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<document ID-DOI="http://dx.doi.org/10.3897/zse.98.82649" ID-Pensoft-Pub="1860-0743-1-137" ID-Pensoft-UUID="58CEEFF0E0A058EA845BB606E20FBB1C" ID-ZooBank="E428377B518645AEA97FA82D4F5C5BD4" LSID="urn:lsid:zoobank.org:act:AFC772CC-2A82-4533-BF66-0317BAACE2AD" ModsDocID="1860-0743-98-1-137" checkinTime="1651791036176" checkinUser="pensoft" docAuthor="Framenau, Volker W. &amp; Castanheira, Pedro de S." docDate="2022" docId="66912C9761705B7AA137364BEDF43EE7" docLanguage="en" docName="ZoosystEvol 98(1): 137-149" docOrigin="Zoosystematics and Evolution 98 (1)" docPubDate="2022-05-05" docSource="http://dx.doi.org/10.3897/zse.98.82649" docTitle="Mangrovia Framenau &amp; Castanheira 2022, gen. nov." docType="treatment" docUuid="AFC772CC-2A82-4533-BF66-0317BAACE2AD" docUuidSource="ZooBank" docVersion="1" id="58CEEFF0E0A058EA845BB606E20FBB1C" lastPageNumber="137" masterDocId="58CEEFF0E0A058EA845BB606E20FBB1C" masterDocTitle="A new genus of Australian orb-weaving spider with extreme sexual size dimorphism (Araneae, Araneidae)" masterLastPageNumber="149" masterPageNumber="137" pageId="0" pageNumber="137" updateTime="1651791036176" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new genus of Australian orb-weaving spider with extreme sexual size dimorphism (Araneae, Araneidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Framenau, Volker W.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7724-3831</mods:nameIdentifier>
<mods:affiliation>Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia &amp; Zoological Museum Hamburg, Leibnitz Institute for the Analysis of Biodiversity Change (LIB), Centre for Taxonomy &amp; Morphology, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
<mods:nameIdentifier type="email">volker.framenau@murdoch.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Castanheira, Pedro de S.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0623-1622</mods:nameIdentifier>
<mods:affiliation>Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-05-05</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>98</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>137</mods:start>
<mods:end>149</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zse.98.82649</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.98.82649</mods:identifier>
<mods:identifier type="Pensoft-Pub">1860-0743-1-137</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:AFC772CC-2A82-4533-BF66-0317BAACE2AD" httpUri="http://treatment.plazi.org/id/66912C9761705B7AA137364BEDF43EE7" lastPageNumber="137" pageId="0" pageNumber="137">
<subSubSection pageId="0" pageNumber="137" type="nomenclature">
<paragraph pageId="0" pageNumber="137">
<taxonomicName LSID="http://zoobank.org/AFC772CC-2A82-4533-BF66-0317BAACE2AD" authority="Framenau &amp; Castanheira, 2022" authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus" status="gen. nov.">Mangrovia</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="137">gen. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="type-species">
<paragraph pageId="0" pageNumber="137">Type-species.</paragraph>
<paragraph pageId="0" pageNumber="137">
<taxonomicName authority="Walckenaer, 1805" authorityName="Framenau &amp; Castanheira" authorityYear="2022" baseAuthorityName="Framenau &amp; Castanheira" baseAuthorityYear="2022" class="Arachnida" family="Araneidae" genus="Epeira" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Epeira albida" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="albida">
<emphasis italics="true" pageId="0" pageNumber="137">Epeira albida</emphasis>
</taxonomicName>
L. Koch, 1871, designated here.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="etymology">
<paragraph pageId="0" pageNumber="137">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="137">The genus-group name is derived from the general habitat preferences of the two species, which are often found in coastal mangroves and woodlands. The gender is feminine.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="diagnosis">
<paragraph pageId="0" pageNumber="137">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="137">
Within an Australian context,
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov. males have only a single patellar spine on the pedipalp, a character considered a putative synapomorphy of the backobourkiines. However,
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen nov. differ considerably from members of the backobourkiines by somatic and genital morphology. Both species in the genus display eSSD with females about 3-5 times larger than males, but eSSD is absent in the backobourkiines (with the exception of
<taxonomicName genus="B." lsidName="B. collina" pageId="0" pageNumber="137" rank="species" species="collina">
<emphasis italics="true" pageId="0" pageNumber="137">B. collina</emphasis>
</taxonomicName>
and species in the
<taxonomicName class="Arachnida" family="Araneidae" genus="Backobourkia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Backobourkia dehaani" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="dehaani">
<emphasis italics="true" pageId="0" pageNumber="137">Backobourkia dehaani</emphasis>
</taxonomicName>
-group (sensu
<bibRefCitation author="Yin, CM" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="137" refId="B31" refString="Yin, CM, Wang, JF, Zhu, MS, Xie, LP, Peng, XJ, Bao, YH, 1997. Fauna Sinica: Arachnida: Araneae: Araneidae. Science Press, Beijing" title="Fauna Sinica: Arachnida: Araneae: Araneidae." year="1997">Yin et al. 1997</bibRefCitation>
)). The median apophysis in backobourkiines is elongate transverse with the base forming an arch over the radix, but it is short with an apical, spine-like projection in
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov. The female epigyne in
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov. has a terminal pocket, absent in all backobourkiines.
</paragraph>
<paragraph pageId="0" pageNumber="137">
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov. differs from the members of the zealaraneines as currently known (see Introduction section) by its eSSD, absent in any known zealaraneines, and the presence of only one patellar spine on the male pedipalp, whereas there are always two in zealaraneines (
<bibRefCitation author="Court, DJ" journalOrPublisher="Otago Museum Bulletin" pageId="0" pageNumber="137" pagination="68 - 124" refId="B3" refString="Court, DJ, Forster, RR, 1988. The spiders of New Zealand: Part VI. Araneidae - Araneinae. Otago Museum Bulletin 6: 68 - 124" title="The spiders of New Zealand: Part VI. Araneidae - Araneinae." volume="6" year="1988">Court and Forster 1988</bibRefCitation>
). The median apophysis of zealaraneines is elongate transverse, with a variable number of spine-like protrusions, but much shorter and only with a single spine-like protrusion in
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov.
</paragraph>
<paragraph pageId="0" pageNumber="137">
Outside the backobourkiines and zealaraneines,
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov. appears most similar to species of
<taxonomicName authorityName="E.Simon" authorityYear="1864" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Neoscona</emphasis>
</taxonomicName>
(
<bibRefCitation author="Berman, JD" journalOrPublisher="Bulletin of the Museum of Comparative Zoology" pageId="0" pageNumber="137" pagination="465 - 500" refId="B1" refString="Berman, JD, Levi, HW, 1971. The orb weaver genus Neoscona in North America (Araneae: Araneidae). Bulletin of the Museum of Comparative Zoology 141: 465 - 500" title="The orb weaver genus Neoscona in North America (Araneae: Araneidae)." volume="141" year="1971">Berman and Levi 1971</bibRefCitation>
;
<bibRefCitation author="Grasshoff, M" journalOrPublisher="Annales de la Musee Royal de L'Afrique Centrale" pageId="0" pageNumber="137" pagination="1 - 123" refId="B10" refString="Grasshoff, M, 1986. Die Radnetzspinnen-Gattung Neoscona in Afrika. Annales de la Musee Royal de L'Afrique Centrale 250: 1 - 123" title="Die Radnetzspinnen-Gattung Neoscona in Afrika." volume="250" year="1986">Grasshoff 1986</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1155/1992/93912" author="Levi, H" journalOrPublisher="Psyche, Cambridge" pageId="0" pageNumber="137" pagination="221 - 239" refId="B19" refString="Levi, H, 1993a. American Neoscona and corrections to previous revisions of Neotropical orb-weavers (Araneae: Araneidae). Psyche, Cambridge 99 (2-3): 221 - 239, DOI: https://doi.org/10.1155/1992/93912" title="American Neoscona and corrections to previous revisions of Neotropical orb-weavers (Araneae: Araneidae)." url="https://doi.org/10.1155/1992/93912" volume="99" year="1993 a">Levi 1993a</bibRefCitation>
), in particular with respect to the male genitalia. However, eSSD is absent in any known
<taxonomicName authorityName="E.Simon" authorityYear="1864" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Neoscona</emphasis>
</taxonomicName>
species and males of
<taxonomicName authorityName="E.Simon" authorityYear="1864" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Neoscona</emphasis>
</taxonomicName>
have distinctly enlarged tibiae on the second leg with numerous and strong spines, often in diagnostic arrangement.
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov. males do not have enlarged tibiae II. In contrast to
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov., males in
<taxonomicName authorityName="E.Simon" authorityYear="1864" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Neoscona</emphasis>
</taxonomicName>
have two patellar spines on the pedipalp, not one, and the embolus of
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov. has a subterminal side branch (e.g. Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mangrovia albida comb. nov., expanded male left pedipalp (QM S 67277). A. Apico-ventral view; B. Mesal view; C. Retrolateral view; D. Dorsal view. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zse.98.82649.figure2" httpUri="https://binary.pensoft.net/fig/680581" pageId="0" pageNumber="137">2B, C</figureCitation>
), not present in
<taxonomicName authorityName="E.Simon" authorityYear="1864" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Neoscona</emphasis>
</taxonomicName>
. An embolus lamella is absent in
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia" order="Araneae" pageId="0" pageNumber="137" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia</emphasis>
</taxonomicName>
gen. nov. males, but present in
<taxonomicName authorityName="E.Simon" authorityYear="1864" class="Arachnida" family="Araneidae" genus="Neoscona" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neoscona" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="137">Neoscona</emphasis>
</taxonomicName>
(e.g.
<bibRefCitation DOI="https://doi.org/10.1155/1992/93912" author="Levi, H" journalOrPublisher="Psyche, Cambridge" pageId="0" pageNumber="137" pagination="221 - 239" refId="B19" refString="Levi, H, 1993a. American Neoscona and corrections to previous revisions of Neotropical orb-weavers (Araneae: Araneidae). Psyche, Cambridge 99 (2-3): 221 - 239, DOI: https://doi.org/10.1155/1992/93912" title="American Neoscona and corrections to previous revisions of Neotropical orb-weavers (Araneae: Araneidae)." url="https://doi.org/10.1155/1992/93912" volume="99" year="1993 a">Levi 1993a</bibRefCitation>
; fig. 6).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="description">
<paragraph pageId="0" pageNumber="137">Description.</paragraph>
<paragraph pageId="0" pageNumber="137">
Small to medium-sized orb-weaving spiders with eSSD (TL males ca. 2.5-3 mm, females ca. 8-10 mm). Carapace (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Mangrovia albida comb. nov., male (QM S 67277). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left pedipalp, ventral view; D. Left pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure1" httpUri="https://binary.pensoft.net/fig/680580" pageId="0" pageNumber="137">1A</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Mangrovia albida comb. nov., female (AM KS. 98754). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view. Scale bars: 2 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zse.98.82649.figure3" httpUri="https://binary.pensoft.net/fig/680582" pageId="0" pageNumber="137">3A</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mangrovia occidentalis sp. nov., male (WAM T 77397). A. Habitus, dorsal view; B. Habitus, ventral view; C. Right pedipalp, ventral view; D. Right pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure6" httpUri="https://binary.pensoft.net/fig/680585" pageId="0" pageNumber="137">6A</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Mangrovia occidentalis sp. nov., female (WAM T 75326). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view. Scale bars: 2 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zse.98.82649.figure8" httpUri="https://binary.pensoft.net/fig/680587" pageId="0" pageNumber="137">8A</figureCitation>
) rounded pear-shaped, longer than wide; colouration variable from light brown to reddish-brown with dusky edges on males. Eyes: AME largest, row of PE slightly recurved, lateral eyes almost touching, PLE separated from PME by approximately their diameter in males and by more than their diameter in females. Chelicerae paturon and fangs yellowish- to reddish-brown. Females with four promarginal teeth and three retromarginal teeth, males with three promarginal teeth and two retromarginal teeth. Labium wider than long, subtriangular, with front end bulging and white. Endites elongate-rounded, beige to light brown with antero-mesal corner shiny and white. Sternum heart-shaped, slightly longer than wide, with dark edges (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Mangrovia albida comb. nov., male (QM S 67277). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left pedipalp, ventral view; D. Left pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure1" httpUri="https://binary.pensoft.net/fig/680580" pageId="0" pageNumber="137">1B</figureCitation>
,
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Mangrovia albida comb. nov., female (AM KS. 98754). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view. Scale bars: 2 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zse.98.82649.figure3" httpUri="https://binary.pensoft.net/fig/680582" pageId="0" pageNumber="137">3B</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mangrovia occidentalis sp. nov., male (WAM T 77397). A. Habitus, dorsal view; B. Habitus, ventral view; C. Right pedipalp, ventral view; D. Right pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure6" httpUri="https://binary.pensoft.net/fig/680585" pageId="0" pageNumber="137">6B</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Mangrovia occidentalis sp. nov., female (WAM T 75326). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view. Scale bars: 2 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zse.98.82649.figure8" httpUri="https://binary.pensoft.net/fig/680587" pageId="0" pageNumber="137">8B</figureCitation>
). Legs: leg formula I&gt; II&gt; IV&gt; III (males) or I&gt; IV&gt; II&gt; III (females). Abdomen about as long as wide, dorsum with dark folium pattern in males, varying from uniformly grey to beige with anterior black area on females; venter black with a pair of white guanine spots. Genitalia: male pedipalp patella with a single strong spine (e.g. Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Mangrovia albida comb. nov., male (QM S 67277). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left pedipalp, ventral view; D. Left pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure1" httpUri="https://binary.pensoft.net/fig/680580" pageId="0" pageNumber="137">1C, D</figureCitation>
); paracymbium poorly developed (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Mangrovia albida comb. nov., male (QM S 67277). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left pedipalp, ventral view; D. Left pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure1" httpUri="https://binary.pensoft.net/fig/680580" pageId="0" pageNumber="137">1D</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mangrovia albida comb. nov., expanded male left pedipalp (QM S 67277). A. Apico-ventral view; B. Mesal view; C. Retrolateral view; D. Dorsal view. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zse.98.82649.figure2" httpUri="https://binary.pensoft.net/fig/680581" pageId="0" pageNumber="137">2C</figureCitation>
); radix elongate (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mangrovia albida comb. nov., expanded male left pedipalp (QM S 67277). A. Apico-ventral view; B. Mesal view; C. Retrolateral view; D. Dorsal view. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zse.98.82649.figure2" httpUri="https://binary.pensoft.net/fig/680581" pageId="0" pageNumber="137">2B</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mangrovia occidentalis sp. nov., expanded right left pedipalp (WAM T 77397). A. Apico-dorsal view; B. Mesal view. Scale bars: 0.1 mm (A, B)." figureDoi="10.3897/zse.98.82649.figure7" httpUri="https://binary.pensoft.net/fig/680586" pageId="0" pageNumber="137">7A</figureCitation>
); no obvious stipes, possibly fused with radix (see Discussion); median apophysis oval, bearing a heavily sclerotized and acute apical process (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Mangrovia albida comb. nov., male (QM S 67277). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left pedipalp, ventral view; D. Left pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure1" httpUri="https://binary.pensoft.net/fig/680580" pageId="0" pageNumber="137">1C</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mangrovia albida comb. nov., expanded male left pedipalp (QM S 67277). A. Apico-ventral view; B. Mesal view; C. Retrolateral view; D. Dorsal view. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zse.98.82649.figure2" httpUri="https://binary.pensoft.net/fig/680581" pageId="0" pageNumber="137">2A</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mangrovia occidentalis sp. nov., male (WAM T 77397). A. Habitus, dorsal view; B. Habitus, ventral view; C. Right pedipalp, ventral view; D. Right pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure6" httpUri="https://binary.pensoft.net/fig/680585" pageId="0" pageNumber="137">6C</figureCitation>
); conductor conspicuous, robust, basally sclerotised but apically fleshy (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Mangrovia albida comb. nov., male (QM S 67277). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left pedipalp, ventral view; D. Left pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure1" httpUri="https://binary.pensoft.net/fig/680580" pageId="0" pageNumber="137">1C</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mangrovia albida comb. nov., expanded male left pedipalp (QM S 67277). A. Apico-ventral view; B. Mesal view; C. Retrolateral view; D. Dorsal view. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zse.98.82649.figure2" httpUri="https://binary.pensoft.net/fig/680581" pageId="0" pageNumber="137">2C</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mangrovia occidentalis sp. nov., male (WAM T 77397). A. Habitus, dorsal view; B. Habitus, ventral view; C. Right pedipalp, ventral view; D. Right pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure6" httpUri="https://binary.pensoft.net/fig/680585" pageId="0" pageNumber="137">6A</figureCitation>
); terminal apophysis well-developed, somewhat spoon-shaped, not sclerotised (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mangrovia albida comb. nov., expanded male left pedipalp (QM S 67277). A. Apico-ventral view; B. Mesal view; C. Retrolateral view; D. Dorsal view. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zse.98.82649.figure2" httpUri="https://binary.pensoft.net/fig/680581" pageId="0" pageNumber="137">2A, B</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mangrovia occidentalis sp. nov., expanded right left pedipalp (WAM T 77397). A. Apico-dorsal view; B. Mesal view. Scale bars: 0.1 mm (A, B)." figureDoi="10.3897/zse.98.82649.figure7" httpUri="https://binary.pensoft.net/fig/680586" pageId="0" pageNumber="137">7A, B</figureCitation>
); subterminal apophysis elongate and thin (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Mangrovia albida comb. nov., male (QM S 67277). A. Habitus, dorsal view; B. Habitus, ventral view; C. Left pedipalp, ventral view; D. Left pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure1" httpUri="https://binary.pensoft.net/fig/680580" pageId="0" pageNumber="137">1D</figureCitation>
,
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mangrovia albida comb. nov., expanded male left pedipalp (QM S 67277). A. Apico-ventral view; B. Mesal view; C. Retrolateral view; D. Dorsal view. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zse.98.82649.figure2" httpUri="https://binary.pensoft.net/fig/680581" pageId="0" pageNumber="137">2A, C</figureCitation>
,
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Mangrovia occidentalis sp. nov., male (WAM T 77397). A. Habitus, dorsal view; B. Habitus, ventral view; C. Right pedipalp, ventral view; D. Right pedipalp, dorsal view. Scale bars: 1 mm (A, B); 0.1 mm (C, D)." figureDoi="10.3897/zse.98.82649.figure6" httpUri="https://binary.pensoft.net/fig/680585" pageId="0" pageNumber="137">6D</figureCitation>
); embolus heavily sclerotized, basally inflated, then straight, thin and elongated, with subterminal short branch (Figs
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Mangrovia albida comb. nov., expanded male left pedipalp (QM S 67277). A. Apico-ventral view; B. Mesal view; C. Retrolateral view; D. Dorsal view. Scale bars: 0.1 mm (A-D)." figureDoi="10.3897/zse.98.82649.figure2" httpUri="https://binary.pensoft.net/fig/680581" pageId="0" pageNumber="137">2B, C</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mangrovia albida comb. nov. (AM KS. 98754) and M. occidentalis sp. nov. (WAM T 75326), cleared female epigynes, posterior view. A. Mangrovia albida comb. nov.; B. M. occidentalis sp. nov. Scale bar: 0.2 mm (A, B)." figureDoi="10.3897/zse.98.82649.figure4" httpUri="https://binary.pensoft.net/fig/680583" pageId="0" pageNumber="137">4C, D</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Mangrovia occidentalis sp. nov., expanded right left pedipalp (WAM T 77397). A. Apico-dorsal view; B. Mesal view. Scale bars: 0.1 mm (A, B)." figureDoi="10.3897/zse.98.82649.figure7" httpUri="https://binary.pensoft.net/fig/680586" pageId="0" pageNumber="137">7A</figureCitation>
). Epigyne base plate wider than long; scape elongate and reaching posteriorly beyond the base plate and bearing a terminal pocket; atrium rounded and very conspicuous, located almost on the border of genital area (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Mangrovia albida comb. nov., female (AM KS. 98754). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view. Scale bars: 2 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zse.98.82649.figure3" httpUri="https://binary.pensoft.net/fig/680582" pageId="0" pageNumber="137">3C-E</figureCitation>
,
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Mangrovia occidentalis sp. nov., female (WAM T 75326). A. Habitus, dorsal view; B. Habitus, ventral view; C. Epigyne, ventral view; D. Epigyne, lateral view; E. Epigyne, posterior view. Scale bars: 2 mm (A, B); 0.2 mm (C-E)." figureDoi="10.3897/zse.98.82649.figure8" httpUri="https://binary.pensoft.net/fig/680587" pageId="0" pageNumber="137">8C-E</figureCitation>
); spermathecae ovoid, separated by less than their diameter; fertilisation duct basally convoluted and attaching posteriorly to spermathecae (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Mangrovia albida comb. nov. (AM KS. 98754) and M. occidentalis sp. nov. (WAM T 75326), cleared female epigynes, posterior view. A. Mangrovia albida comb. nov.; B. M. occidentalis sp. nov. Scale bar: 0.2 mm (A, B)." figureDoi="10.3897/zse.98.82649.figure4" httpUri="https://binary.pensoft.net/fig/680583" pageId="0" pageNumber="137">4A, B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="composition">
<paragraph pageId="0" pageNumber="137">Composition.</paragraph>
<paragraph pageId="0" pageNumber="137">
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" baseAuthorityName="L. Koch" baseAuthorityYear="1871" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia albida" order="Araneae" pageId="0" pageNumber="137" rank="species" species="albida">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia albida</emphasis>
</taxonomicName>
(L. Koch, 1871) comb. nov. and
<taxonomicName authorityName="Framenau &amp; Castanheira" authorityYear="2022" family="Araneidae" genus="Mangrovia" kingdom="Animalia" lsidName="Mangrovia occidentalis" order="Araneae" pageId="0" pageNumber="137" rank="species" species="occidentalis">
<emphasis italics="true" pageId="0" pageNumber="137">Mangrovia occidentalis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="137" type="distribution">
<paragraph pageId="0" pageNumber="137">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="137">
Australia (Queensland and Western Australia) (Figs
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution records of Mangrovia albida comb. nov." figureDoi="10.3897/zse.98.82649.figure5" httpUri="https://binary.pensoft.net/fig/680584" pageId="0" pageNumber="137">5</figureCitation>
,
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Distribution records of Mangrovia occidentalis sp. nov." figureDoi="10.3897/zse.98.82649.figure9" httpUri="https://binary.pensoft.net/fig/680588" pageId="0" pageNumber="137">9</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

121
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<mods:title id="846FCEAB6AA82791C49D35B9DA816B6F">Chapter 7: Linnaean Plant Names and their Types (part E)</mods:title>
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<paragraph id="7139BCAA7069B6DCDD381C4D8B9D7EC7" pageNumber="504">
<treatmentCitation id="848A48BDA7347C74E4069DC0F5356E1E" author="Linnaeus" pageNumber="738" year="1753">
<bibCitation id="CF628FA369E4C15A25501B9F5C43C00B" author="Linnaeus" pagination="738" title="Ervum tetraspermum" volumeTitle="Species Plantarum 2" year="1753">
<taxonomicName id="FDB4BE2DC1CD1482FECF0F2298B2F93F" ID-CoL="6H326" authority="Linnaeus" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Fabaceae" genus="Ervum" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="tetraspermum">
<emphasis id="B1F41F07AC094A2AD4EC754B06E46F7D" bold="true" pageNumber="504">Ervum tetraspermum</emphasis>
Linnaeus
</taxonomicName>
,
<volumeTitle id="944F5B0DD36FDA5B26BA52789EFB7FF3">
<emphasis id="4115EFC4A14504A324BB0524A63A3904" italics="true" pageNumber="504">Species Plantarum</emphasis>
2
</volumeTitle>
: 738. 1753
</bibCitation>
</treatmentCitation>
.
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<subSubSection id="EBBAC6C1E9CC1B7DC16EA8D2910F8E40" pageNumber="504" type="distribution">
<paragraph id="0774F88DFE74015318AFE810CF20D7EC" pageNumber="504">&quot;Habitat inter Europae segetes.&quot; RCN: 5425.</paragraph>
</subSubSection>
<subSubSection id="F71B8D22A7C99F50BF038FB1281BCC0C" pageNumber="504" type="materials_examined">
<paragraph id="B7A406AEE0DDF578E7FFD0EC05B53D1D" pageNumber="504">
<materialsCitation id="A7685B65016EE13187FCF04D10445D87" ID-GBIF-Occurrence="2611986417" collectionCode="LINN" specimenCode="907.3" typeStatus="lectotype">
<typeStatus id="A3002CDB4B2F4A2187F38EBF2773ED39">Lectotype</typeStatus>
(Ali in
<emphasis id="4B74B30F952608B0343A7ECB4AF39741" italics="true" pageNumber="504">Bot. Not.</emphasis>
120: 51. 1967):
<emphasis id="76CF49B138717E5480E23E28CDEC4EEA" italics="true" pageNumber="504">Magnol</emphasis>
, Herb. Linn. No. 907.3 (
<collectionCode id="8A8EA39AAD32D1F244CDCF1446AAE231">LINN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="246E46EF561DA67C4CF7DFDDE34AB9AF" pageNumber="504" type="materials_examined">
<paragraph id="F557CD31DB2EFF4BF883A81535396757" pageNumber="504">
<typeStatus id="24471D848822CD2964017B7FC4191413">Generitype</typeStatus>
of
<taxonomicName id="6F97274E8E05FF42D81B16F083045B93" authority="Linnaeus" authorityName="Linnaeus" class="Magnoliopsida" family="Fabaceae" genus="Ervum" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="genus">
<emphasis id="F37AD35A0CF36CD79A60F8E6F1A51643" italics="true" pageNumber="504">Ervum</emphasis>
Linnaeus
</taxonomicName>
(vide Green,
<emphasis id="1AB90F49D4427A996967B1A135E5CBB8" italics="true" pageNumber="504">Prop. Brit. Bot.</emphasis>
: 175. 1929).
</paragraph>
</subSubSection>
<subSubSection id="70555A80438CFC68BE05977E803A6DCA" pageNumber="504" type="nomenclature">
<paragraph id="ACEF463104514E6681A2478ED9C279D6" pageNumber="504">
Current name:
<taxonomicName id="79DA2B8AFB1CA4CE3CB4BE7B415B60A5" ID-CoL="5BCYC" authority="(L.) Schreb." authorityName="Schreb." baseAuthorityName="Linnaeus" baseAuthorityYear="1753" class="Magnoliopsida" family="Fabaceae" genus="Vicia" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="tetrasperma">
<emphasis id="118F3342A6A7447A795385255DBF17C9" bold="true" italics="true" pageNumber="504">Vicia tetrasperma</emphasis>
(L.) Schreb.
</taxonomicName>
(
<taxonomicName id="957CF9EBA4BFF7EC6D356DE13F7152BB" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
:
<taxonomicName id="7FB24849D9EE1816F98167B73BA70575" authorityName="Rudd" authorityYear="1968" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="subFamily" subFamily="Faboideae">Faboideae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="F61A9CFB43B61CE4BA48E9856119DC62" pageNumber="504" type="discussion">
<paragraph id="DD46580378FC18506D90E50ED43F12E3" pageNumber="504">
<emphasis id="88D31F033A6974CB7E6AFFA636EE5C56" italics="true" pageNumber="504">Note:</emphasis>
Jarvis &amp; al. (
<emphasis id="F7A52355D0B93C674F7E7E6790443C62" italics="true" pageNumber="504">Regnum Veg.</emphasis>
127: 46. 1993) wrongly listed
<taxonomicName id="59101F5200BAE2FE79449F66F9E5D185" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Fabaceae" genus="Ervum" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="hirsutum">
<emphasis id="671D08E212CEF336A507C316FA1EB414" italics="true" pageNumber="504">E. hirsutum</emphasis>
</taxonomicName>
as the
<typeStatus id="0E8BBB786D541155CD2355F3A0685957">generitype</typeStatus>
, in error for
<emphasis id="B6BEE9F08E411D5DEFED5B3FCB776E26" italics="true" pageNumber="504">
<taxonomicName id="F793E1E41AAD0E1DFD95D4390FEAC81F" authorityName="Linnaeus" authorityYear="1753" class="Magnoliopsida" family="Fabaceae" genus="Ervum" higherTaxonomySource="CoL" kingdom="Plantae" order="Fabales" phylum="Tracheophyta" rank="species" species="tetraspermum">E. tetraspermum</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1002.58017" ID-GBIF-Dataset="8bedb86f-0c38-4738-b1b5-1dab270c95d2" ID-PMC="PMC7746671" ID-Pensoft-Pub="1313-2970-1002-1" ID-Pensoft-UUID="92722606C9CB50B6A63A1345E68167F9" ID-PubMed="33363429" ID-ZooBank="C9EAF9DCD87D402FAC35CC4983D2E142" ModsDocID="1313-2970-1002-1" checkinTime="1607628106955" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Garces, Jhoana M. &amp; Gattolliat, Jean-Luc" docDate="2020" docId="6691A0BB0CFF590B936416B44221754F" docLanguage="en" docName="ZooKeys 1002: 1-114" docOrigin="ZooKeys 1002" docPubDate="2020-12-10" docSource="http://dx.doi.org/10.3897/zookeys.1002.58017" docTitle="Labiobaetis delocadoi Kaltenbach &amp; Garces &amp; Gattolliat 2020, sp. nov." docType="treatment" docUuid="61631C84-8BAE-4C88-8CDE-AC051839D419" docUuidSource="ZooBank" docVersion="4" id="92722606C9CB50B6A63A1345E68167F9" lastPageNumber="1" masterDocId="92722606C9CB50B6A63A1345E68167F9" masterDocTitle="The success story of Labiobaetis Novikova &amp; Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species" masterLastPageNumber="114" masterPageNumber="1" pageNumber="1" updateTime="1668146493680" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>The success story of Labiobaetis Novikova &amp; Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kaltenbach, Thomas</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8052-0388</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garces, Jhoana M.</mods:namePart>
<mods:affiliation>Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Metro Manila, Philippines</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-12-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1002</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>114</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1002.58017</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1002.58017</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1002-1</mods:identifier>
<mods:identifier type="ZooBank">C9EAF9DCD87D402FAC35CC4983D2E142</mods:identifier>
<mods:identifier type="Pensoft-UUID">92722606C9CB50B6A63A1345E68167F9</mods:identifier>
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<treatment ID-GBIF-Taxon="170725677" LSID="urn:lsid:zoobank.org:act:61631C84-8BAE-4C88-8CDE-AC051839D419" httpUri="http://treatment.plazi.org/id/6691A0BB0CFF590B936416B44221754F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/61631C84-8BAE-4C88-8CDE-AC051839D419" authority="Kaltenbach &amp; Garces &amp; Gattolliat, 2020" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis delocadoi" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="delocadoi" status="sp. nov.">Labiobaetis delocadoi</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis delocadoi sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium j apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1">Figures 31</figureCitation>
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis delocadoi sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape." figureDoi="10.3897/zookeys.1002.58017.figure32" httpUri="https://binary.pensoft.net/fig/484697" pageId="0" pageNumber="1">, 32</figureCitation>
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Habitus, larvae, dorsal view a Labiobaetis valdezorum sp. nov. b Labiobaetis wantzeni sp. nov. c Labiobaetis baganii sp. nov. d Labiobaetis delocadoi sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure44" httpUri="https://binary.pensoft.net/fig/484709" pageId="0" pageNumber="1">, 44d</figureCitation>
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Distribution of Labiobaetis in the Philippines a Group Labiobaetis operosus b Group Labiobaetis sumigarensis c Group Labiobaetis vallus." figureDoi="10.3897/zookeys.1002.58017.figure49" httpUri="https://binary.pensoft.net/fig/484714" pageId="0" pageNumber="1">, 49b</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
Following combinations of characters: A) dorsal surface of labrum with submarginal arc of 16-19 long, clavate setae; B) labial palp segment II with a hook-like distomedial protuberance, segment III slightly pentagonal; C) left mandible without setae at apex of mola; D) fore femur rather slender, length ca. 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width, dorsal margin with ca. eleven curved, spine-like setae; E) paraproct distally expanded, with 36-39 stout, marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva</emphasis>
(Figs
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis delocadoi sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium j apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1">31</figureCitation>
,
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis delocadoi sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape." figureDoi="10.3897/zookeys.1002.58017.figure32" httpUri="https://binary.pensoft.net/fig/484697" pageId="0" pageNumber="1">32</figureCitation>
,
<figureCitation captionStart="Figure 44" captionStartId="F44" captionText="Figure 44. Habitus, larvae, dorsal view a Labiobaetis valdezorum sp. nov. b Labiobaetis wantzeni sp. nov. c Labiobaetis baganii sp. nov. d Labiobaetis delocadoi sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure44" httpUri="https://binary.pensoft.net/fig/484709" pageId="0" pageNumber="1">44d</figureCitation>
). Body length 3.2-4.6 mm. Cerci ca. 2/3 of body length. Paracercus ca. 2/3 of cerci length. Antenna approx. twice as long as head length.
</paragraph>
<caption doi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1" start="Figure 31" startId="F31">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 31.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis delocadoi" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="delocadoi">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis delocadoi</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
labrum
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
seta of arc on dorsal surface of labrum
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
right mandible
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
right prostheca
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
left mandible
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
left prostheca
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
hypopharynx and superlinguae
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
maxilla
<emphasis bold="true" pageId="0" pageNumber="1">i</emphasis>
labium
<emphasis bold="true" pageId="0" pageNumber="1">j</emphasis>
apex of paraglossa.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
.
</emphasis>
Head, thorax, and abdomen dorsally light brown, fore protoptera light brown with bright striation. Head, thorax, and abdomen ventrally light brown. Legs light brown, femur with a distomedial brown spot and a brown streak at dorsal margin, apex brown, tibia darker in distomedial area. Caudalii light brown, with a brown band at ca. 1/2 of cerci length.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
(Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis delocadoi sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape." figureDoi="10.3897/zookeys.1002.58017.figure32" httpUri="https://binary.pensoft.net/fig/484697" pageId="0" pageNumber="1">32f</figureCitation>
) with scape and pedicel subcylindrical, with very poorly developed distolateral process at scape.
</paragraph>
<caption doi="10.3897/zookeys.1002.58017.figure32" httpUri="https://binary.pensoft.net/fig/484697" pageId="0" pageNumber="1" start="Figure 32" startId="F32">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 32.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis delocadoi" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="delocadoi">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis delocadoi</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
foreleg
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
fore claw
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
tergum IV
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
gill IV
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
paraproct
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
antennal scape.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis delocadoi sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium j apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1">31a, b</figureCitation>
). Rectangular, length 0.7
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of 16-19 long, clavate setae. Ventrally with marginal row of setae composed of anterolateral long, feathered setae and medial long, bifid setae; ventral surface with ca. three short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis delocadoi sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium j apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1">31c, d</figureCitation>
). Incisor and kinetodontium fused. Incisor with five denticles; kinetodontium with three denticles, inner margin of innermost denticle with a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex, with minute denticles. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis delocadoi sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium j apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1">31e, f</figureCitation>
). Incisor and kinetodontium fused. Incisor with five denticles; kinetodontium with three denticles. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight, with minute denticles towards subtriangular process. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis delocadoi sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium j apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1">31g</figureCitation>
). Lingua shorter than superlinguae. Lingua longer than broad; medial tuft of stout setae well developed; distal half laterally expanded. Superlinguae distally straight; lateral margin rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis delocadoi sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium j apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1">31h</figureCitation>
). Galea-lacinia ventrally with two simple, apical setae under canines. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and three or four medium to long, simple setae. Maxillary palp 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of galea-lacinia; 2-segmented; palp segment II 1.6
<normalizedToken originalValue="×">x</normalizedToken>
length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment rounded, with slight excavation at inner distolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labium</emphasis>
(Fig.
<figureCitation captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Labiobaetis delocadoi sp. nov., larva morphology a labrum b seta of arc on dorsal surface of labrum c right mandible d right prostheca e left mandible f left prostheca g hypopharynx and superlinguae h maxilla i labium j apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure31" httpUri="https://binary.pensoft.net/fig/484696" pageId="0" pageNumber="1">31i, j</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with ca. seven spine-like setae increasing in length distally; apex with two long and one medium, robust, pectinate setae; outer margin with four or five long, spine-like setae; ventral surface with short, fine, simple and short, spine-like setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and a row of four medium, simple setae in anteromedial area; dorsally with a row of three long, spine-like setae near inner margin. Labial palp with segment I 0.6
<normalizedToken originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with hook-like distomedial protuberance; distomedial protuberance 0.8
<normalizedToken originalValue="×">x</normalizedToken>
width of base of segment III; ventral surface with short, fine, simple setae; dorsally with one or two long, spine-like setae near outer margin. Segment III slightly pentagonal; apex slightly pointed; length 1.3
<normalizedToken originalValue="×">x</normalizedToken>
width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hind protoptera</emphasis>
absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
(Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis delocadoi sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape." figureDoi="10.3897/zookeys.1002.58017.figure32" httpUri="https://binary.pensoft.net/fig/484697" pageId="0" pageNumber="1">32a, b</figureCitation>
). Ratio of foreleg segments 1.1:1.0:0.4:0.2.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Femur</emphasis>
.
</emphasis>
Length ca. 4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with ten or eleven long, curved, spine-like setae; length of setae 0.28
<normalizedToken originalValue="×">x</normalizedToken>
maximum width of femur. Apex rounded, with a pair of long, curved, spine-like setae and one or two short, stout setae and some fine, simple setae. Many stout, lanceolate setae scattered along ventral margin; femoral patch absent.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Tibia</emphasis>
.
</emphasis>
Dorsal margin with a partial row of short, spine-like setae and a row of fine, simple setae, on apex one longer, spine-like seta. Ventral margin with a row of short, curved, spine-like setae, on apex one longer, bipectinate, spine-like seta and a tuft of fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal 1/3.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Tarsus</emphasis>
.
</emphasis>
Dorsal margin with some fine, simple setae. Ventral margin with a row of curved, spine-like setae. Claw with one row of 11-13 denticles; distally pointed; with ca. three stripes; subapical setae absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Terga</emphasis>
(Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis delocadoi sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape." figureDoi="10.3897/zookeys.1002.58017.figure32" httpUri="https://binary.pensoft.net/fig/484697" pageId="0" pageNumber="1">32c</figureCitation>
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae and micropores. Posterior margin of tergum IV with triangular spines, wider than long.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gills</emphasis>
(Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis delocadoi sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape." figureDoi="10.3897/zookeys.1002.58017.figure32" httpUri="https://binary.pensoft.net/fig/484697" pageId="0" pageNumber="1">32d</figureCitation>
). Present on segments II-VII. Margin with small denticles intercalating fine simple setae. Tracheae partly extending from main trunk towards outer and inner margins. Gill IV as long as length of segments V and 1/3 VI combined. Gill VII little longer than length of segment VIII.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
(Fig.
<figureCitation captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Labiobaetis delocadoi sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape." figureDoi="10.3897/zookeys.1002.58017.figure32" httpUri="https://binary.pensoft.net/fig/484697" pageId="0" pageNumber="1">32e</figureCitation>
). Distally expanded, with 36-39 stout, marginal spines. Surface scattered with U-shaped scale bases and micropores. Cercotractor with numerous small, marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Dedicated to Mr. Emmanuel Delocado (Philippines), entomologist and member of the AdMU Biodiversity Laboratory.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Philippines: Cebu and Leyte (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Distribution of Labiobaetis in the Philippines a Group Labiobaetis operosus b Group Labiobaetis sumigarensis c Group Labiobaetis vallus." figureDoi="10.3897/zookeys.1002.58017.figure49" httpUri="https://binary.pensoft.net/fig/484714" pageId="0" pageNumber="1">49b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biological aspects">
<paragraph pageId="0" pageNumber="1">Biological aspects.</paragraph>
<paragraph pageId="0" pageNumber="1">The specimens were collected at altitudes of 50 m and 700 m, partly on rock surface in riffles or runs.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Philippines • larva; Cebu, Cantipla;
<geoCoordinate degrees="10" direction="north" minutes="24" orientation="latitude" precision="15" seconds="55" value="10.4152775">10°24'55&quot;N</geoCoordinate>
,
<geoCoordinate degrees="123" direction="east" minutes="49" orientation="longitude" precision="15" seconds="05" value="123.818054">123°49'05&quot;E</geoCoordinate>
; 750 m; 30.X.1995; leg. Mendoza; on slide; GenBank: MT830976; GBIFCH 00654886; PNM.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
Philippines • 2 larvae; same data as holotype; 1 on slide; GBIFCH 00592307; ZSM; 1 in alcohol; GBIFCH 00515451; AdMU • 86 larvae; Leyte, Southern Leyte, Brgy. Malico, San Francisco, Taglibas River;
<geoCoordinate degrees="10" direction="north" minutes="01" orientation="latitude" precision="15" seconds="07" value="10.018611">10°01'07&quot;N</geoCoordinate>
, 125°12'35'E; 50 m; 19-20.X.2019; leg. Garces and Pelingen; 1 on slide; MT830977; GBIFCH 00763668; AdMU; 6 in alcohol; GBIFCH 00515467; ZSM; 79 in alcohol; GBIFCH 00515494; AdMU.
</paragraph>
</subSubSection>
</treatment>
</document>