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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1009.57857" ID-GBIF-Dataset="a9e0f82e-7a5b-4494-aa30-2e77fcc1f4e9" ID-PMC="PMC7813806" ID-Pensoft-Pub="1313-2970-1009-123" ID-Pensoft-UUID="782AEBF1DA175A52A75908053031B979" ID-PubMed="33519257" ID-ZooBank="5CD8C6607CDB4664AF88B79090215BA3" ModsDocID="1313-2970-1009-123" checkinTime="1610662657732" checkinUser="pensoft" docAuthor="Yu, Li, Liu, Fengxiang, Zhang, Zengtao, Li, Daiqin &amp; Xu, Xin" docDate="2021" docId="66AD90D319485037B27EAEA229378ADC" docLanguage="en" docName="ZooKeys 1009: 123-138" docOrigin="ZooKeys 1009" docPubDate="2021-01-11" docSource="http://dx.doi.org/10.3897/zookeys.1009.57857" docTitle="Qiongthela nankai Yu &amp; Liu &amp; Zhang &amp; Li &amp; Xu 2021, sp. nov." docType="treatment" docUuid="7A75CB1F-E032-40BC-906D-F79CEAC64B96" docUuidSource="ZooBank" docVersion="4" id="782AEBF1DA175A52A75908053031B979" lastPageNumber="123" masterDocId="782AEBF1DA175A52A75908053031B979" masterDocTitle="Three new species of the segmented spider genus Qiongthela (Mesothelae, Liphistiidae) from Hainan Island, China" masterLastPageNumber="138" masterPageNumber="123" pageNumber="123" updateTime="1668149676036" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of the segmented spider genus Qiongthela (Mesothelae, Liphistiidae) from Hainan Island, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yu, Li</mods:namePart>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Fengxiang</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan, Hubei 430062, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Zengtao</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan, Hubei 430062, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Li, Daiqin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8269-7734</mods:nameIdentifier>
<mods:affiliation>Department of Biological Sciences, National University of Singapore, 14 Science Drive 4, Singapore 117543, Singapore</mods:affiliation>
<mods:nameIdentifier type="email">dbslidq@nus.edu.sg</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Xu, Xin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5632-6622</mods:nameIdentifier>
<mods:affiliation>College of Life Sciences, Hunan Normal University, Changsha, Hunan 410081, China &amp; State Key Laboratory of Biocatalysis and Enzyme Engineering, and Centre for Behavioural Ecology and Evolution (CBEE), School of Life Sciences, Hubei University, 368 Youyi Road, Wuhan, Hubei 430062, China</mods:affiliation>
<mods:nameIdentifier type="email">xuxin_09@163.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-01-11</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1009</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>123</mods:start>
<mods:end>138</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1009.57857</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1009.57857</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1009-123</mods:identifier>
<mods:identifier type="ZooBank">5CD8C6607CDB4664AF88B79090215BA3</mods:identifier>
<mods:identifier type="Pensoft-UUID">782AEBF1DA175A52A75908053031B979</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="176584324" LSID="urn:lsid:zoobank.org:act:7A75CB1F-E032-40BC-906D-F79CEAC64B96" httpUri="http://treatment.plazi.org/id/66AD90D319485037B27EAEA229378ADC" lastPageNumber="123" pageId="0" pageNumber="123">
<subSubSection pageId="0" pageNumber="123" type="nomenclature">
<paragraph pageId="0" pageNumber="123">
<taxonomicName LSID="http://zoobank.org/7A75CB1F-E032-40BC-906D-F79CEAC64B96" authority="Yu &amp; Liu &amp; Zhang &amp; Li &amp; Xu, 2021" authorityName="Yu &amp; Liu &amp; Zhang &amp; Li &amp; Xu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela nankai" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="nankai" status="sp. nov.">Qiongthela nankai</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="123">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Microhabitat and general somatic morphology of Qiongthela nankai sp. nov. A microhabitat B, C trapdoor with door closed and open D male (XUX- 2019 - 174, holotype) E female (XUX- 2019 - 173); Scale bars: 2 mm (D, E)." figureDoi="10.3897/zookeys.1009.57857.figure4" httpUri="https://binary.pensoft.net/fig/496168" pageId="0" pageNumber="123">Figures 4</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">, 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="type material">
<paragraph pageId="0" pageNumber="123">Type material.</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Holotype</emphasis>
♂: China, Hainan Province, Baisha City, Nankai Town, Nankai Village,
<geoCoordinate degrees="19.04" direction="north" orientation="latitude" precision="555" value="19.04">19.04°N</geoCoordinate>
,
<geoCoordinate degrees="109.39" direction="east" orientation="longitude" precision="555" value="109.39">109.39°E</geoCoordinate>
, alt. 300 m, 26 August 2019, D. Li, F.X. Liu, X. Xu and L. Yu leg., XUX-2019-174 (matured on 10 September 2019 at HNU).
<emphasis bold="true" italics="true" pageId="0" pageNumber="123">Paratypes</emphasis>
: 1 ♂, 2 ♀♀, same data as holotype, XUX-2019-172, 173, 175.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="diagnosis">
<paragraph pageId="0" pageNumber="123">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="123">
Males of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. nankai" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="nankai">
<emphasis italics="true" pageId="0" pageNumber="123">Q. nankai</emphasis>
</taxonomicName>
sp. nov. can be distinguished from those of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. qiongzhong" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="qiongzhong">
<emphasis italics="true" pageId="0" pageNumber="123">Q. qiongzhong</emphasis>
</taxonomicName>
,
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. yalin" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="yalin">
<emphasis italics="true" pageId="0" pageNumber="123">Q. yalin</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. yinggezui" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="yinggezui">
<emphasis italics="true" pageId="0" pageNumber="123">Q. yinggezui</emphasis>
</taxonomicName>
by the straight tegular marginal apophysis (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A, D</figureCitation>
), and the cymbial projection short and thick (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5G</figureCitation>
); from those of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. australis" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="australis">
<emphasis italics="true" pageId="0" pageNumber="123">Q. australis</emphasis>
</taxonomicName>
by the conductor with a bent apex (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5C, E, G</figureCitation>
), and the longer tegular marginal apophysis (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A, C, D, F</figureCitation>
); from those of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. dongfang" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="dongfang">
<emphasis italics="true" pageId="0" pageNumber="123">Q. dongfang</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. jianfeng" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="jianfeng">
<emphasis italics="true" pageId="0" pageNumber="123">Q. jianfeng</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. sanya" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="sanya">
<emphasis italics="true" pageId="0" pageNumber="123">Q. sanya</emphasis>
</taxonomicName>
by the tegular terminal apophysis with an abruptly narrowed and hooked apex (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A-E</figureCitation>
); from those of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. nui" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="nui">
<emphasis italics="true" pageId="0" pageNumber="123">Q. nui</emphasis>
</taxonomicName>
by the tegular marginal apophysis with a blunt edge (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A-G</figureCitation>
); from those of the rest of
<taxonomicName authorityName="Xu &amp; Kuntner" authorityYear="2015" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="123">Qiongthela</emphasis>
</taxonomicName>
species by the contrategulum with two distal edges (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A, D</figureCitation>
). Females of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. nankai" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="nankai">
<emphasis italics="true" pageId="0" pageNumber="123">Q. nankai</emphasis>
</taxonomicName>
sp. nov. differ from those of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. dongfang" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="dongfang">
<emphasis italics="true" pageId="0" pageNumber="123">Q. dongfang</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. wuzhi" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="wuzhi">
<emphasis italics="true" pageId="0" pageNumber="123">Q. wuzhi</emphasis>
</taxonomicName>
by the middle receptacular clusters situated close to each other (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5H-K</figureCitation>
); from those of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. bawang" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="bawang">
<emphasis italics="true" pageId="0" pageNumber="123">Q. bawang</emphasis>
</taxonomicName>
and
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. qiongzhong" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="qiongzhong">
<emphasis italics="true" pageId="0" pageNumber="123">Q. qiongzhong</emphasis>
</taxonomicName>
by the two pairs of receptacular clusters similar in size and shape (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5H-K</figureCitation>
); from those of the rest of
<taxonomicName authorityName="Xu &amp; Kuntner" authorityYear="2015" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="123">Qiongthela</emphasis>
</taxonomicName>
species by similar-sized receptacular clusters, and all situated along the anterior margin of the bursa copulatrix (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5H-K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="description">
<paragraph pageId="0" pageNumber="123">Description.</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Male</emphasis>
(holotype, Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Microhabitat and general somatic morphology of Qiongthela nankai sp. nov. A microhabitat B, C trapdoor with door closed and open D male (XUX- 2019 - 174, holotype) E female (XUX- 2019 - 173); Scale bars: 2 mm (D, E)." figureDoi="10.3897/zookeys.1009.57857.figure4" httpUri="https://binary.pensoft.net/fig/496168" pageId="0" pageNumber="123">4D</figureCitation>
). Carapace reddish brown; opisthosoma brown, with 12 yellow tergites, close to each other, the first 2-7 larger than the others, and the fourth largest; sternum narrow, much longer than wide; a few fine pointed hairs running over the ocular area; chelicerae with promargin of cheliceral groove bearing 11 denticles of variable size; legs with firm hairs and spines; 7 spinnerets. Measurements: BL 11.44, CL 5.31, CW 5.12, OL 5.63, OW 3.62; ALE&gt; PLE&gt; PME&gt; AME; leg I 14.01 (3.91 + 0.95 + 3.18 + 3.94 + 2.03), leg II 14.76 (4.05 + 1.30 + 3.28 + 4.00 + 2.13), leg III 15.64 (3.94 + 1.53 + 3.09 + 4.61 + 2.47), leg IV 21.12 (5.48 + 1.34 + 4.56 + 6.43 + 3.31).
</paragraph>
<caption doi="10.3897/zookeys.1009.57857.figure4" httpUri="https://binary.pensoft.net/fig/496168" pageId="0" pageNumber="123" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Figure 4.</emphasis>
Microhabitat and general somatic morphology of
<taxonomicName authorityName="Yu &amp; Liu &amp; Zhang &amp; Li &amp; Xu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela nankai" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="nankai">
<emphasis italics="true" pageId="0" pageNumber="123">Qiongthela nankai</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="123">A</emphasis>
microhabitat
<emphasis bold="true" pageId="0" pageNumber="123">B, C</emphasis>
trapdoor with door closed and open
<emphasis bold="true" pageId="0" pageNumber="123">D</emphasis>
male (XUX-2019-174, holotype)
<emphasis bold="true" pageId="0" pageNumber="123">E</emphasis>
female (XUX-2019-173); Scale bars: 2 mm (
<emphasis bold="true" pageId="0" pageNumber="123">D, E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">
<emphasis italics="true" pageId="0" pageNumber="123">Palp</emphasis>
.
</emphasis>
Cymbium with a short projection dorsally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5G</figureCitation>
); paracymbium unpigmented and unsclerotised prolaterally, with numerous setae at the tip (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A-C</figureCitation>
). Contrategulum with an irregular dentate edge proximally and two edges distally: the inner one dentate, and the outer one sharp, semi-translucent (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A, D, F</figureCitation>
). Marginal apophysis of tegulum long, with a blunt apex distally, distal portion of similar width as basal portion (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A, D</figureCitation>
), a proximally directed terminal apophysis of the tegulum with few denticles, abruptly narrowing to a hooked apex (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A-E</figureCitation>
). Conductor situated ventro-proximally on the embolus, the basal portion fused with the embolus, distal portion free, narrowing to a bent apex (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A-E</figureCitation>
). Embolus largely sclerotised, smooth ventrally, with several longitudinal ribs retrolaterally, and with a wide, flat opening of the sperm duct distally (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5A, D, E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Figure 5.</emphasis>
Male and female genital anatomy of
<taxonomicName authorityName="Yu &amp; Liu &amp; Zhang &amp; Li &amp; Xu" authorityYear="2021" class="Arachnida" family="Liphistiidae" genus="Qiongthela" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Qiongthela nankai" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="nankai">
<emphasis italics="true" pageId="0" pageNumber="123">Qiongthela nankai</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="123">A</emphasis>
left palp, prolateral view
<emphasis bold="true" pageId="0" pageNumber="123">B, E</emphasis>
left palp, ventral view
<emphasis bold="true" pageId="0" pageNumber="123">C</emphasis>
left palp, retrolateral view
<emphasis bold="true" pageId="0" pageNumber="123">D</emphasis>
left palp, distal view
<emphasis bold="true" pageId="0" pageNumber="123">F, G</emphasis>
left palp, dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">H, I</emphasis>
vulva, dorsal view
<emphasis bold="true" pageId="0" pageNumber="123">J, K</emphasis>
vulva, ventral view
<emphasis bold="true" pageId="0" pageNumber="123">A-C, G</emphasis>
XUX-2019-174 (holotype)
<emphasis bold="true" pageId="0" pageNumber="123">D-F</emphasis>
XUX-2019-172
<emphasis bold="true" pageId="0" pageNumber="123">H, J</emphasis>
XUX-2019-173
<emphasis bold="true" pageId="0" pageNumber="123">I, K</emphasis>
XUX-2019-175; Scale bars: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">Female</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Microhabitat and general somatic morphology of Qiongthela nankai sp. nov. A microhabitat B, C trapdoor with door closed and open D male (XUX- 2019 - 174, holotype) E female (XUX- 2019 - 173); Scale bars: 2 mm (D, E)." figureDoi="10.3897/zookeys.1009.57857.figure4" httpUri="https://binary.pensoft.net/fig/496168" pageId="0" pageNumber="123">4E</figureCitation>
). Carapace and opisthosoma color like in male, 12 opisthosoma tergites, close to each other, the first 2-7 larger than the others, and the fourth largest; sternum narrow, much longer than wide; a few fine pointed hairs running over the ocular area; chelicerae robust with promargin of cheliceral groove containing 10 denticles of variable size; legs with firm hairs and spines; 7 spinnerets. Measurements: BL 17.48, CL 8.45, CW 7.56, OL 8.47, OW 6.49; ALE&gt; PLE&gt; PME&gt; AME; palp 14.11 (5.02 + 1.93 + 3.41 + 3.75), leg I 15.49 (4.90 + 1.72 + 3.72+ 3.21 + 1.94), leg II 15.79 (4.91 + 1.97 + 3.41 + 3.38 + 2.12), leg III 16.31 (4.64 + 1.93 + 3.50 + 4.00 + 2.24), leg IV 23.11 (6.47 + 2.18 + 5.08 + 6.21 + 3.17).
</paragraph>
<paragraph pageId="0" pageNumber="123">
<emphasis bold="true" pageId="0" pageNumber="123">
<emphasis italics="true" pageId="0" pageNumber="123">Female genitalia</emphasis>
.
</emphasis>
Two pairs of receptacular clusters along the anterior margin of the bursa copulatrix, of similar size and shape, and the middle ones close to each other, with very short genital stalks (Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male and female genital anatomy of Qiongthela nankai sp. nov. A left palp, prolateral view B, E left palp, ventral view C left palp, retrolateral view D left palp, distal view F, G left palp, dorsal view H, I vulva, dorsal view J, K vulva, ventral view A-C, G XUX- 2019 - 174 (holotype) D-F XUX- 2019 - 172 H, J XUX- 2019 - 173 I, K XUX- 2019 - 175; Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1009.57857.figure5" httpUri="https://binary.pensoft.net/fig/496169" pageId="0" pageNumber="123">5H-K</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="variation">
<paragraph pageId="0" pageNumber="123">Variation.</paragraph>
<paragraph pageId="0" pageNumber="123">
Males and females vary in body size. Range of measurements in males (
<emphasis italics="true" pageId="0" pageNumber="123">N</emphasis>
= 2): BL 11.44-12.13, CL 5.31-5.65, CW 5.12-5.56, OL 5.63-6.35, OW 3.62-4.69; in females (
<emphasis italics="true" pageId="0" pageNumber="123">N</emphasis>
= 2): BL 13.09-17.48, CL 6.63-8.45, CW 6.04-7.56, OL 6.32-8.47, OW 5.05-6.49.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="etymology">
<paragraph pageId="0" pageNumber="123">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="123">The species epithet, a noun in apposition, refers to the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="distribution">
<paragraph pageId="0" pageNumber="123">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="123">Hainan (Baisha), China</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="genbank accession number">
<paragraph pageId="0" pageNumber="123">GenBank accession number.</paragraph>
<paragraph pageId="0" pageNumber="123">XUX-2019-174: MT900752.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="123" type="remarks">
<paragraph pageId="0" pageNumber="123">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="123">
The maximum and mean intraspecific genetic distances of
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. nankai" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="nankai">
<emphasis italics="true" pageId="0" pageNumber="123">Q. nankai</emphasis>
</taxonomicName>
sp. nov. are 0.6% and 0.3% based on K2P, respectively (
<emphasis italics="true" pageId="0" pageNumber="123">N</emphasis>
= 4). The interspecific genetic distances between
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. nankai" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="nankai">
<emphasis italics="true" pageId="0" pageNumber="123">Q. nankai</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. baishensis" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="baishensis">
<emphasis italics="true" pageId="0" pageNumber="123">Q. baishensis</emphasis>
</taxonomicName>
(GenBank accession code: KP229805), and
<taxonomicName class="Arachnida" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Q. yalin" order="Araneae" pageId="0" pageNumber="123" phylum="Arthropoda" rank="species" species="yalin">
<emphasis italics="true" pageId="0" pageNumber="123">Q. yalin</emphasis>
</taxonomicName>
sp. nov. are 10.3% and 16.1% based on K2P, respectively.
</paragraph>
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114
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Erzurum (Horasan,
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, Kandilli, Tatos
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); Erzurum (
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, Horasan,
<normalizedToken originalValue="İspir">Ispir</normalizedToken>
, Tortum) (
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1976
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<paragraph pageId="32" pageNumber="91">Material examined.</paragraph>
<paragraph pageId="32" pageNumber="91">
Ankara: Hacettepe
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, Beytepe
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,
<geoCoordinate degrees="32" direction="east" minutes="45" orientation="longitude" precision="15" seconds="06" value="32.751667">32°45'06&quot;E</geoCoordinate>
, 18.V.2005, 2 ♂♂, 29.V.2005, 1 ♂, 3.VI.2005, 1 ♀, 1 ♂, leg. E. Scheuchl; Konya:
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yolu,
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,
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<mods:identifier type="Zenodo-Dep">133270</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182226962" LSID="urn:lsid:plazi:treatment:66ADBD41923859BD99F8EF6C2E33B1C2" httpUri="http://treatment.plazi.org/id/66ADBD41923859BD99F8EF6C2E33B1C2" lastPageId="210" lastPageNumber="211" pageId="209" pageNumber="210">
<subSubSection pageId="209" pageNumber="210" type="nomenclature">
<paragraph pageId="209" pageNumber="210">
<taxonomicName LSID="66ADBD41-9238-59BD-99F8-EF6C2E33B1C2" authority="L. O. Williams, Fieldiana" authorityName="L. O. Williams, Fieldiana" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia plumosa" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="plumosa">Conostegia plumosa L.O. Williams, Fieldiana</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="209" pageNumber="210" type="reference_group">
<paragraph pageId="209" pageNumber="210">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia plumosa" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="plumosa">Conostegia plumosa</taxonomicName>
L.O. Williams, Fieldiana, Bot. 29: 562. 1963. Type: British Honduras [= Belize]. Middlesex: 200 ft elev, 10 July, 1929, W. Schipp 232 (holotype: F!, isotypes: A, CAS!, GH, MO!, US!).
</paragraph>
</subSubSection>
<subSubSection pageId="209" pageNumber="210" type="description">
<paragraph pageId="209" pageNumber="210">Description.</paragraph>
<paragraph pageId="209" pageNumber="210">
Trees 2.7-15 m tall with flattened branches which soon become terete and are covered with stellate and dendritic hairs in younger parts; the nodal line inconspicuous. Leaves of a pair equal to somewhat unequal in length. Petiole 0.9-4.5 cm. Leaf blade 6-21
<normalizedToken originalValue="×">x</normalizedToken>
2-6 cm, 3-5 plinerved, with innermost pair of veins diverging from the midvein above the base in opposite or alternate fashion, elliptic to elliptic ovate, the base acute to attenuate, the apex acute to acuminate, the margin serrulate, the adaxial surface stellate pubescent when young and glabrous with age, the abaxial surface tan to white color from the stellate and plumose hairs. Inflorescence a terminal panicle 3.3-12 cm long, the rachis covered with stellate and plumose hairs, linear, the bracteoles 2-8 mm long, deciduous. Pedicel absent or to 0.5 mm. Flowers 6 merous, calyptrate. Floral buds 4-6
<normalizedToken originalValue="×">x</normalizedToken>
2.4-3.25 mm, ovoid, the base rounded, the apex obtuse, slightly constricted in the middle, with six linear appendages ca. 2.5 mm long appressed to the top of the calyptra. Hypanthium 2.5-3
<normalizedToken originalValue="×">x</normalizedToken>
3-3.25 mm, with stellate and plumose hairs. Petals 4-6
<normalizedToken originalValue="×">x</normalizedToken>
1.5-2 mm, white, spathulate, spreading, glabrous. Stamens 12(-13), 4.25-5.5 mm long, the filament 2-2.5 mm long, white, the anthers 2.25-2.75
<normalizedToken originalValue="×">x</normalizedToken>
0.5-0.75 mm, linear-oblong and somewhat recirved, yellow, the pore 0.1-0.2 mm in diameter, subterminal. Ovary 6 locular, inferior, the apex glabrous and forming an irregular collar around the style. Style 3.5-6 mm, apparently straight, distance between the anthers and stigma ca. 0-1 mm, stigma truncate, 0.35-0.65 mm wide. Berry 5-6
<normalizedToken originalValue="×">x</normalizedToken>
5-6 mm, purple-black. Seeds 0.65-1.2 mm, obliquely pyramidal, the testa angulate and smooth.
</paragraph>
</subSubSection>
<subSubSection lastPageId="210" lastPageNumber="211" pageId="209" pageNumber="210" type="distribution">
<paragraph pageId="209" pageNumber="210">Distribution</paragraph>
<paragraph pageId="209" pageNumber="210">
(Fig.
<figureCitation captionStart="Figure 185" captionStartId="F185" captionText="Figure 185. Distribution of Conostegia plumosa." figureDoi="10.3897/phytokeys.67.6703.figure185" httpUri="https://binary.pensoft.net/fig/96543" pageId="209" pageNumber="210">185</figureCitation>
). Belize, Guatemala, Honduras and in Chiapas in Mexico, at 50-1000 m in elevation.
</paragraph>
<caption ID-Zenodo-Dep="974027" doi="10.3897/phytokeys.67.6703.figure185" httpUri="https://binary.pensoft.net/fig/96543" pageId="209" pageNumber="210" start="Figure 185" startId="F185">
<paragraph pageId="209" pageNumber="210">
<emphasis bold="true" pageId="209" pageNumber="210">Figure 185.</emphasis>
Distribution of
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia plumosa" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="plumosa">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia plumosa</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph lastPageId="210" lastPageNumber="211" pageId="209" pageNumber="210">
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia plumosa" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="plumosa">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia plumosa</emphasis>
</taxonomicName>
is a rare species and one of the few in the genus to have the abaxial leaf surface tan colored from the dense indument covering it. This characteristic leaf underside is shared with
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel, Almeda &amp; A. Estrada" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia dissitinervia" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="dissitinervia">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia dissitinervia</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia centrosperma" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="centrosperma">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia centrosperma</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Gleason" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia incurva" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="incurva">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia incurva</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Donn. Sm." baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia oligocephala" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="oligocephala">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia oligocephala</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia xalapensis" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="xalapensis">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia xalapensis</emphasis>
</taxonomicName>
. In particular,
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia plumosa" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="plumosa">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia plumosa</emphasis>
</taxonomicName>
has been confused with
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Donn. Sm." baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia oligocephala" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="oligocephala">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia oligocephala</emphasis>
</taxonomicName>
and
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia xalapensis" order="Myrtales" pageId="209" pageNumber="210" phylum="Tracheophyta" rank="species" species="xalapensis">
<emphasis italics="true" pageId="209" pageNumber="210">Conostegia xalapensis</emphasis>
</taxonomicName>
, both of which occur in the same general area of northern Central America. The other species mentioned are endemic to southern Central America.
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia plumosa" order="Myrtales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="plumosa">
<emphasis italics="true" pageId="210" pageNumber="211">
<pageBreakToken pageId="210" pageNumber="211" start="start">Conostegia</pageBreakToken>
plumosa
</emphasis>
</taxonomicName>
shares the calyptrate calyx and short style with
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia xalapensis" order="Myrtales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="xalapensis">
<emphasis italics="true" pageId="210" pageNumber="211">Conostegia xalapensis</emphasis>
</taxonomicName>
but not with
<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Donn. Sm." baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia oligocephala" order="Myrtales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="oligocephala">
<emphasis italics="true" pageId="210" pageNumber="211">Conostegia oligocephala</emphasis>
</taxonomicName>
which lacks the calyptrate calyx and has an exserted style.
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia plumosa" order="Myrtales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="plumosa">
<emphasis italics="true" pageId="210" pageNumber="211">Conostegia plumosa</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia xalapensis" order="Myrtales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="xalapensis">
<emphasis italics="true" pageId="210" pageNumber="211">Conostegia xalapensis</emphasis>
</taxonomicName>
on the basis of its evident calyptra appendices which are absent in
<taxonomicName class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia xalapensis" order="Myrtales" pageId="210" pageNumber="211" phylum="Tracheophyta" rank="species" species="xalapensis">
<emphasis italics="true" pageId="210" pageNumber="211">Conostegia xalapensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="210" pageNumber="211" type="materials_examined">
<paragraph pageId="210" pageNumber="211">Specimens examined.</paragraph>
<paragraph pageId="210" pageNumber="211">
<emphasis bold="true" pageId="210" pageNumber="211">
<collectingCountry name="Mexico">MEXICO</collectingCountry>
</emphasis>
(fide Schnell).
<emphasis bold="true" pageId="210" pageNumber="211">
<collectingRegion country="Mexico" name="Chiapas">Chiapas</collectingRegion>
</emphasis>
: km 33 S of SE on the road to Mal Paso, near
<collectingRegion country="Mexico" name="Tabasco">Tabasco</collectingRegion>
border, Roe et al. 1371 (
<collectingCountry name="United States of America">US</collectingCountry>
).
</paragraph>
<paragraph pageId="210" pageNumber="211">
<materialsCitation collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" country="BELIZE" location="Mountain Pine Ridge" specimenCount="1" stateProvince="Cayo">
<emphasis bold="true" pageId="210" pageNumber="211">
<collectingCountry name="Belize">BELIZE</collectingCountry>
.
<collectingRegion country="Belize" name="Cayo">Cayo</collectingRegion>
</emphasis>
: Road to Caracol at Guacamallo Bridge, Balick 3104 (NY); along Hummingbird Highway at mile 28.5, Dwyer 11208 (MO, NY); Humming Bird Highway, Gentle 8388 (NY);
<location LSID="urn:lsid:plazi:treatment:66ADBD41923859BD99F8EF6C2E33B1C2:4D4A21FF12AB51A2A6F03F663B08938F" country="BELIZE" name="Mountain Pine Ridge" stateProvince="Cayo">Mountain Pine Ridge</location>
, San Agustin, Lundell 6608, 6770 (NY)
</materialsCitation>
.
<emphasis bold="true" pageId="210" pageNumber="211">
<collectingRegion country="Belize" name="Stann Creek">Stann Creek</collectingRegion>
</emphasis>
: beyond Middlesex, Gentle 2758 (NY);
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.7358848" unit="mi" value="17.0">17 miles</quantity>
section Stann Creek Valley, Gentle 9258 (NY).
</paragraph>
<paragraph pageId="210" pageNumber="211">
<emphasis bold="true" pageId="210" pageNumber="211">
<collectingCountry name="Guatemala">GUATEMALA</collectingCountry>
.
<collectingRegion country="Guatemala" name="Izabal">Izabal</collectingRegion>
</emphasis>
: slopes WNW of (above) El Estor, along margin of open pit nickel mine, Stevens and Martinez 25247 (NY).
</paragraph>
<paragraph pageId="210" pageNumber="211">
<emphasis bold="true" pageId="210" pageNumber="211">
<collectingCountry name="Honduras">HONDURAS</collectingCountry>
.
<collectingRegion country="Honduras" name="Olancho">Olancho</collectingRegion>
</emphasis>
:
<normalizedToken originalValue="Montaña">Montana</normalizedToken>
La Bellota 20 kms al N.O. de Campamento, Molina 13436 (NY).
</paragraph>
</subSubSection>
</treatment>
</document>

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data/66/AD/CA/66ADCACABC3D2E6ADE685EC58D0F1EA1.xml Normal file
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<treatment ID-GBIF-Taxon="159359421" LSID="urn:lsid:plazi:treatment:66ADCACABC3D2E6ADE685EC58D0F1EA1" httpUri="http://treatment.plazi.org/id/66ADCACABC3D2E6ADE685EC58D0F1EA1" lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19">
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<taxonomicName LSID="A10B9961-8C77-503D-BA93-6EF4305E08E8" authority="(Hochr.) Dorr" authorityName="Dorr" authorityYear="2011" baseAuthorityName="Hochr." class="Magnoliopsida" family="Malvaceae" genus="Trichospermum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Trichospermum lessertianum" order="Malvales" pageId="2" pageNumber="19" phylum="Magnoliophyta" rank="species" species="lessertianum" status="comb. nov.">Trichospermum lessertianum (Hochr.) Dorr</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="19">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19" type="reference_group">
<paragraph lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19">
<taxonomicName authorityName="Hochr" authorityYear="1914" class="Magnoliopsida" family="Malvaceae" genus="Belotia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="2" pageNumber="19" phylum="Tracheophyta" rank="species" species="lessertiana">Belotia lessertiana</taxonomicName>
Hochr., Annuaire Conserv. Jard Bot.
<normalizedToken originalValue="Genève">Geneve</normalizedToken>
18-19: 90. 1914, as &quot;
<taxonomicName genus="Lessertiana" lsidName="" pageId="2" pageNumber="19" rank="genus">Lessertiana</taxonomicName>
.&quot;
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Belotia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="2" pageNumber="19" phylum="Tracheophyta" rank="species" species="grewiifolia">Belotia grewiifolia</taxonomicName>
var.
<emphasis italics="true" pageId="2" pageNumber="19">lessertiana</emphasis>
(Hochr.) Vict., Contr. Inst.
<pageBreakToken pageId="3" pageNumber="20" start="start">Bot</pageBreakToken>
. Univ.
<normalizedToken originalValue="Montréal">Montreal</normalizedToken>
63: 13. 1948. TYPE: CUBA. La Havane, s.d. (fl),
<taxonomicName genus="Delessert" lsidName="" pageId="3" pageNumber="20" rank="genus">Delessert</taxonomicName>
s.n. (holotype: G-DEL; isotypes: K, NY! [00084148], P). (Basionym)
</paragraph>
<paragraph pageId="3" pageNumber="20">
<taxonomicName authorityName="Sprague" authorityYear="1921" class="Magnoliopsida" family="Malvaceae" genus="Belotia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="caribaea">Belotia caribaea</taxonomicName>
Sprague, Bull. Misc. Inform. Kew 1921(7): 276. 1921.
<taxonomicName authorityName="Kosterm" authorityYear="1962" baseAuthorityName="Sprague" class="Magnoliopsida" family="Malvaceae" genus="Trichospermum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="caribaeum">Trichospermum caribaeum</taxonomicName>
(Sprague) Kosterm., Reinwardtia 6(3): 278. 1962. TYPE: ST. LUCIA, s.d. (fl, fr),
<taxonomicName genus="Anderson" lsidName="" pageId="3" pageNumber="20" rank="genus">Anderson</taxonomicName>
s.n. (holotype: K! [K000381875]).
</paragraph>
<paragraph pageId="3" pageNumber="20">
<taxonomicName authorityName="Sprague" authorityYear="1921" class="Magnoliopsida" family="Malvaceae" genus="Belotia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="reticulata">Belotia reticulata</taxonomicName>
Sprague, Bull. Misc. Inform. Kew 1921(7): 277. 1921.
<taxonomicName authorityName="Kosterm" authorityYear="1962" baseAuthorityName="Sprague" class="Magnoliopsida" family="Malvaceae" genus="Trichospermum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="reticulatum">Trichospermum reticulatum</taxonomicName>
(Sprague) Kosterm., Reinwardtia 6(3): 279. 1962. TYPE: NICARAGUA. Chontales,
<emphasis italics="true" pageId="3" pageNumber="20">Seemann 11</emphasis>
(holotype: K).
</paragraph>
<paragraph pageId="3" pageNumber="20">
<taxonomicName authorityName="Sprague" authorityYear="1921" class="Magnoliopsida" family="Malvaceae" genus="Belotia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="campbellii">Belotia campbellii</taxonomicName>
Sprague, Bull. Misc. Inform. Kew 1921(7): 277. 1921, as &quot;
<taxonomicName authorityName="Korf" authorityYear="1973" baseAuthorityName="Sacc." genus="Campbellii" lsidName="" pageId="3" pageNumber="20" rank="genus">Campbellii</taxonomicName>
.&quot; TYPE: BELIZE. Seven Hills Estate, s.d. (fl, fr),
<emphasis italics="true" pageId="3" pageNumber="20">E.J.F. Campbell 75</emphasis>
(holotype: K! [K000381880]; isotype: F).
</paragraph>
<paragraph pageId="3" pageNumber="20">
<taxonomicName authorityName="Sprague" authorityYear="1921" class="Magnoliopsida" family="Malvaceae" genus="Belotia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="tabascana">Belotia tabascana</taxonomicName>
Sprague, Bull. Misc. Inform. Kew 1921(7): 278. 1921.
<taxonomicName authorityName="Kosterm" authorityYear="1962" baseAuthorityName="Sprague" class="Magnoliopsida" family="Malvaceae" genus="Trichospermum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="tabascanum">Trichospermum tabascanum</taxonomicName>
(Sprague) Kosterm., Reinwardtia 6(3): 279. 1962. TYPE: MEXICO. Tabasco: Lomas de San
<normalizedToken originalValue="Sebastián">Sebastian</normalizedToken>
, 26 Mar 1889 (fl),
<emphasis italics="true" pageId="3" pageNumber="20">Rovirosa 416</emphasis>
(holotype: K; isotypes: F, NY! [00546807], US! [00098426]).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="20" type="distribution">
<paragraph pageId="3" pageNumber="20">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="20">
Southern Mexico to Costa Rica, and in western Cuba. A collection stated to be from St. Lucia (the type of
<taxonomicName authorityName="Sprague" authorityYear="1921" class="Magnoliopsida" family="Malvaceae" genus="Belotia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="caribaea">Belotia caribaea</taxonomicName>
) represents either material cultivated in the St. Vincent Botanic Garden or mislabeled material from Cuba (
<bibRefCitation author="Bornstein, AJ" editor="Howard, RA" journalOrPublisher="Jamaica Plain, Massachusetts: Arnold Arboretum, Harvard University" pageId="4" pageNumber="21" pagination="184 - 199" refId="B1" refString="Bornstein, AJ, 1989. Tiliaceae. In: Howard, RA, Ed., Flora of the Lesser Antilles, vol. 5. Jamaica Plain, Massachusetts: Arnold Arboretum, Harvard University: 184 - 199" title="Tiliaceae." volumeTitle="Flora of the Lesser Antilles, vol. 5." year="1989">Bornstein 1989: 185-186</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="20" type="note">
<paragraph pageId="3" pageNumber="20">Note.</paragraph>
<paragraph pageId="3" pageNumber="20">
The name
<taxonomicName authorityName="Kosterm" authorityYear="1962" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Malvaceae" genus="Trichospermum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="grewiifolium">Trichospermum grewiifolium</taxonomicName>
(A. Rich.) Kosterm. is frequently applied to this species, but it is illegitimate because as explained in the text
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Belotia" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Tracheophyta" rank="species" species="grewiifolia">Belotia grewiifolia</taxonomicName>
A. Rich. was nomenclaturally superfluous when published (
<bibRefCitation author="McNeill, J" journalOrPublisher="Regnum Vegetabile 146" pageId="4" pageNumber="21" pagination="1 - 568" refId="B11" refString="McNeill, J, Barrie, FR, Burdet, HM, Demoulin, V, Hawksworth, DL, Marhold, K, Nicolson, DH, Prado, J, Silva, PC, Skog, JE, Wiersema, JH, Turland, NJ, 2006. International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005. Regnum Vegetabile 146 i-xviii: 1 - 568" title="International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005." volume="i-xviii" volumeTitle="Regnum Vegetabile 146" year="2006">McNeill et al. 2006</bibRefCitation>
: Art. 52.1) and cannot serve as the basionym for this combination. More recently,
<bibRefCitation author="Rodriguez Fuentes, A" editor="Greuter, W" journalOrPublisher="Flora de la Republica de Cuba: Serie A, Plantas Vasculares" pageId="5" pageNumber="22" pagination="1 - 38" refId="B16" refString="Rodriguez Fuentes, A, 2000. Tiliaceae. Flora de la Republica de Cuba: Serie A, Plantas Vasculares 3 (5): 1 - 38" title="Tiliaceae." volume="3" volumeTitle="Flora de la Republica de Cuba: Serie A, Plantas Vasculares" year="2000">
<normalizedToken originalValue="Rodríguez">Rodriguez</normalizedToken>
Fuentes (2000: 32)
</bibRefCitation>
accepted
<taxonomicName authorityName="Kosterm" authorityYear="1962" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Malvaceae" genus="Trichospermum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="grewiifolium">Trichospermum grewiifolium</taxonomicName>
as a new species published by
<bibRefCitation author="Kostermans, AJGH" journalOrPublisher="Reinwardtia" pageId="4" pageNumber="21" pagination="277 - 279" refId="B9" refString="Kostermans, AJGH, 1962. The genera Belotia Rich. and Trichospermum Bl. (Tiliaceae). Reinwardtia 6 (3): 277 - 279" title="The genera Belotia Rich. and Trichospermum Bl. (Tiliaceae)." volume="6" year="1962">Kostermans (1962)</bibRefCitation>
apparently in the belief that Kostermans had explicitly excluded the purported
<normalizedToken originalValue="basionym’s">basionym's</normalizedToken>
type, but this is debatable and in any case
<taxonomicName authorityName="Kosterm" authorityYear="1962" baseAuthorityName="A. Rich." class="Magnoliopsida" family="Malvaceae" genus="Trichospermum" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Malvales" pageId="3" pageNumber="20" phylum="Magnoliophyta" rank="species" species="grewiifolium">Trichospermum grewiifolium</taxonomicName>
is not a valid name as Kostermans failed to designate a nomenclatural type (
<bibRefCitation author="McNeill, J" journalOrPublisher="Regnum Vegetabile 146" pageId="4" pageNumber="21" pagination="1 - 568" refId="B11" refString="McNeill, J, Barrie, FR, Burdet, HM, Demoulin, V, Hawksworth, DL, Marhold, K, Nicolson, DH, Prado, J, Silva, PC, Skog, JE, Wiersema, JH, Turland, NJ, 2006. International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005. Regnum Vegetabile 146 i-xviii: 1 - 568" title="International Code of Botanical Nomenclature (Vienna Code) adopted by the Seventeenth International Botanical Congress, Vienna, Austria, July 2005." volume="i-xviii" volumeTitle="Regnum Vegetabile 146" year="2006">McNeill et al. 2006</bibRefCitation>
: Art. 37.1).
</paragraph>
</subSubSection>
</treatment>
</document>