber="27">
<treatmentCitation author="Morelet A." box="[189,616,841,867]" page="340" pageId="26" pageNumber="27" year="1877">
<taxonomicName ID-CoL="33XQL" authority="Morelet, 1877: 340" authorityName="Morelet" authorityPageNumber="340" authorityYear="1877" box="[189,616,841,867]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[189,251,842,867]" italics="true" pageId="26" pageNumber="27">Pupa</emphasis>
<emphasis box="[258,379,841,867]" italics="true" pageId="26" pageNumber="27">minuscula</emphasis>
<bibRefCitation author="Morelet A." box="[386,616,841,867]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877: 340</bibRefCitation>
<subSubSection id="85A20680D83EFFDDFDCCFC87F19FFCC6" box="[616,972,775,801]" pageId="26" pageNumber="27" type="description">
<paragraph id="CD07550BD83EFFDDFDCCFC87F19FFCC6" blockId="26.[420,1168,739,801]" box="[616,972,775,801]" pageId="26" pageNumber="27">
<figureCitation id="5583498ED83EFFDDFDCCFC87F0BAFCC6" box="[616,745,775,801]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="26" pageNumber="27">Figs 38-39</figureCitation>
,
<figureCitation id="5583498ED83EFFDDFD51FC87F16EFCC6" box="[757,829,775,801]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="26" pageNumber="27">62-63</figureCitation>
,
<figureCitation id="5583498ED83EFFDDFCECFC87F13AFCC6" box="[840,873,775,801]" captionStart="Fig" captionStartId="40.[189,232,1930,1956]" captionTargetBox="[193,1393,858,1884]" captionTargetId="figure@40.[189,1399,855,1890]" captionTargetPageId="40" captionText="Fig. 84. Distribution of Dadagulella gen. nov. species, based on published records and material examined here (for additional records of D. meredithae comb. nov. in northern Malawi see van Bruggen 2000). The inset on the left corresponds to the map on the right." figureDoi="http://doi.org/10.5281/zenodo.3806812" httpUri="https://zenodo.org/record/3806812/files/figure.png" pageId="26" pageNumber="27">84</figureCitation>
;
<tableCitation id="803A60B0D83EFFDDFCD1FC87F19FFCC6" box="[885,972,775,801]" captionStart="Table 1" captionStartId="3.[187,256,550,578]" captionTargetBox="[202,1925,710,1733]" captionTargetId="graphics@3.[187,1941,696,1740]" captionTargetPageId="3" captionText="Table 1. Measurements and dentition scores for adult shells of Dadagulella gen. nov. species and subspecies. Ranges are given with the mean (for measurements) or mode (for tooth counts) in brackets when n (number of shells measured) was greater than 2. Scores and measurements mentioned in the Table were taken either from all available adult material or from literature." httpUri="http://table.plazi.org/id/99C70583D827FFC4FF1FFDA6F602FD6F" pageId="26" pageNumber="27" tableUuid="99C70583D827FFC4FF1FFDA6F602FD6F">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="85A20680D83EFFDDFF19FCCAF011FB22" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="CD07550BD83EFFDDFF19FCCAF038FC84" blockId="26.[189,988,841,904]" box="[189,619,841,867]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="EDA87225D83EFFDDFF19FCCAF038FC84" box="[189,619,841,867]" pageId="26" pageNumber="27">
<treatmentCitation id="4C19731AD83EFFDDFF19FCCAF03BFC84" author="Morelet A." box="[189,616,841,867]" page="340" pageId="26" pageNumber="27" year="1877">
<taxonomicName id="0AB82E88D83EFFDDFF19FCCAF03BFC84" ID-CoL="33XQL" authority="Morelet, 1877: 340" authorityName="Morelet" authorityPageNumber="340" authorityYear="1877" box="[189,616,841,867]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="minuscula">
<emphasis id="FFCC8919D83EFFDDFF19FCCAF328FC84" box="[189,379,841,867]" italics="true" pageId="26" pageNumber="27">Pupa minuscula</emphasis>
<bibRefCitation id="A92928FAD83EFFDDFE26FCC9F03BFC84" author="Morelet A." box="[386,616,841,867]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877: 340</bibRefCitation>
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.
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</paragraph>
<paragraph blockId="26.[189,988,841,904]" box="[189,988,877,904]" pageId="26" pageNumber="27">
<emphasis box="[189,251,878,903]" italics="true" pageId="26" pageNumber="27">Pupa</emphasis>
<taxonomicName authority="Morelet, 1877" authorityName="Morelet" authorityYear="1877" box="[258,566,877,903]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[258,388,877,903]" italics="true" pageId="26" pageNumber="27">fischeriana</emphasis>
<bibRefCitation author="Morelet A." box="[396,566,877,903]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877</bibRefCitation>
<paragraph id="CD07550BD83EFFDDFF19FCEEF18FFC6F" blockId="26.[189,988,841,904]" box="[189,988,877,904]" pageId="26" pageNumber="27">
<emphasis id="FFCC8919D83EFFDDFF19FCEEF2A8FC60" box="[189,251,878,903]" italics="true" pageId="26" pageNumber="27">Pupa</emphasis>
<taxonomicName id="0AB82E88D83EFFDDFEA6FCEDF065FC60" authority="Morelet, 1877" authorityName="Morelet" authorityYear="1877" box="[258,566,877,903]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D83EFFDDFEA6FCEDF3D7FC60" box="[258,388,877,903]" italics="true" pageId="26" pageNumber="27">fischeriana</emphasis>
<bibRefCitation id="A92928FAD83EFFDDFE28FCEDF065FC60" author="Morelet A." box="[396,566,877,903]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877</bibRefCitation>
</taxonomicName>
: pl. XII, fig. 5 (objective synonym).
</paragraph>
<paragraph blockId="26.[189,1399,943,1221]" box="[189,1271,943,970]" pageId="26" pageNumber="27">
<treatmentCitationGroup box="[189,1271,943,970]" pageId="26" pageNumber="27">
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[189,401,943,969]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[189,401,943,969]" italics="true" pageId="26" pageNumber="27">Ennea fischeriana</emphasis>
<paragraph id="CD07550BD83EFFDDFF19FC30F6A4FC2D" blockId="26.[189,1399,943,1221]" box="[189,1271,943,970]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="EDA87225D83EFFDDFF19FC30F6A4FC2D" box="[189,1271,943,970]" pageId="26" pageNumber="27">
<taxonomicName id="0AB82E88D83EFFDDFF19FC30F3C2FC2E" authorityName="Morelet" authorityYear="1877" box="[189,401,943,969]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D83EFFDDFF19FC30F3C2FC2E" box="[189,401,943,969]" italics="true" pageId="26" pageNumber="27">Ennea fischeriana</emphasis>
</taxonomicName>
–
<treatmentCitation author="Morelet A." box="[431,706,943,969]" page="230" pageId="26" pageNumber="27" year="1881">
<bibRefCitation author="Morelet A." box="[431,706,943,969]" pageId="26" pageNumber="27" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet 1881: 230-231</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFE0BFC2FF091FC2E" author="Morelet A." box="[431,706,943,969]" page="230" pageId="26" pageNumber="27" year="1881">
<bibRefCitation id="A92928FAD83EFFDDFE0BFC2FF091FC2E" author="Morelet A." box="[431,706,943,969]" pageId="26" pageNumber="27" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet 1881: 230-231</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Richardson C. L." box="[754,1013,943,969]" page="103" pageId="26" pageNumber="27" year="1988">
<bibRefCitation author="Richardson C. L." box="[754,1013,943,969]" pageId="26" pageNumber="27" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">Richardson 1988: 103</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFD56FC2FF1A6FC2E" author="Richardson C. L." box="[754,1013,943,969]" page="103" pageId="26" pageNumber="27" year="1988">
<bibRefCitation id="A92928FAD83EFFDDFD56FC2FF1A6FC2E" author="Richardson C. L." box="[754,1013,943,969]" pageId="26" pageNumber="27" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">Richardson 1988: 103</bibRefCitation>
</treatmentCitation>
(objective synonym).
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="26.[189,1399,943,1221]" box="[189,920,979,1006]" pageId="26" pageNumber="27">
<treatmentCitationGroup box="[189,920,979,1006]" pageId="26" pageNumber="27">
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[189,516,979,1006]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana" subGenus="Gulella">
<emphasis box="[189,264,980,1005]" italics="true" pageId="26" pageNumber="27">Ennea</emphasis>
<paragraph id="CD07550BD83EFFDDFF19FC54F1CBFC0A" blockId="26.[189,1399,943,1221]" box="[189,920,979,1006]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="EDA87225D83EFFDDFF19FC54F1CBFC0A" box="[189,920,979,1006]" pageId="26" pageNumber="27">
<taxonomicName id="0AB82E88D83EFFDDFF19FC54F057FC0A" authorityName="Morelet" authorityYear="1877" box="[189,516,979,1006]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana" subGenus="Gulella">
<emphasis id="FFCC8919D83EFFDDFF19FC54F35BFC0A" box="[189,264,980,1005]" italics="true" pageId="26" pageNumber="27">Ennea</emphasis>
(
<emphasis box="[281,369,979,1005]" italics="true" pageId="26" pageNumber="27">Gulella</emphasis>
<emphasis id="FFCC8919D83EFFDDFEBDFC53F322FC0A" box="[281,369,979,1005]" italics="true" pageId="26" pageNumber="27">Gulella</emphasis>
)
<emphasis box="[386,516,979,1005]" italics="true" pageId="26" pageNumber="27">fischeriana</emphasis>
<emphasis id="FFCC8919D83EFFDDFE26FC53F057FC0A" box="[386,516,979,1005]" italics="true" pageId="26" pageNumber="27">fischeriana</emphasis>
</taxonomicName>
–
<treatmentCitation author="Tryon G. W." box="[545,744,979,1006]" page="100" pageId="26" pageNumber="27" year="1885">
<bibRefCitation author="Tryon G. W." box="[545,744,979,1006]" pageId="26" pageNumber="27" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon 1885: 100</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFD85FC54F0BBFC0A" author="Tryon G. W." box="[545,744,979,1006]" page="100" pageId="26" pageNumber="27" year="1885">
<bibRefCitation id="A92928FAD83EFFDDFD85FC54F0BBFC0A" author="Tryon G. W." box="[545,744,979,1006]" pageId="26" pageNumber="27" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon 1885: 100</bibRefCitation>
</treatmentCitation>
, pl. 20, fig. 39.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="26.[189,1399,943,1221]" box="[189,1112,1015,1042]" pageId="26" pageNumber="27">
<treatmentCitationGroup box="[189,1112,1015,1042]" pageId="26" pageNumber="27">
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[189,565,1015,1042]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana" subGenus="Uniplicaria">
<emphasis box="[189,264,1016,1041]" italics="true" pageId="26" pageNumber="27">Ennea</emphasis>
<paragraph id="CD07550BD83EFFDDFF19FC78F60BFBF5" blockId="26.[189,1399,943,1221]" box="[189,1112,1015,1042]" pageId="26" pageNumber="27">
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<emphasis id="FFCC8919D83EFFDDFF19FC78F35BFBF6" box="[189,264,1016,1041]" italics="true" pageId="26" pageNumber="27">Ennea</emphasis>
(
<emphasis box="[281,418,1015,1041]" italics="true" pageId="26" pageNumber="27">Uniplicaria</emphasis>
<emphasis id="FFCC8919D83EFFDDFEBDFC77F3F1FBF6" box="[281,418,1015,1041]" italics="true" pageId="26" pageNumber="27">Uniplicaria</emphasis>
)
<emphasis box="[435,565,1015,1041]" italics="true" pageId="26" pageNumber="27">fischeriana</emphasis>
<emphasis id="FFCC8919D83EFFDDFE17FC77F066FBF6" box="[435,565,1015,1041]" italics="true" pageId="26" pageNumber="27">fischeriana</emphasis>
</taxonomicName>
–
<treatmentCitation author="Kobelt W." box="[595,803,1015,1042]" page="166" pageId="26" pageNumber="27" year="1905">
<bibRefCitation author="Kobelt W." box="[595,803,1015,1042]" pageId="26" pageNumber="27" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt 1905: 166</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFDF7FC77F170FBF6" author="Kobelt W." box="[595,803,1015,1042]" page="166" pageId="26" pageNumber="27" year="1905">
<bibRefCitation id="A92928FAD83EFFDDFDF7FC77F170FBF6" author="Kobelt W." box="[595,803,1015,1042]" pageId="26" pageNumber="27" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt 1905: 166</bibRefCitation>
</treatmentCitation>
, pl. 22, fig. 10; 1910: 158.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="26.[189,1399,943,1221]" box="[189,1082,1051,1078]" pageId="26" pageNumber="27">
<treatmentCitationGroup box="[189,1082,1051,1078]" pageId="26" pageNumber="27">
<taxonomicName ID-CoL="33XQL" authorityName="Morelet" authorityYear="1877" box="[189,379,1051,1077]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[189,379,1051,1077]" italics="true" pageId="26" pageNumber="27">Pupa minuscula</emphasis>
<paragraph id="CD07550BD83EFFDDFF19FB9CF669FBD2" blockId="26.[189,1399,943,1221]" box="[189,1082,1051,1078]" pageId="26" pageNumber="27">
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<emphasis id="FFCC8919D83EFFDDFF19FB9CF328FBD2" box="[189,379,1051,1077]" italics="true" pageId="26" pageNumber="27">Pupa minuscula</emphasis>
</taxonomicName>
–
<treatmentCitation author="Kobelt W." box="[408,616,1051,1078]" page="166" pageId="26" pageNumber="27" year="1905">
<bibRefCitation author="Kobelt W." box="[408,616,1051,1078]" pageId="26" pageNumber="27" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt 1905: 166</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFE3CFB9BF03BFBD2" author="Kobelt W." box="[408,616,1051,1078]" page="166" pageId="26" pageNumber="27" year="1905">
<bibRefCitation id="A92928FAD83EFFDDFE3CFB9BF03BFBD2" author="Kobelt W." box="[408,616,1051,1078]" pageId="26" pageNumber="27" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt 1905: 166</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Kobelt W." box="[629,750,1051,1078]" page="158" pageId="26" pageNumber="27" year="1910">
<bibRefCitation author="Kobelt W." box="[629,750,1051,1078]" pageId="26" pageNumber="27" pagination="138 - 196" refId="ref26795" refString="Kobelt W. 1910. Katalog der lebenden schalentragenden Mollusken der Abteilung Agnatha. Jahrbucher des Nassauischen Vereins fur Naturkunde 63: 138 - 196." type="journal article" year="1910">1910: 158</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFDD1FB9BF0BDFBD1" author="Kobelt W." box="[629,750,1051,1078]" page="158" pageId="26" pageNumber="27" year="1910">
<bibRefCitation id="A92928FAD83EFFDDFDD1FB9BF0BDFBD1" author="Kobelt W." box="[629,750,1051,1078]" pageId="26" pageNumber="27" pagination="138 - 196" refId="ref26795" refString="Kobelt W. 1910. Katalog der lebenden schalentragenden Mollusken der Abteilung Agnatha. Jahrbucher des Nassauischen Vereins fur Naturkunde 63: 138 - 196." type="journal article" year="1910">1910: 158</bibRefCitation>
</treatmentCitation>
. — van
<treatmentCitation author="Bruggen A. C. van" box="[848,1078,1051,1078]" page="166" pageId="26" pageNumber="27" year="1975">
<bibRefCitation author="Bruggen A. C. van" box="[848,1078,1051,1078]" pageId="26" pageNumber="27" pagination="157 - 175" refId="ref26057" refString="Bruggen A. C. van 1975. Streptaxidae (Mollusca, Gastropoda: Pulmonata) from Aldabra Island, Western Indian Ocean. Bulletin of the British Museum (Natural History) Zoology 28: 157 - 175." type="journal article" year="1975">Bruggen 1975: 166</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFCF4FB9BF665FBD2" author="Bruggen A. C. van" box="[848,1078,1051,1078]" page="166" pageId="26" pageNumber="27" year="1975">
<bibRefCitation id="A92928FAD83EFFDDFCF4FB9BF665FBD2" author="Bruggen A. C. van" box="[848,1078,1051,1078]" pageId="26" pageNumber="27" pagination="157 - 175" refId="ref26057" refString="Bruggen A. C. van 1975. Streptaxidae (Mollusca, Gastropoda: Pulmonata) from Aldabra Island, Western Indian Ocean. Bulletin of the British Museum (Natural History) Zoology 28: 157 - 175." type="journal article" year="1975">Bruggen 1975: 166</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="26.[189,1399,943,1221]" box="[189,642,1087,1114]" pageId="26" pageNumber="27">
<treatmentCitationGroup box="[189,642,1087,1114]" pageId="26" pageNumber="27">
<taxonomicName box="[189,392,1087,1113]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[189,392,1087,1113]" italics="true" pageId="26" pageNumber="27">Ennea minuscula</emphasis>
<paragraph id="CD07550BD83EFFDDFF19FBC0F0D1FBBD" blockId="26.[189,1399,943,1221]" box="[189,642,1087,1114]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="EDA87225D83EFFDDFF19FBC0F0D1FBBD" box="[189,642,1087,1114]" pageId="26" pageNumber="27">
<taxonomicName id="0AB82E88D83EFFDDFF19FBC0F3DBFBBE" box="[189,392,1087,1113]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="minuscula">
<emphasis id="FFCC8919D83EFFDDFF19FBC0F3DBFBBE" box="[189,392,1087,1113]" italics="true" pageId="26" pageNumber="27">Ennea minuscula</emphasis>
</taxonomicName>
–
<treatmentCitation author="Preston H. B." box="[421,639,1087,1114]" page="529" pageId="26" pageNumber="27" year="1910">
<bibRefCitation author="Preston H. B." box="[421,639,1087,1114]" pageId="26" pageNumber="27" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston 1910: 529</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFE01FBBFF02CFBBD" author="Preston H. B." box="[421,639,1087,1114]" page="529" pageId="26" pageNumber="27" year="1910">
<bibRefCitation id="A92928FAD83EFFDDFE01FBBFF02CFBBD" author="Preston H. B." box="[421,639,1087,1114]" pageId="26" pageNumber="27" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston 1910: 529</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="26.[189,1399,943,1221]" pageId="26" pageNumber="27">
<treatmentCitationGroup pageId="26" pageNumber="27">
<taxonomicName ID-CoL="6L5MK" authority="Morelet" authorityName="Morelet" authorityYear="1877" box="[189,516,1123,1149]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[189,411,1123,1149]" italics="true" pageId="26" pageNumber="27">Gulella minuscula</emphasis>
<paragraph id="CD07550BD83EFFDDFF19FBE3F011FB22" blockId="26.[189,1399,943,1221]" pageId="26" pageNumber="27">
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<emphasis id="FFCC8919D83EFFDDFF19FBE3F3C8FB9A" box="[189,411,1123,1149]" italics="true" pageId="26" pageNumber="27">Gulella minuscula</emphasis>
Morelet
</taxonomicName>
(
<emphasis box="[539,583,1124,1149]" italics="true" pageId="26" pageNumber="27">non</emphasis>
<taxonomicName authority="Emberton &amp; Pearce, 2000" authorityName="Emberton &amp; Pearce" authorityYear="2000" box="[595,1152,1123,1149]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[595,683,1123,1149]" italics="true" pageId="26" pageNumber="27">Gulella</emphasis>
<emphasis box="[696,817,1123,1149]" italics="true" pageId="26" pageNumber="27">minuscula</emphasis>
<bibRefCitation author="Emberton K. C. &amp; Pearce T. A." box="[829,1152,1123,1149]" pageId="26" pageNumber="27" pagination="126 - 153" refId="ref26524" refString="Emberton K. C. &amp; Pearce T. A. 2000. Small, High-Spired Pulmonates from Mounts Mahermana, Ilapiry, and Vasiha, Southeastern Madagascar, with Description of a New Genus, and with Conservation Statuses of 15 Streptaxid Species. The Veliger 43 (2): 126 - 153." type="journal article" year="2000">Emberton &amp; Pearce, 2000</bibRefCitation>
<emphasis id="FFCC8919D83EFFDDFDBFFBE4F014FB9A" box="[539,583,1124,1149]" italics="true" pageId="26" pageNumber="27">non</emphasis>
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<emphasis id="FFCC8919D83EFFDDFDF7FBE3F162FB9A" box="[595,817,1123,1149]" italics="true" pageId="26" pageNumber="27">Gulella minuscula</emphasis>
<bibRefCitation id="A92928FAD83EFFDDFC99FBE3F6D3FB9A" author="Emberton K. C. &amp; Pearce T. A." box="[829,1152,1123,1149]" pageId="26" pageNumber="27" pagination="126 - 153" refId="ref26524" refString="Emberton K. C. &amp; Pearce T. A. 2000. Small, High-Spired Pulmonates from Mounts Mahermana, Ilapiry, and Vasiha, Southeastern Madagascar, with Description of a New Genus, and with Conservation Statuses of 15 Streptaxid Species. The Veliger 43 (2): 126 - 153." type="journal article" year="2000">Emberton &amp; Pearce, 2000</bibRefCitation>
</taxonomicName>
) –
<treatmentCitation author="Fischer-Piette E. &amp; Vukadinovic D." page="55" pageId="26" pageNumber="27" year="1974">
<bibRefCitation author="Fischer-Piette E. &amp; Vukadinovic D." pageId="26" pageNumber="27" refId="ref26580" refString="Fischer-Piette E. &amp; Vukadinovic D. 1974. Les Mollusques Terrestres des Iles Comores. Memoires du Museum National d'Histoire Naturelle (Nouvelle Serie: Serie A, Zoologie) 84, Museum national d'Histoire naturelle, Paris." type="book" year="1974">Fischer-Piette &amp; Vukadinovic 1974: 55</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFB15FBE3F398FB46" author="Fischer-Piette E. &amp; Vukadinovic D." page="55" pageId="26" pageNumber="27" year="1974">
<bibRefCitation id="A92928FAD83EFFDDFB15FBE3F398FB46" author="Fischer-Piette E. &amp; Vukadinovic D." pageId="26" pageNumber="27" refId="ref26580" refString="Fischer-Piette E. &amp; Vukadinovic D. 1974. Les Mollusques Terrestres des Iles Comores. Memoires du Museum National d'Histoire Naturelle (Nouvelle Serie: Serie A, Zoologie) 84, Museum national d'Histoire naturelle, Paris." type="book" year="1974">Fischer-Piette &amp; Vukadinovic 1974: 55</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Richardson C. L." box="[512,780,1158,1185]" page="103" pageId="26" pageNumber="27" year="1988">
<bibRefCitation author="Richardson C. L." box="[512,780,1158,1185]" pageId="26" pageNumber="27" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">Richardson 1988: 103</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFDA4FB06F15FFB46" author="Richardson C. L." box="[512,780,1158,1185]" page="103" pageId="26" pageNumber="27" year="1988">
<bibRefCitation id="A92928FAD83EFFDDFDA4FB06F15FFB46" author="Richardson C. L." box="[512,780,1158,1185]" pageId="26" pageNumber="27" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">Richardson 1988: 103</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Rowson B." box="[834,1132,1159,1185]" page="441" pageId="26" pageNumber="27" year="2007">
<bibRefCitation author="Rowson B." box="[834,1132,1159,1185]" pageId="26" pageNumber="27" pagination="425 - 466" refId="ref27203" refString="Rowson B. 2007 a. Land molluscs of Zanzibar Island (Unguja), Tanzania, including a new species of Gulella (Pulmonata: Streptaxidae). Journal of Conchology 39 (4): 425 - 466." type="journal article" year="2007">Rowson 2007a: 441-442</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFCE6FB07F63FFB46" author="Rowson B." box="[834,1132,1159,1185]" page="441" pageId="26" pageNumber="27" year="2007">
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</treatmentCitation>
. —
<treatmentCitation author="Rowson B. &amp; Lange C. N." page="31" pageId="26" pageNumber="27" year="2007">
<bibRefCitation author="Rowson B. &amp; Lange C. N." pageId="26" pageNumber="27" pagination="21 - 32" refId="ref27291" refString="Rowson B. &amp; Lange C. N. 2007. Two new species of Gulella (Mollusca: Pulmonata: Streptaxidae) from the Taita Hills, Kenya. African Invertebrates 48 (2): 21 - 32." type="journal article" year="2007">Rowson &amp; Lange 2007: 31</bibRefCitation>
<treatmentCitation id="4C19731AD83EFFDDFB06FB07F374FB22" author="Rowson B. &amp; Lange C. N." page="31" pageId="26" pageNumber="27" year="2007">
<bibRefCitation id="A92928FAD83EFFDDFB06FB07F374FB22" author="Rowson B. &amp; Lange C. N." pageId="26" pageNumber="27" pagination="21 - 32" refId="ref27291" refString="Rowson B. &amp; Lange C. N. 2007. Two new species of Gulella (Mollusca: Pulmonata: Streptaxidae) from the Taita Hills, Kenya. African Invertebrates 48 (2): 21 - 32." type="journal article" year="2007">Rowson &amp; Lange 2007: 31</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation author="Muratov I. V." box="[344,574,1195,1221]" page="277" pageId="26" pageNumber="27" year="2010">
<bibRefCitation author="Muratov I. V." box="[344,574,1195,1221]" pageId="26" pageNumber="27" pagination="255 - 288" refId="ref26968" refString="Muratov I. V. 2010. Terrestrial molluscs of Cabo Delgado and adjacent inland areas of north-eastern Mozambique. African Invertebrates 51 (2): 255 - 288. http: // dx. doi. org / 10.5733 / afin. 051.0203" type="journal article" year="2010">Muratov 2010: 277</bibRefCitation>
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</treatmentCitation>
.
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</paragraph>
</subSubSection>
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<emphasis bold="true" box="[189,492,1260,1287]" pageId="26" pageNumber="27">
<typeStatus box="[189,250,1261,1287]" pageId="26" pageNumber="27">Type</typeStatus>
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<paragraph id="CD07550BD83EFFDDFF19FB6DF3BFFAE1" blockId="26.[189,492,1260,1287]" box="[189,492,1260,1287]" pageId="26" pageNumber="27">
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<typeStatus id="1203EBA9D83EFFDDFF19FB6DF2A9FAE0" box="[189,250,1261,1287]" pageId="26" pageNumber="27">Type</typeStatus>
material examined
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<paragraph blockId="26.[189,1399,1308,1478]" pageId="26" pageNumber="27">
<collectingCountry box="[189,338,1308,1334]" name="Comoros" pageId="26" pageNumber="27">COMOROS</collectingCountry>
<paragraph id="CD07550BD83EFFDDFF19FA9CF620FA21" blockId="26.[189,1399,1308,1478]" pageId="26" pageNumber="27">
<materialsCitation id="7DD05F56D83EFFDDFF19FA9CF0C2FAD0" box="[189,657,1308,1335]" collectionCode="NHMUK" country="Comoros" determinerName="Morelet" latitude="-12.22" location="COMOROS" longLatPrecision="776" longitude="44.45" pageId="26" pageNumber="27" specimenCode="1893.2.4.87-88" specimenCount="1" stateProvince="Anjouan I." typeStatus="lectotype">
<collectingCountry id="B5AF159BD83EFFDDFF19FA9CF301FAD1" box="[189,338,1308,1334]" name="Comoros" pageId="26" pageNumber="27">COMOROS</collectingCountry>
:
<typeStatus box="[347,453,1308,1334]" pageId="26" pageNumber="27">lectotype</typeStatus>
(here designated) and
<typeStatus box="[710,866,1308,1334]" pageId="26" pageNumber="27">paralectotype</typeStatus>
<collectionCode box="[872,988,1308,1334]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="26" pageNumber="27" type="Museum">NHMUK</collectionCode>
.1893.2.4.87-88: 2 ads, “
<collectingRegion box="[1260,1363,1308,1334]" country="Comoros" name="Anjouan" pageId="26" pageNumber="27">Anjouan</collectingRegion>
I.” (
<collectingRegion box="[197,289,1344,1370]" country="Comoros" name="Anjouan" pageId="26" pageNumber="27">Nzwani</collectingRegion>
I., approx.
<geoCoordinate box="[415,510,1344,1370]" degrees="12.22" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="555" value="-12.22">12.22°S</geoCoordinate>
<typeStatus id="1203EBA9D83EFFDDFEFFFA9CF396FAD1" box="[347,453,1308,1334]" pageId="26" pageNumber="27" type="lectotype">lectotype</typeStatus>
(here designated)
</materialsCitation>
<materialsCitation id="7DD05F56D83EFFDDFD33FA9CF3F1FA45" collectionCode="NHMUK" determinerName="Morelet" latitude="-12.22" location="Nzwani I." longLatPrecision="776" longitude="44.45" pageId="26" pageNumber="27" specimenCode="1893.2.4.87-88" specimenCount="2" stateProvince="Anjouan I." typeStatus="paralectotype">
and
<typeStatus id="1203EBA9D83EFFDDFD62FA9CF131FAD1" box="[710,866,1308,1334]" pageId="26" pageNumber="27" type="paralectotype">paralectotype</typeStatus>
<collectionCode id="ABA9CDCED83EFFDDFCCCFA9CF18FFAD1" box="[872,988,1308,1334]" collectionName="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="26" pageNumber="27" type="Museum">NHMUK</collectionCode>
.
<specimenCode id="9D1EFD70D83EFFDDFC7BFA9CF6C1FAD1" box="[991,1170,1308,1334]" pageId="26" pageNumber="27">1893.2.4.87-88</specimenCode>
:
<specimenCount id="DBBE9E82D83EFFDDFB3FFA9CF68BFAD1" box="[1179,1240,1308,1334]" pageId="26" pageNumber="27">2 ads</specimenCount>
, “
<collectingRegion id="0F7C9BE9D83EFFDDFB48FA9CF739FAD1" box="[1260,1386,1308,1334]" pageId="26" pageNumber="27">Anjouan I.</collectingRegion>
” (
<location id="C86703D0D83EFFDDFF61FAC0F36BFABD" LSID="urn:lsid:plazi:treatment:4511E41DD83EFFDCFE00FD63F6D1F80C:C86703D0D83EFFDDFF61FAC0F36BFABD" box="[197,312,1344,1370]" latitude="-12.22" longLatPrecision="776" longitude="44.45" name="Nzwani I." pageId="26" pageNumber="27" stateProvince="Anjouan I.">Nzwani I.</location>
, approx.
<geoCoordinate id="A88C33CCD83EFFDDFE3BFAC0F3ADFABD" box="[415,510,1344,1370]" degrees="12.22" direction="south" orientation="latitude" pageId="26" pageNumber="27" precision="555" value="-12.22">12.22°S</geoCoordinate>
,
<geoCoordinate box="[519,618,1343,1370]" degrees="44.45" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="555" value="44.45">44.45°E</geoCoordinate>
), labelled “
<typeStatus box="[743,811,1344,1370]" pageId="26" pageNumber="27">types</typeStatus>
”.
<collectingMunicipality box="[830,875,1344,1370]" pageId="26" pageNumber="27">The</collectingMunicipality>
smaller of the two
<specimenCount box="[1087,1208,1344,1370]" pageId="26" pageNumber="27" type="generic">specimens</specimenCount>
is apparently the shell figured in
<bibRefCitation author="Morelet A." box="[368,538,1379,1406]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">
<location LSID="urn:lsid:plazi:treatment:4511E41DD83EFFDCFE00FD63F6D1F80C:C86703D0D83EFFDDFED4FAE3F39FFA9A" box="[368,460,1379,1405]" country="Comoros" latitude="-12.22" longLatPrecision="776" longitude="44.45" municipality="The" name="Morelet" pageId="26" pageNumber="27" stateProvince="Anjouan">Morelet</location>
<geoCoordinate id="A88C33CCD83EFFDDFDA3FABFF039FABD" box="[519,618,1343,1370]" degrees="44.45" direction="east" orientation="longitude" pageId="26" pageNumber="27" precision="555" value="44.45">44.45°E</geoCoordinate>
), labelled “types”. The smaller of the two specimens is apparently the shell figured in
<bibRefCitation id="A92928FAD83EFFDDFED4FAE3F049FA99" author="Morelet A." box="[368,538,1379,1406]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">
<determinerName id="38DBB9CFD83EFFDDFED4FAE3F39FFA9A" box="[368,460,1379,1405]" pageId="26" pageNumber="27">Morelet</determinerName>
(1877
</bibRefCitation>
: pl.
<collectionCode box="[584,627,1380,1406]" pageId="26" pageNumber="27">XII</collectionCode>
, fig. 5) under the name “
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[916,1045,1379,1405]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[916,1045,1379,1405]" italics="true" pageId="26" pageNumber="27">fischeriana</emphasis>
: pl. XII, fig. 5) under the name “
<taxonomicName id="0AB82E88D83EFFDDFC30FAE3F646FA9A" authorityName="Morelet" authorityYear="1877" box="[916,1045,1379,1405]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D83EFFDDFC30FAE3F646FA9A" box="[916,1045,1379,1405]" italics="true" pageId="26" pageNumber="27">fischeriana</emphasis>
</taxonomicName>
”.
<location LSID="urn:lsid:plazi:treatment:4511E41DD83EFFDCFE00FD63F6D1F80C:C86703D0D83EFFDDFB8BFAE4F612FA99" box="[1071,1089,1380,1406]" country="Comoros" latitude="-12.22" longLatPrecision="776" longitude="44.45" municipality="The" name="It" pageId="26" pageNumber="27" stateProvince="Anjouan">It</location>
was evidently formerly on a white card mount.
<location LSID="urn:lsid:plazi:treatment:4511E41DD83EFFDCFE00FD63F6D1F80C:C86703D0D83EFFDDFE0CFA07F38FFA46" box="[424,476,1415,1441]" country="Comoros" latitude="-12.22" longLatPrecision="776" longitude="44.45" municipality="The" name="This" pageId="26" pageNumber="27" stateProvince="Anjouan">This</location>
”. It was evidently formerly on a white card mount.
</materialsCitation>
<location id="C86703D0D83EFFDDFE0CFA07F38FFA46" LSID="urn:lsid:plazi:treatment:4511E41DD83EFFDCFE00FD63F6D1F80C:C86703D0D83EFFDDFE0CFA07F38FFA46" box="[424,476,1415,1441]" country="Comoros" latitude="-12.22" longLatPrecision="776" longitude="44.45" municipality="The" name="This" pageId="26" pageNumber="27" stateProvince="Anjouan">This</location>
shell is here designated
<typeStatus box="[757,866,1415,1441]" pageId="26" pageNumber="27">lectotype</typeStatus>
<typeStatus id="1203EBA9D83EFFDDFD51FA07F131FA46" box="[757,866,1415,1441]" pageId="26" pageNumber="27" type="lectotype">lectotype</typeStatus>
, while the larger is considered a
<typeStatus box="[1242,1398,1415,1441]" pageId="26" pageNumber="27">paralectotype</typeStatus>
<typeStatus id="1203EBA9D83EFFDDFB7EFA07F725FA46" box="[1242,1398,1415,1441]" pageId="26" pageNumber="27" type="paralectotype">paralectotype</typeStatus>
(although Morelet’s description could in fact have been made from a single shell).
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="distribution">
<paragraph blockId="26.[189,349,1517,1544]" box="[189,349,1517,1544]" pageId="26" pageNumber="27">
<heading bold="true" box="[189,349,1517,1544]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis bold="true" box="[189,349,1517,1544]" pageId="26" pageNumber="27">
<typeStatus box="[189,250,1518,1544]" pageId="26" pageNumber="27">Type</typeStatus>
<paragraph id="CD07550BD83EFFDDFF19FA6EF30EF9E0" blockId="26.[189,349,1517,1544]" box="[189,349,1517,1544]" pageId="26" pageNumber="27">
<heading id="964FE267D83EFFDDFF19FA6EF30EF9E0" bold="true" box="[189,349,1517,1544]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="FFCC8919D83EFFDDFF19FA6EF30EF9E0" bold="true" box="[189,349,1517,1544]" pageId="26" pageNumber="27">
<typeStatus id="1203EBA9D83EFFDDFF19FA6EF2A9F9EF" box="[189,250,1518,1544]" pageId="26" pageNumber="27">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph blockId="26.[189,1009,1564,1591]" box="[189,1009,1564,1591]" pageId="26" pageNumber="27">
<paragraph id="CD07550BD83EFFDDFF19F99DF1A2F9D0" blockId="26.[189,1009,1564,1591]" box="[189,1009,1564,1591]" pageId="26" pageNumber="27">
“Dans l’île d’Anjouan (Johanna)” (
<bibRefCitation author="Morelet A." box="[601,765,1564,1591]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet 1877</bibRefCitation>
<bibRefCitation id="A92928FAD83EFFDDFDFDF99CF0AEF9D0" author="Morelet A." box="[601,765,1564,1591]" pageId="26" pageNumber="27" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet 1877</bibRefCitation>
; from title of paper).
</paragraph>
</subSubSection>
<subSubSection pageId="26" pageNumber="27" type="description">
<paragraph blockId="26.[189,334,1630,1656]" box="[189,334,1630,1656]" pageId="26" pageNumber="27">
<heading bold="true" box="[189,334,1630,1656]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis bold="true" box="[189,334,1630,1656]" pageId="26" pageNumber="27">Description</emphasis>
<subSubSection id="85A20680D83EFFDDFF19F9DEF361F80C" pageId="26" pageNumber="27" type="description">
<paragraph id="CD07550BD83EFFDDFF19F9DEF31DF99F" blockId="26.[189,334,1630,1656]" box="[189,334,1630,1656]" pageId="26" pageNumber="27">
<heading id="964FE267D83EFFDDFF19F9DEF31DF99F" bold="true" box="[189,334,1630,1656]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="FFCC8919D83EFFDDFF19F9DEF31DF99F" bold="true" box="[189,334,1630,1656]" pageId="26" pageNumber="27">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="26.[189,1399,1678,2027]" pageId="26" pageNumber="27">
<paragraph id="CD07550BD83EFFDDFF19F90EF361F80C" blockId="26.[189,1399,1678,2027]" pageId="26" pageNumber="27">
SHELL (
<figureCitation box="[277,407,1678,1704]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="26" pageNumber="27">Figs 38-39</figureCitation>
<figureCitation id="5583498ED83EFFDDFEB1F90EF3C4F94F" box="[277,407,1678,1704]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="26" pageNumber="27">Figs 38-39</figureCitation>
,
<figureCitation box="[421,494,1678,1705]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="26" pageNumber="27">62-63</figureCitation>
<figureCitation id="5583498ED83EFFDDFE01F90EF3BDF94E" box="[421,494,1678,1705]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="26" pageNumber="27">62-63</figureCitation>
). Variable in size, shape and dentition, small to large (3.00
<emphasis box="[1197,1207,1679,1704]" italics="true" pageId="26" pageNumber="27">-</emphasis>
<quantity box="[1207,1314,1678,1704]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="26" pageNumber="27" unit="mm" value="4.1">4.10 mm</quantity>
<emphasis id="FFCC8919D83EFFDDFB09F90FF6E4F94F" box="[1197,1207,1679,1704]" italics="true" pageId="26" pageNumber="27">-</emphasis>
<quantity id="0A40F8EED83EFFDDFB13F90EF771F94F" box="[1207,1314,1678,1704]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="26" pageNumber="27" unit="mm" value="4.1">4.10 mm</quantity>
high x 1.90
<emphasis box="[240,250,1715,1740]" italics="true" pageId="26" pageNumber="27">-</emphasis>
<emphasis id="FFCC8919D83EFFDDFF54F933F2A9F92B" box="[240,250,1715,1740]" italics="true" pageId="26" pageNumber="27">-</emphasis>
2.00 mm wide), of 6.5
<emphasis box="[509,519,1715,1740]" italics="true" pageId="26" pageNumber="27">-</emphasis>
<emphasis id="FFCC8919D83EFFDDFE59F933F054F92B" box="[509,519,1715,1740]" italics="true" pageId="26" pageNumber="27">-</emphasis>
7.0 whorls. Ovate-acuminate, spire broadly acuminate (spire angle 48
<emphasis box="[1330,1340,1715,1740]" italics="true" pageId="26" pageNumber="27">-</emphasis>
<emphasis id="FFCC8919D83EFFDDFA96F933F76FF92B" box="[1330,1340,1715,1740]" italics="true" pageId="26" pageNumber="27">-</emphasis>
63°). Apex pointed. Embryonic whorls smoothly granulate. Later whorls with relatively fine, numerous ribs (10
<emphasis box="[228,238,1786,1811]" italics="true" pageId="26" pageNumber="27">-</emphasis>
<emphasis id="FFCC8919D83EFFDDFF40F97AF2BDF8F4" box="[228,238,1786,1811]" italics="true" pageId="26" pageNumber="27">-</emphasis>
13 per mm on penultimate whorl). Sutures relatively deep. Umbilicus closed or nearly so. Peristome complete, or incomplete parietally. Outer palatal surface of aperture with a depression corresponding to the palatal tooth. Dentition 4-fold (
<typeStatus box="[651,763,1857,1883]" pageId="26" pageNumber="27">lectotype</typeStatus>
<typeStatus id="1203EBA9D83EFFDDFD2FF8C1F0A8F8BC" box="[651,763,1857,1883]" pageId="26" pageNumber="27" type="lectotype">lectotype</typeStatus>
,
<figureCitation box="[779,873,1857,1883]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="26" pageNumber="27">Figs 38</figureCitation>
<figureCitation id="5583498ED83EFFDDFCAFF8C1F13AF8BC" box="[779,873,1857,1883]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="26" pageNumber="27">Figs 38</figureCitation>
,
<figureCitation box="[889,920,1857,1884]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="26" pageNumber="27">62</figureCitation>
<figureCitation id="5583498ED83EFFDDFCDDF8C1F1CBF8BB" box="[889,920,1857,1884]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="26" pageNumber="27">62</figureCitation>
) to 5-fold (
<typeStatus box="[1064,1227,1857,1883]" pageId="26" pageNumber="27">paralectotype</typeStatus>
<typeStatus id="1203EBA9D83EFFDDFB8CF8C1F698F8BC" box="[1064,1227,1857,1883]" pageId="26" pageNumber="27" type="paralectotype">paralectotype</typeStatus>
,
<figureCitation box="[1242,1336,1857,1883]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="26" pageNumber="27">Figs 39</figureCitation>
<figureCitation id="5583498ED83EFFDDFB7EF8C1F76BF8BC" box="[1242,1336,1857,1883]" captionStart="Figs 36-40" captionStartId="24.[189,241,1860,1886]" captionTargetBox="[189,1399,1268,1819]" captionTargetId="figure@24.[189,1399,1268,1819]" captionTargetPageId="24" captionText="Figs 36-40. Adult shells of Dadagulella gen. nov. species. 36. D. selene (van Bruggen &amp; Van Goethem, 1999) comb. nov., holotype, Virunga NP (after van Bruggen &amp; Van Goethem 1999). 37. D. meredithae (van Bruggen, 2000), holotype, Nyika NP. 38. D. minuscula minuscula (Morelet, 1877), lectotype, Nzwani. 39. D. minuscula minuscula, paralectotype, Nzwani. 40. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806784" httpUri="https://zenodo.org/record/3806784/files/figure.png" pageId="26" pageNumber="27">Figs 39</figureCitation>
,
<figureCitation box="[1352,1384,1857,1884]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="26" pageNumber="27">63</figureCitation>
<figureCitation id="5583498ED83EFFDDFAECF8C1F73BF8BB" box="[1352,1384,1857,1884]" captionStart="Figs 51-64" captionStartId="35.[189,241,1719,1745]" captionTargetBox="[203,1389,740,1651]" captionTargetId="figure@35.[189,1398,725,1678]" captionTargetPageId="35" captionText="Figs 51-64. Apertures of adult shells of Dadagulella gen. nov. species (not to scale). 51. D. radius calva, lectotype, Taru Desert. 52. D. radius radius, lectotype, Shimba Hills. 53. D. radius radius, Amboni. 54. D. browni mafiensis subsp. nov., holotype, Mlula. 55. D. browni semulikiensis subsp. nov., holotype, Semuliki NP. 56. D. ecclesiola sp. nov., holotype, Kimboza (two views of shell showing basal tooth only visible when shell angled). 57. D. cresswelli sp. nov., holotype, Ngorongoro. 58. D. minareta sp. nov., holotype, Kimboza. 59. D. cuspidata comb. nov., holotype, Shume (note shell angled slightly differently to that in Fig. 31). 60. D. pembensis sp. nov., holotype, Ngezi. 61. D. frontierarum sp. nov., holotype, Mtai. 62. D. minuscula minuscula, lectotype, Nzwani. 63. D. minuscula minuscula, paralectotype, Nzwani. 64. D. minuscula mahorana subsp. nov., holotype, Mayotte." figureDoi="http://doi.org/10.5281/zenodo.3806796" httpUri="https://zenodo.org/record/3806796/files/figure.png" pageId="26" pageNumber="27">63</figureCitation>
), consisting of: one lamella-like parietal tooth; one slab-like palatal tooth, often bifid, but with parietopalatal sinus barely present; one basal denticle; and one deep-set columellar baffle, always visible. An additional shallow columellar denticle is present in the
<typeStatus box="[846,1008,1965,1991]" pageId="26" pageNumber="27">paralectotype</typeStatus>
<typeStatus id="1203EBA9D83EFFDDFCEAF82DF1A3F820" box="[846,1008,1965,1991]" pageId="26" pageNumber="27" type="paralectotype">paralectotype</typeStatus>
. Shells and anatomy of juveniles unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="multiple">
<paragraph blockId="27.[189,421,267,293]" box="[189,421,267,293]" pageId="27" pageNumber="28">
<heading bold="true" box="[189,421,267,293]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis bold="true" box="[189,421,267,293]" pageId="27" pageNumber="28">Range and habitat</emphasis>
<subSubSection id="85A20680D83FFFDCFF19FE8BF65DFEB3" pageId="27" pageNumber="28" type="biology_ecology">
<paragraph id="CD07550BD83FFFDCFF19FE8BF3F6FEC2" blockId="27.[189,421,267,293]" box="[189,421,267,293]" pageId="27" pageNumber="28">
<heading id="964FE267D83FFFDCFF19FE8BF3F6FEC2" bold="true" box="[189,421,267,293]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="FFCC8919D83FFFDCFF19FE8BF3F6FEC2" bold="true" box="[189,421,267,293]" pageId="27" pageNumber="28">Range and habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="27.[189,1038,314,341]" box="[189,1038,314,341]" pageId="27" pageNumber="28">
<collectingRegion box="[189,280,314,340]" country="Comoros" name="Anjouan" pageId="27" pageNumber="28">Nzwani</collectingRegion>
<paragraph id="CD07550BD83FFFDCFF19FEBAF65DFEB3" blockId="27.[189,1038,314,341]" box="[189,1038,314,341]" pageId="27" pageNumber="28">
<collectingRegion id="0F7C9BE9D83FFFDCFF19FEBAF34BFEB3" box="[189,280,314,340]" country="Comoros" name="Anjouan" pageId="27" pageNumber="28">Nzwani</collectingRegion>
(
<collectingRegion box="[295,400,314,341]" country="Comoros" name="Anjouan" pageId="27" pageNumber="28">Anjouan</collectingRegion>
<collectingRegion id="0F7C9BE9D83FFFDCFE83FEBAF3C3FEB2" box="[295,400,314,341]" country="Comoros" name="Anjouan" pageId="27" pageNumber="28">Anjouan</collectingRegion>
) island in the
<collectingCountry box="[566,674,314,340]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
<collectingCountry id="B5AF159BD83FFFDCFD92FEBAF0F1FEB3" box="[566,674,314,340]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
archipelago; habitat unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="discussion">
<paragraph blockId="27.[189,303,381,407]" box="[189,303,381,407]" pageId="27" pageNumber="28">
<heading bold="true" box="[189,303,381,407]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis bold="true" box="[189,303,381,407]" pageId="27" pageNumber="28">Remarks</emphasis>
<subSubSection id="85A20680D83FFFDCFF19FEFDF6D1F80C" pageId="27" pageNumber="28" type="discussion">
<paragraph id="CD07550BD83FFFDCFF19FEFDF37CFE70" blockId="27.[189,303,381,407]" box="[189,303,381,407]" pageId="27" pageNumber="28">
<heading id="964FE267D83FFFDCFF19FEFDF37CFE70" bold="true" box="[189,303,381,407]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="FFCC8919D83FFFDCFF19FEFDF37CFE70" bold="true" box="[189,303,381,407]" pageId="27" pageNumber="28">Remarks</emphasis>
</heading>
</paragraph>
<paragraph blockId="27.[189,1399,428,1499]" pageId="27" pageNumber="28">
<bibRefCitation author="Morelet A." box="[189,370,428,454]" pageId="27" pageNumber="28" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet (1877)</bibRefCitation>
<paragraph id="CD07550BD83FFFDCFF19FE2CF160FA3C" blockId="27.[189,1399,428,1499]" pageId="27" pageNumber="28">
<bibRefCitation id="A92928FAD83FFFDCFF19FE2CF321FE21" author="Morelet A." box="[189,370,428,454]" pageId="27" pageNumber="28" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet (1877)</bibRefCitation>
described this species from Anjouan (type locality implied by the paper’s title) under the name “
<emphasis box="[320,513,464,490]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFEE4FE51F052FE0D" box="[320,513,464,490]" italics="true" pageId="27" pageNumber="28">
Pupa
<taxonomicName authorityName="Emberton &amp; Pearce" authorityYear="2000" box="[392,513,464,490]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFE2CFE50F052FE0D" authorityName="Emberton &amp; Pearce" authorityYear="2000" box="[392,513,464,490]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
</emphasis>
” but figured it under the name “
<emphasis box="[908,1110,464,490]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFC28FE51F605FE0D" box="[908,1110,464,490]" italics="true" pageId="27" pageNumber="28">
Pupa
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[980,1110,464,490]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">fischeriana</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFC70FE50F605FE0D" authorityName="Morelet" authorityYear="1877" box="[980,1110,464,490]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">fischeriana</taxonomicName>
</emphasis>
” (pl. XII, fig. 5). In the paper he compared it to three species from “Île Bourbon” (Réunion), but none is very like it, suggesting Morelet was unaware of other
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[541,688,536,562]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[541,688,536,562]" italics="true" pageId="27" pageNumber="28">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFDB9FD98F0E3FDD5" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[541,688,536,562]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83FFFDCFDB9FD98F0E3FDD5" box="[541,688,536,562]" italics="true" pageId="27" pageNumber="28">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[693,798,536,562]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFD11FD98F14DFDD5" box="[693,798,536,562]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
species and probably meaning that
<emphasis box="[1209,1399,536,562]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFB1DFD99F724FDD5" box="[1209,1399,536,562]" italics="true" pageId="27" pageNumber="28">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[1278,1399,536,562]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFB5AFD98F724FDD5" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[1278,1399,536,562]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[189,317,572,598]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFF19FDBCF36EFDB1" box="[189,317,572,598]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
was the first species of the group to be collected. Two of the Réunion species are streptaxids of the Mascarene genus
<taxonomicName authority="Swainson, 1840" authorityName="Swainson" authorityYear="1840" box="[442,763,608,634]" class="Gastropoda" family="Streptaxidae" genus="Gonospira" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[442,567,608,634]" italics="true" pageId="27" pageNumber="28">Gonospira</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFE1EFDE0F0A8FD9D" authority="Swainson, 1840" authorityName="Swainson" authorityYear="1840" box="[442,763,608,634]" class="Gastropoda" family="Streptaxidae" genus="Gonospira" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis id="FFCC8919D83FFFDCFE1EFDE0F064FD9D" box="[442,567,608,634]" italics="true" pageId="27" pageNumber="28">Gonospira</emphasis>
Swainson, 1840
</taxonomicName>
while the third belongs to the
<taxonomicName authority="Turton, 1831" authorityName="Turton" authorityYear="1831" box="[1121,1399,608,634]" class="Gastropoda" family="Pupillidae" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="family">Pupillidae Turton, 1831</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFBC5FDE0F724FD9D" authority="Turton, 1831" authorityName="Turton" authorityYear="1831" box="[1121,1399,608,634]" class="Gastropoda" family="Pupillidae" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="family">Pupillidae Turton, 1831</taxonomicName>
(see
<bibRefCitation author="Griffiths O. L. &amp; Florens V. F. B." box="[246,549,644,671]" pageId="27" pageNumber="28" refId="ref26620" refString="Griffiths O. L. &amp; Florens V. F. B. 2006. A Field Guide to the Non-Marine Molluscs of the Mascarene Islands (Mauritius, Rodrigues and Reunion) and the Northern Dependencies of Mauritius. Bioculture Press, Mauritius." type="book" year="2006">Griffiths &amp; Florens 2006</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFF52FD04F076FD78" author="Griffiths O. L. &amp; Florens V. F. B." box="[246,549,644,671]" pageId="27" pageNumber="28" refId="ref26620" refString="Griffiths O. L. &amp; Florens V. F. B. 2006. A Field Guide to the Non-Marine Molluscs of the Mascarene Islands (Mauritius, Rodrigues and Reunion) and the Northern Dependencies of Mauritius. Bioculture Press, Mauritius." type="book" year="2006">Griffiths &amp; Florens 2006</bibRefCitation>
). Although
<bibRefCitation author="Kobelt W." box="[689,849,644,671]" pageId="27" pageNumber="28" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt (1905</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFD15FD04F102FD78" author="Kobelt W." box="[689,849,644,671]" pageId="27" pageNumber="28" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt (1905</bibRefCitation>
,
<bibRefCitation author="Kobelt W." box="[864,924,644,670]" pageId="27" pageNumber="28" pagination="138 - 196" refId="ref26795" refString="Kobelt W. 1910. Katalog der lebenden schalentragenden Mollusken der Abteilung Agnatha. Jahrbucher des Nassauischen Vereins fur Naturkunde 63: 138 - 196." type="journal article" year="1910">1910</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFCC4FD04F1CFFD79" author="Kobelt W." box="[864,924,644,670]" pageId="27" pageNumber="28" pagination="138 - 196" refId="ref26795" refString="Kobelt W. 1910. Katalog der lebenden schalentragenden Mollusken der Abteilung Agnatha. Jahrbucher des Nassauischen Vereins fur Naturkunde 63: 138 - 196." type="journal article" year="1910">1910</bibRefCitation>
) classified
<emphasis box="[1061,1258,644,670]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFB81FD05F6B9FD79" box="[1061,1258,644,670]" italics="true" pageId="27" pageNumber="28">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[1137,1258,644,670]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFBD5FD04F6B9FD79" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[1137,1258,644,670]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[1267,1399,644,670]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFB57FD04F724FD79" box="[1267,1399,644,670]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
(under
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[274,404,680,706]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[274,404,680,706]" italics="true" pageId="27" pageNumber="28">fischeriana</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFEB6FD28F3C7FD25" authorityName="Morelet" authorityYear="1877" box="[274,404,680,706]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D83FFFDCFEB6FD28F3C7FD25" box="[274,404,680,706]" italics="true" pageId="27" pageNumber="28">fischeriana</emphasis>
</taxonomicName>
) in the subgenus
<taxonomicName authority="L. Pfeiffer, 1856" authorityName="L. Pfeiffer" authorityYear="1856" box="[613,1058,680,707]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="subGenus" subGenus="Uniplicaria">
<emphasis box="[613,688,681,706]" italics="true" pageId="27" pageNumber="28">Ennea</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFDC1FD29F671FD24" authority="L. Pfeiffer, 1856" authorityName="L. Pfeiffer" authorityYear="1856" box="[613,1058,680,707]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="subGenus" subGenus="Uniplicaria">
<emphasis id="FFCC8919D83FFFDCFDC1FD29F0E3FD25" box="[613,688,681,706]" italics="true" pageId="27" pageNumber="28">Ennea</emphasis>
(
<emphasis box="[706,843,680,706]" italics="true" pageId="27" pageNumber="28">Uniplicaria</emphasis>
<emphasis id="FFCC8919D83FFFDCFD66FD28F118FD25" box="[706,843,680,706]" italics="true" pageId="27" pageNumber="28">Uniplicaria</emphasis>
) L. Pfeiffer, 1856
</taxonomicName>
, it is very unlike
<emphasis box="[1264,1326,681,706]" italics="true" pageId="27" pageNumber="28">Pupa</emphasis>
<taxonomicName authority="Dunker, 1848" authorityName="Dunker" authorityYear="1848" pageId="27" pageNumber="28" rank="species" species="cerea">
<emphasis box="[1334,1398,681,706]" italics="true" pageId="27" pageNumber="28">cerea</emphasis>
<emphasis id="FFCC8919D83FFFDCFB54FD29F77DFD25" box="[1264,1326,681,706]" italics="true" pageId="27" pageNumber="28">Pupa</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFA92FD29F333FD01" authority="Dunker, 1848" authorityName="Dunker" authorityYear="1848" pageId="27" pageNumber="28" rank="species" species="cerea">
<emphasis id="FFCC8919D83FFFDCFA92FD29F725FD25" box="[1334,1398,681,706]" italics="true" pageId="27" pageNumber="28">cerea</emphasis>
Dunker, 1848
</taxonomicName>
, the type species of
<emphasis box="[592,729,716,742]" italics="true" pageId="27" pageNumber="28">Uniplicaria</emphasis>
<emphasis id="FFCC8919D83FFFDCFDF4FD4CF08AFD01" box="[592,729,716,742]" italics="true" pageId="27" pageNumber="28">Uniplicaria</emphasis>
.
<bibRefCitation author="Preston H. B." box="[744,918,716,742]" pageId="27" pageNumber="28" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston (1910)</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFD4CFD4CF1C5FD01" author="Preston H. B." box="[744,918,716,742]" pageId="27" pageNumber="28" pagination="526 - 536" refId="ref27068" refString="Preston H. B. 1910. Additions to the non-Marine Molluscan Fauna of British and German East Africa and Lake Albert Edward. Annals and Magazine of Natural History, Series 8 6: 526 - 536. http: // dx. doi. org / 10.1080 / 00222931008692885" type="journal article" year="1910">Preston (1910)</bibRefCitation>
made a more relevant comparison in his description of
<taxonomicName authorityName="Preston" authorityYear="1910" box="[360,519,752,778]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="radius">
<emphasis box="[360,519,752,778]" italics="true" pageId="27" pageNumber="28">Ennea radius</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFECCFD71F054FCED" authorityName="Preston" authorityYear="1910" box="[360,519,752,778]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="radius">
<emphasis id="FFCC8919D83FFFDCFECCFD71F054FCED" box="[360,519,752,778]" italics="true" pageId="27" pageNumber="28">Ennea radius</emphasis>
</taxonomicName>
, noting that his species had coarser ribs, a columellar denticle, and a bifid palatal tooth that were absent in
<emphasis box="[590,789,788,814]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFDEAFC95F146FCC9" box="[590,789,788,814]" italics="true" pageId="27" pageNumber="28">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[668,789,788,814]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFD38FC94F146FCC9" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[668,789,788,814]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[799,932,788,814]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFCBBFC94F1F7FCC9" box="[799,932,788,814]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
Given these remarks and the fact that Preston cited only
<collectingRegion box="[411,512,824,850]" country="Comoros" name="Anjouan" pageId="27" pageNumber="28">Anjouan</collectingRegion>
<collectingRegion id="0F7C9BE9D83FFFDCFE3FFCB8F053FCB5" box="[411,512,824,850]" country="Comoros" name="Anjouan" pageId="27" pageNumber="28">Anjouan</collectingRegion>
as a locality for
<emphasis box="[716,911,824,850]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFD68FCB9F1DCFCB5" box="[716,911,824,850]" italics="true" pageId="27" pageNumber="28">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[790,911,824,850]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFCB2FCB8F1DCFCB5" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[790,911,824,850]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[920,1053,824,850]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFC3CFCB8F64EFCB5" box="[920,1053,824,850]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
, it thus appears he examined only the
<typeStatus box="[288,396,860,886]" pageId="27" pageNumber="28">lectotype</typeStatus>
<typeStatus id="1203EBA9D83FFFDCFE84FCDCF3DFFC91" box="[288,396,860,886]" pageId="27" pageNumber="28" type="lectotype">lectotype</typeStatus>
of the latter or material that was very like it. We agree that these features distinguish the
<typeStatus box="[189,297,896,922]" pageId="27" pageNumber="28">lectotype</typeStatus>
<typeStatus id="1203EBA9D83FFFDCFF19FC00F37AFC7D" box="[189,297,896,922]" pageId="27" pageNumber="28" type="lectotype">lectotype</typeStatus>
of
<emphasis box="[333,523,896,922]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFEE9FC01F058FC7D" box="[333,523,896,922]" italics="true" pageId="27" pageNumber="28">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[402,523,896,922]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFE36FC00F058FC7D" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[402,523,896,922]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[529,658,896,922]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFDB5FC00F0C1FC7D" box="[529,658,896,922]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
from that of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[807,938,896,922]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis box="[807,938,896,922]" italics="true" pageId="27" pageNumber="28">D. r. radius</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFC83FC01F1F9FC7D" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[807,938,896,922]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis id="FFCC8919D83FFFDCFC83FC01F1F9FC7D" box="[807,938,896,922]" italics="true" pageId="27" pageNumber="28">D. r. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[945,1073,896,922]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFC15FC00F662FC7D" box="[945,1073,896,922]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
However, the
<typeStatus box="[1240,1398,896,922]" pageId="27" pageNumber="28">paralectotype</typeStatus>
<typeStatus id="1203EBA9D83FFFDCFB7CFC00F725FC7D" box="[1240,1398,896,922]" pageId="27" pageNumber="28" type="paralectotype">paralectotype</typeStatus>
(if it is such; see also below) of
<emphasis box="[585,783,932,958]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFDEDFC25F15CFC59" box="[585,783,932,958]" italics="true" pageId="27" pageNumber="28">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[662,783,932,958]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFD32FC24F15CFC59" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[662,783,932,958]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[793,927,932,958]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFCBDFC24F1CCFC59" box="[793,927,932,958]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
, while having similarly fine ribs, has a stronger shallow columellar denticle that resembles that of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[896,1033,968,994]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis box="[896,1033,968,994]" italics="true" pageId="27" pageNumber="28">D. r. radius</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFC24FC49F65AFC05" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[896,1033,968,994]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis id="FFCC8919D83FFFDCFC24FC49F65AFC05" box="[896,1033,968,994]" italics="true" pageId="27" pageNumber="28">D. r. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1042,1173,968,994]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFBB6FC48F6C6FC05" box="[1042,1173,968,994]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
Although we have seen no mainland specimen that exactly matches either of them, both specimens of
<emphasis box="[1201,1399,1004,1030]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFB15FC6DF724FBE1" box="[1201,1399,1004,1030]" italics="true" pageId="27" pageNumber="28">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[1278,1399,1004,1030]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFB5AFC6CF724FBE1" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[1278,1399,1004,1030]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[189,322,1040,1066]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFF19FB90F311FBCD" box="[189,322,1040,1066]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
fall within the range of variation seen in
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[838,978,1040,1066]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis box="[838,978,1040,1066]" italics="true" pageId="27" pageNumber="28">D. r. radius</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFCE2FB91F181FBCD" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Rowson &amp; Tattersfield" baseAuthorityYear="2013" box="[838,978,1040,1066]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="subSpecies" species="radius" status="comb. nov." subSpecies="radius">
<emphasis id="FFCC8919D83FFFDCFCE2FB91F181FBCD" box="[838,978,1040,1066]" italics="true" pageId="27" pageNumber="28">D. r. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[989,1122,1040,1066]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFC79FB90F631FBCD" box="[989,1122,1040,1066]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
in other respects. It is therefore possible that the two taxa are synonymous, in which case Morelet’s name would take priority over Preston’s. It is even possible that
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[639,796,1112,1138]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
<emphasis box="[639,796,1112,1138]" italics="true" pageId="27" pageNumber="28">
D.
<taxonomicName box="[675,796,1112,1138]" pageId="27" pageNumber="28" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFDDBFBD9F14FFB95" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[639,796,1112,1138]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
<emphasis id="FFCC8919D83FFFDCFDDBFBD9F14FFB95" box="[639,796,1112,1138]" italics="true" pageId="27" pageNumber="28">D. minuscula</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[803,932,1112,1139]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[939,974,1112,1138]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
<taxonomicNameLabel id="E4FF3462D83FFFDCFC87FBD8F1F7FB94" box="[803,932,1112,1139]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D83FFFDCFC0FFBD8F19DFB95" box="[939,974,1112,1138]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
consists of populations of
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1288,1398,1112,1138]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis box="[1288,1398,1112,1138]" italics="true" pageId="27" pageNumber="28">D. radius</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFAACFBD9F725FB95" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1288,1398,1112,1138]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis id="FFCC8919D83FFFDCFAACFBD9F725FB95" box="[1288,1398,1112,1138]" italics="true" pageId="27" pageNumber="28">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[189,320,1148,1175]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[328,363,1148,1174]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
<taxonomicNameLabel id="E4FF3462D83FFFDCFF19FBFCF313FB70" box="[189,320,1148,1175]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D83FFFDCFEECFBFCF338FB71" box="[328,363,1148,1174]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
derived from specimens introduced from the African mainland (the alternative, that
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis italics="true" pageId="27" pageNumber="28">D. radius</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFAFEFBFDF35BFB5D" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis id="FFCC8919D83FFFDCFAFEFBFDF35BFB5D" italics="true" pageId="27" pageNumber="28">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[274,406,1184,1211]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[416,451,1184,1210]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
<taxonomicNameLabel id="E4FF3462D83FFFDCFEB6FB20F3C5FB5C" box="[274,406,1184,1211]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D83FFFDCFE04FB20F390FB5D" box="[416,451,1184,1210]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
was introduced into East Africa from the
<collectingCountry box="[964,1072,1184,1210]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
<collectingCountry id="B5AF159BD83FFFDCFC60FB20F663FB5D" box="[964,1072,1184,1210]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
appears unlikely given the variation seen in East African populations and the many similar species there). That said, the isolated, volcanic
<collectingCountry box="[297,405,1256,1282]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
<collectingCountry id="B5AF159BD83FFFDCFE8DFB68F3C6FAE5" box="[297,405,1256,1282]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
archipelago comprises a region of endemism in itself and
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[1102,1260,1256,1282]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
<emphasis box="[1102,1260,1256,1282]" italics="true" pageId="27" pageNumber="28">
D.
<taxonomicName box="[1139,1260,1256,1282]" pageId="27" pageNumber="28" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFBEAFB69F6BFFAE5" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[1102,1260,1256,1282]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">
<emphasis id="FFCC8919D83FFFDCFBEAFB69F6BFFAE5" box="[1102,1260,1256,1282]" italics="true" pageId="27" pageNumber="28">D. minuscula</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1268,1399,1256,1283]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[189,224,1292,1318]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
<taxonomicNameLabel id="E4FF3462D83FFFDCFB50FB68F724FAE4" box="[1268,1399,1256,1283]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D83FFFDCFF19FA8CF2B3FAC1" box="[189,224,1292,1318]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
could easily be a
<collectingCountry box="[442,550,1292,1318]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
<collectingCountry id="B5AF159BD83FFFDCFE1EFA8CF075FAC1" box="[442,550,1292,1318]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
endemic whose overlapping variation with
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1074,1187,1292,1318]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis box="[1074,1187,1292,1318]" italics="true" pageId="27" pageNumber="28">D. radius</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFB96FA8DF6F0FAC1" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Preston" baseAuthorityYear="1910" box="[1074,1187,1292,1318]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="radius" status="comb. nov.">
<emphasis id="FFCC8919D83FFFDCFB96FA8DF6F0FAC1" box="[1074,1187,1292,1318]" italics="true" pageId="27" pageNumber="28">D. radius</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1196,1327,1292,1319]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis box="[1336,1370,1292,1318]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
<taxonomicNameLabel id="E4FF3462D83FFFDCFB08FA8CF77CFAC0" box="[1196,1327,1292,1319]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<emphasis id="FFCC8919D83FFFDCFA9CFA8CF709FAC1" box="[1336,1370,1292,1318]" italics="true" pageId="27" pageNumber="28">s.l.</emphasis>
is due to homoplasy. Resolving the relationships between these two taxa, and
<emphasis box="[1070,1261,1328,1354]" italics="true" pageId="27" pageNumber="28">D. m. mahorana</emphasis>
<emphasis id="FFCC8919D83FFFDCFB8AFAB1F6BEFAAD" box="[1070,1261,1328,1354]" italics="true" pageId="27" pageNumber="28">D. m. mahorana</emphasis>
subsp. nov. below, is likely to require anatomical or molecular data. Until such information becomes available, and while we focus on other issues within
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[643,790,1400,1426]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[643,790,1400,1426]" italics="true" pageId="27" pageNumber="28">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFD27FAF8F145FA75" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[643,790,1400,1426]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83FFFDCFD27FAF8F145FA75" box="[643,790,1400,1426]" italics="true" pageId="27" pageNumber="28">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[798,908,1401,1427]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFCBAFAF9F1DFFA74" box="[798,908,1401,1427]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
, we maintain the two species as separate. Van
<bibRefCitation author="Bruggen A. C. van" box="[242,428,1436,1463]" pageId="27" pageNumber="28" pagination="71 - 77" refId="ref26097" refString="Bruggen A. C. van 1986. Additional notes on Gulella (Gastropoda Pulmonata: Streptaxidae) from the Aldabra group, western Indian Ocean. Basteria 50: 71 - 77." type="journal article" year="1986">Bruggen (1986)</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFF56FA1DF3FFFA50" author="Bruggen A. C. van" box="[242,428,1436,1463]" pageId="27" pageNumber="28" pagination="71 - 77" refId="ref26097" refString="Bruggen A. C. van 1986. Additional notes on Gulella (Gastropoda Pulmonata: Streptaxidae) from the Aldabra group, western Indian Ocean. Basteria 50: 71 - 77." type="journal article" year="1986">Bruggen (1986)</bibRefCitation>
took a similar approach with two other streptaxids, one described by Preston from the Shimba Hills, the other by Morelet from Mayotte.
</paragraph>
<paragraph blockId="27.[189,1399,1538,1888]" pageId="27" pageNumber="28">
<paragraph id="CD07550BD83FFFDCFF19F982F1D7F887" blockId="27.[189,1399,1538,1888]" pageId="27" pageNumber="28">
There is a nomenclatural issue concerning the name
<taxonomicName authority="Morelet, 1877" authorityName="Morelet" authorityYear="1877" box="[813,1113,1538,1564]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[813,934,1538,1564]" italics="true" pageId="27" pageNumber="28">minuscula</emphasis>
<bibRefCitation author="Morelet A." box="[942,1113,1538,1564]" pageId="27" pageNumber="28" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877</bibRefCitation>
<taxonomicName id="0AB82E88D83FFFDCFC89F982F60AF9FB" authority="Morelet, 1877" authorityName="Morelet" authorityYear="1877" box="[813,1113,1538,1564]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis id="FFCC8919D83FFFDCFC89F982F1F5F9FB" box="[813,934,1538,1564]" italics="true" pageId="27" pageNumber="28">minuscula</emphasis>
<bibRefCitation id="A92928FAD83FFFDCFC0AF982F60AF9FB" author="Morelet A." box="[942,1113,1538,1564]" pageId="27" pageNumber="28" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">Morelet, 1877</bibRefCitation>
</taxonomicName>
.
<bibRefCitation author="Morelet A." box="[1126,1305,1538,1564]" pageId="27" pageNumber="28" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet (1881)</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFBC2F982F74AF9FB" author="Morelet A." box="[1126,1305,1538,1564]" pageId="27" pageNumber="28" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet (1881)</bibRefCitation>
tried to replace the name used for the (1877) description,
<emphasis box="[784,976,1574,1600]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFCB4F9A7F183F9A7" box="[784,976,1574,1600]" italics="true" pageId="27" pageNumber="28">
Pupa
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[855,976,1574,1600]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFCF3F9A6F183F9A7" authorityName="Morelet" authorityYear="1877" box="[855,976,1574,1600]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">minuscula</taxonomicName>
</emphasis>
, with the name used for the (1877) figure,
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[271,483,1610,1636]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[271,483,1610,1636]" italics="true" pageId="27" pageNumber="28">Ennea fischeriana</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFEABF9CBF3B0F983" authorityName="Morelet" authorityYear="1877" box="[271,483,1610,1636]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D83FFFDCFEABF9CBF3B0F983" box="[271,483,1610,1636]" italics="true" pageId="27" pageNumber="28">Ennea fischeriana</emphasis>
</taxonomicName>
.
<bibRefCitation author="Morelet A." box="[497,675,1610,1636]" pageId="27" pageNumber="28" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet (1881)</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFE55F9CAF0F0F983" author="Morelet A." box="[497,675,1610,1636]" pageId="27" pageNumber="28" pagination="212 - 240" refId="ref26900" refString="Morelet A. 1881. Malacologie des Comores. Recolte de M. Marie a l'ile Mayotte. Journal de Conchyliologie 29: 212 - 240." type="journal article" year="1881">Morelet (1881)</bibRefCitation>
wrote that the name
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[919,1068,1610,1636]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[919,1068,1610,1636]" italics="true" pageId="27" pageNumber="28">P. minuscula</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFC33F9CBF67FF983" authorityName="Morelet" authorityYear="1877" box="[919,1068,1610,1636]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis id="FFCC8919D83FFFDCFC33F9CBF67FF983" box="[919,1068,1610,1636]" italics="true" pageId="27" pageNumber="28">P. minuscula</emphasis>
</taxonomicName>
had been published in error. Subsequently both
<bibRefCitation author="Tryon G. W." box="[418,637,1646,1673]" pageId="27" pageNumber="28" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon (1885: 100)</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFE06F9EFF02EF96F" author="Tryon G. W." box="[418,637,1646,1673]" pageId="27" pageNumber="28" refId="ref27788" refString="Tryon G. W. 1885. Testacellidae, Oleacinidae, Streptaxidae, Helicoidea, Vitrinidae, Limacidae, Arionidae. Manual of Conchology: Structrual and Systematic, with Illustrations of the Species. Second Series: Pulmonata 1, Tyron G. W., Philadelphia." type="journal volume" year="1885">Tryon (1885: 100)</bibRefCitation>
and
<bibRefCitation author="Kobelt W." box="[699,922,1646,1673]" pageId="27" pageNumber="28" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt (1905: 166</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFD1FF9EEF1C9F96F" author="Kobelt W." box="[699,922,1646,1673]" pageId="27" pageNumber="28" pagination="1 - 211" refId="ref26749" refString="Kobelt W. 1905. Die Raublungenschnecken (Agnatha). Zweite Abtheilung. Streptaxidae und Daudebardiidae. In: Martini &amp; Chemnitz (eds) Systematisches Conchylien-Cabinet I. 12 B (2): 1 - 211. Bauer und Raspe, Nurnberg." type="journal article" year="1905">Kobelt (1905: 166</bibRefCitation>
;
<bibRefCitation author="Kobelt W." box="[937,1058,1646,1673]" pageId="27" pageNumber="28" pagination="138 - 196" refId="ref26795" refString="Kobelt W. 1910. Katalog der lebenden schalentragenden Mollusken der Abteilung Agnatha. Jahrbucher des Nassauischen Vereins fur Naturkunde 63: 138 - 196." type="journal article" year="1910">1910: 158</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFC0DF9EEF671F96E" author="Kobelt W." box="[937,1058,1646,1673]" pageId="27" pageNumber="28" pagination="138 - 196" refId="ref26795" refString="Kobelt W. 1910. Katalog der lebenden schalentragenden Mollusken der Abteilung Agnatha. Jahrbucher des Nassauischen Vereins fur Naturkunde 63: 138 - 196." type="journal article" year="1910">1910: 158</bibRefCitation>
) accepted
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[1189,1319,1646,1672]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[1189,1319,1646,1672]" italics="true" pageId="27" pageNumber="28">fischeriana</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFB01F9EEF774F96F" authorityName="Morelet" authorityYear="1877" box="[1189,1319,1646,1672]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D83FFFDCFB01F9EEF774F96F" box="[1189,1319,1646,1672]" italics="true" pageId="27" pageNumber="28">fischeriana</emphasis>
</taxonomicName>
as the species name. However, when revising the fauna of the
<collectingCountry box="[842,953,1682,1708]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
<collectingCountry id="B5AF159BD83FFFDCFCEEF912F1EAF94B" box="[842,953,1682,1708]" name="Comoros" pageId="27" pageNumber="28">Comoros</collectingCountry>
,
<bibRefCitation author="Fischer-Piette E. &amp; Vukadinovic D." box="[964,1398,1682,1708]" pageId="27" pageNumber="28" refId="ref26580" refString="Fischer-Piette E. &amp; Vukadinovic D. 1974. Les Mollusques Terrestres des Iles Comores. Memoires du Museum National d'Histoire Naturelle (Nouvelle Serie: Serie A, Zoologie) 84, Museum national d'Histoire naturelle, Paris." type="book" year="1974">Fischer-Piette &amp; Vukadinovic (1974)</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFC60F912F725F94B" author="Fischer-Piette E. &amp; Vukadinovic D." box="[964,1398,1682,1708]" pageId="27" pageNumber="28" refId="ref26580" refString="Fischer-Piette E. &amp; Vukadinovic D. 1974. Les Mollusques Terrestres des Iles Comores. Memoires du Museum National d'Histoire Naturelle (Nouvelle Serie: Serie A, Zoologie) 84, Museum national d'Histoire naturelle, Paris." type="book" year="1974">Fischer-Piette &amp; Vukadinovic (1974)</bibRefCitation>
disagreed. They treated
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[472,635,1718,1744]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[472,635,1718,1744]" italics="true" pageId="27" pageNumber="28">
E.
<taxonomicName box="[505,635,1718,1744]" pageId="27" pageNumber="28" rank="species" species="fischeriana">fischeriana</taxonomicName>
</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFE7CF937F028F937" authorityName="Morelet" authorityYear="1877" box="[472,635,1718,1744]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D83FFFDCFE7CF937F028F937" box="[472,635,1718,1744]" italics="true" pageId="27" pageNumber="28">E. fischeriana</emphasis>
</taxonomicName>
is an objective synonym of
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[970,1120,1718,1744]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[970,1120,1718,1744]" italics="true" pageId="27" pageNumber="28">P. minuscula</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFC6EF937F633F937" authorityName="Morelet" authorityYear="1877" box="[970,1120,1718,1744]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis id="FFCC8919D83FFFDCFC6EF937F633F937" box="[970,1120,1718,1744]" italics="true" pageId="27" pageNumber="28">P. minuscula</emphasis>
</taxonomicName>
since the latter was not preoccupied and was validly introduced. This point of view was followed by
<bibRefCitation author="Richardson C. L." box="[1110,1334,1754,1781]" pageId="27" pageNumber="28" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">Richardson (1988)</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFBF2F95AF765F912" author="Richardson C. L." box="[1110,1334,1754,1781]" pageId="27" pageNumber="28" refId="ref27169" refString="Richardson C. L. 1988. Streptaxacea: catalog of species. Part I: Streptaxidae. Tryonia 16, Department of Malacology, Academy of Natural Sciences of Philadelphia, Philadelphia." type="journal volume" year="1988">Richardson (1988)</bibRefCitation>
. As a consequence, the
<typeStatus box="[400,508,1790,1816]" pageId="27" pageNumber="28">lectotype</typeStatus>
<typeStatus id="1203EBA9D83FFFDCFE34F97EF3AFF8FF" box="[400,508,1790,1816]" pageId="27" pageNumber="28" type="lectotype">lectotype</typeStatus>
of
<emphasis box="[550,745,1790,1816]" italics="true" pageId="27" pageNumber="28">
<emphasis id="FFCC8919D83FFFDCFD82F97FF0BAF8FF" box="[550,745,1790,1816]" italics="true" pageId="27" pageNumber="28">
D. m.
<taxonomicName authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[624,745,1790,1816]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFDD4F97EF0BAF8FF" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" baseAuthorityName="Morelet" baseAuthorityYear="1877" box="[624,745,1790,1816]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula" status="comb. nov.">minuscula</taxonomicName>
</emphasis>
<taxonomicNameLabel box="[754,885,1790,1817]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFD56F97EF126F8FE" box="[754,885,1790,1817]" pageId="27" pageNumber="28" rank="species">comb. nov.</taxonomicNameLabel>
in NHMUK also becomes the
<typeStatus box="[1258,1366,1790,1816]" pageId="27" pageNumber="28">lectotype</typeStatus>
<typeStatus id="1203EBA9D83FFFDCFB4EF97EF705F8FF" box="[1258,1366,1790,1816]" pageId="27" pageNumber="28" type="lectotype">lectotype</typeStatus>
of
<taxonomicName authorityName="Morelet" authorityYear="1877" box="[189,353,1826,1852]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis box="[189,353,1826,1852]" italics="true" pageId="27" pageNumber="28">
E.
<taxonomicName box="[223,353,1826,1852]" pageId="27" pageNumber="28" rank="species" species="fischeriana">fischeriana</taxonomicName>
</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFF19F8A3F332F8DB" authorityName="Morelet" authorityYear="1877" box="[189,353,1826,1852]" class="Gastropoda" family="Streptaxidae" genus="Ennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="fischeriana">
<emphasis id="FFCC8919D83FFFDCFF19F8A3F332F8DB" box="[189,353,1826,1852]" italics="true" pageId="27" pageNumber="28">E. fischeriana</emphasis>
</taxonomicName>
and the latter name remains unavailable for a taxon founded on another specimen. This includes the
<typeStatus box="[337,498,1862,1888]" pageId="27" pageNumber="28">paralectotype</typeStatus>
<typeStatus id="1203EBA9D83FFFDCFEF5F8C6F3A1F887" box="[337,498,1862,1888]" pageId="27" pageNumber="28" type="paralectotype">paralectotype</typeStatus>
, which is further discussed below.
</paragraph>
<paragraph blockId="27.[189,1399,1929,2027]" pageId="27" pageNumber="28">
<paragraph id="CD07550BD83FFFDCFF19F809F6D1F80C" blockId="27.[189,1399,1929,2027]" pageId="27" pageNumber="28">
Finally, we note that, although
<bibRefCitation author="Emberton K. C. &amp; Pearce T. A." box="[562,884,1929,1955]" pageId="27" pageNumber="28" pagination="126 - 153" refId="ref26524" refString="Emberton K. C. &amp; Pearce T. A. 2000. Small, High-Spired Pulmonates from Mounts Mahermana, Ilapiry, and Vasiha, Southeastern Madagascar, with Description of a New Genus, and with Conservation Statuses of 15 Streptaxid Species. The Veliger 43 (2): 126 - 153." type="journal article" year="2000">Emberton &amp; Pearce (2000)</bibRefCitation>
<bibRefCitation id="A92928FAD83FFFDCFD96F809F127F844" author="Emberton K. C. &amp; Pearce T. A." box="[562,884,1929,1955]" pageId="27" pageNumber="28" pagination="126 - 153" refId="ref26524" refString="Emberton K. C. &amp; Pearce T. A. 2000. Small, High-Spired Pulmonates from Mounts Mahermana, Ilapiry, and Vasiha, Southeastern Madagascar, with Description of a New Genus, and with Conservation Statuses of 15 Streptaxid Species. The Veliger 43 (2): 126 - 153." type="journal article" year="2000">Emberton &amp; Pearce (2000)</bibRefCitation>
described a junior homonym of
<taxonomicName authority="Morelet, 1877" authorityName="Morelet" authorityYear="1877" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[1278,1399,1929,1955]" italics="true" pageId="27" pageNumber="28">minuscula</emphasis>
<bibRefCitation author="Morelet A." box="[189,358,1965,1991]" pageId="27" pageNumber="28" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">
<taxonomicName id="0AB82E88D83FFFDCFB5AF809F335F820" authority="Morelet, 1877" authorityName="Morelet" authorityYear="1877" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis id="FFCC8919D83FFFDCFB5AF809F724F844" box="[1278,1399,1929,1955]" italics="true" pageId="27" pageNumber="28">minuscula</emphasis>
<bibRefCitation id="A92928FAD83FFFDCFF19F82DF335F820" author="Morelet A." box="[189,358,1965,1991]" pageId="27" pageNumber="28" pagination="325 - 347" refId="ref26877" refString="Morelet A. 1877. Excursion conchyliologique dans i'ile d'Anjouan (Johanna). Journal de Conchyliologie 25: 325 - 347." type="journal article" year="1877">
Morelet,
<quantity box="[299,358,1965,1991]" metricMagnitude="1" metricUnit="m" metricValue="4.76758" pageId="27" pageNumber="28" unit="in" value="1877.0">1877</quantity>
<quantity id="0A40F8EED83FFFDCFE8FF82DF335F820" box="[299,358,1965,1991]" metricMagnitude="1" metricUnit="m" metricValue="4.76758" pageId="27" pageNumber="28" unit="in" value="1877.0">1877</quantity>
</bibRefCitation>
</taxonomicName>
in
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1856" box="[400,488,1965,1991]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[400,488,1965,1991]" italics="true" pageId="27" pageNumber="28">Gulella</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFE34F82DF3BBF820" authorityName="L. Pfeiffer" authorityYear="1856" box="[400,488,1965,1991]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis id="FFCC8919D83FFFDCFE34F82DF3BBF820" box="[400,488,1965,1991]" italics="true" pageId="27" pageNumber="28">Gulella</emphasis>
</taxonomicName>
(
<taxonomicName authority="Emberton &amp; Pearce, 2000" authorityName="Emberton &amp; Pearce" authorityYear="2000" box="[507,988,1965,1991]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[507,535,1965,1991]" italics="true" pageId="27" pageNumber="28">G.</emphasis>
<taxonomicName authority="Emberton &amp; Pearce, 2000" authorityName="Emberton &amp; Pearce" authorityYear="2000" box="[545,988,1965,1991]" pageId="27" pageNumber="28" rank="species" species="minuscula">
<emphasis box="[545,666,1965,1991]" italics="true" pageId="27" pageNumber="28">minuscula</emphasis>
<bibRefCitation author="Emberton K. C. &amp; Pearce T. A." box="[675,988,1965,1991]" pageId="27" pageNumber="28" pagination="126 - 153" refId="ref26524" refString="Emberton K. C. &amp; Pearce T. A. 2000. Small, High-Spired Pulmonates from Mounts Mahermana, Ilapiry, and Vasiha, Southeastern Madagascar, with Description of a New Genus, and with Conservation Statuses of 15 Streptaxid Species. The Veliger 43 (2): 126 - 153." type="journal article" year="2000">Emberton &amp; Pearce, 2000</bibRefCitation>
</taxonomicName>
<taxonomicName id="0AB82E88D83FFFDCFE5FF82DF18FF820" authority="Emberton &amp; Pearce, 2000" authorityName="Emberton &amp; Pearce" authorityYear="2000" box="[507,988,1965,1991]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis id="FFCC8919D83FFFDCFE5FF82DF044F820" box="[507,535,1965,1991]" italics="true" pageId="27" pageNumber="28">G.</emphasis>
<emphasis id="FFCC8919D83FFFDCFD85F82DF0C9F820" box="[545,666,1965,1991]" italics="true" pageId="27" pageNumber="28">minuscula</emphasis>
<bibRefCitation id="A92928FAD83FFFDCFD07F82DF18FF820" author="Emberton K. C. &amp; Pearce T. A." box="[675,988,1965,1991]" pageId="27" pageNumber="28" pagination="126 - 153" refId="ref26524" refString="Emberton K. C. &amp; Pearce T. A. 2000. Small, High-Spired Pulmonates from Mounts Mahermana, Ilapiry, and Vasiha, Southeastern Madagascar, with Description of a New Genus, and with Conservation Statuses of 15 Streptaxid Species. The Veliger 43 (2): 126 - 153." type="journal article" year="2000">Emberton &amp; Pearce, 2000</bibRefCitation>
</taxonomicName>
) no replacement name is required since
<taxonomicName authority="Morelet" authorityName="Morelet" authorityYear="1877" box="[257,477,2001,2027]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis box="[257,378,2001,2027]" italics="true" pageId="27" pageNumber="28">minuscula</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFEA5F851F38EF80C" authority="Morelet" authorityName="Morelet" authorityYear="1877" box="[257,477,2001,2027]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="minuscula">
<emphasis id="FFCC8919D83FFFDCFEA5F851F329F80C" box="[257,378,2001,2027]" italics="true" pageId="27" pageNumber="28">minuscula</emphasis>
Morelet
</taxonomicName>
is here transferred from
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1856" box="[768,856,2001,2027]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[768,856,2001,2027]" italics="true" pageId="27" pageNumber="28">Gulella</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFCA4F851F10BF80C" authorityName="L. Pfeiffer" authorityYear="1856" box="[768,856,2001,2027]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis id="FFCC8919D83FFFDCFCA4F851F10BF80C" box="[768,856,2001,2027]" italics="true" pageId="27" pageNumber="28">Gulella</emphasis>
</taxonomicName>
to
<taxonomicName authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[894,1041,2001,2027]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis box="[894,1041,2001,2027]" italics="true" pageId="27" pageNumber="28">Dadagulella</emphasis>
<taxonomicName id="0AB82E88D83FFFDCFCDAF851F642F80C" authority="Rowson &amp; Tattersfield, 2013" authorityName="Rowson &amp; Tattersfield" authorityYear="2013" box="[894,1041,2001,2027]" class="Gastropoda" family="Streptaxidae" genus="Dadagulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="FFCC8919D83FFFDCFCDAF851F642F80C" box="[894,1041,2001,2027]" italics="true" pageId="27" pageNumber="28">Dadagulella</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1048,1154,2001,2027]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
<taxonomicNameLabel id="E4FF3462D83FFFDCFBBCF851F6D1F80C" box="[1048,1154,2001,2027]" pageId="27" pageNumber="28" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>