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<mods:title id="5B2491407B1C772CF9C1CC2E7FC5A4D5">The identity of catfishes identified as Mystus cavasius (Hamilton, 1822) (Teleostei: Bagridae), with a description of a new species from Myanmar.</mods:title>
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<mods:namePart id="A62ECD31B1EC0CF5A5DCF9122DE7E80B">Prosanta Chakrabarty</mods:namePart>
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[[ Genus
<taxonomicName id="B909ADDB8D29196B86F1C427415A22D7" ID-CoL="63CDY" ID-ENA="205131" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="genus">Mystus Scopoli</taxonomicName>
]]
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<taxonomicName id="DB344B68531AA344B9CC4A8045A966AE" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="genus">Mystus Scopoli, 1777</taxonomicName>
, is a diverse group of small- to medium-sized bagrid catfishes, from South Asia, with 23 of the 46 nominal species known from there (Talwar &amp; Jhingran, 1991). Despite two previous studies (Jayaram, 1954; Jayaram &amp; Anuradha Sanyal, 2003), the diversity and distribution of the group in South Asia is not well known. Phylogenetic relationships within the genus are also poorly understood, although Mo (1991) suggested there are two major lineages. Hardman (2005) recently confirmed the paraphyly of
<taxonomicName id="35DD9A85BEAB20B36F7A16FE5B9FD90F" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="genus">Mystus</taxonomicName>
.
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<paragraph id="5D5E62EFBEED16F2C2F2FFE34605711E" lastPageNumber="2" pageNumber="1">
One group of species (considered
<taxonomicName id="A0D850C82F641F3C50D60FF03E9705C8" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="genus">Mystus</taxonomicName>
s. str. by Roberts, 1994) can be easily distinguished from other congeners in having a very long adipose-fin base that spans almost the entire postdorsal distance. This group (hereafter referred to in the diagnoses as “congeners with a long-based adipose fin”), known from West, South and Southeast Asia, was revised by Roberts (1994), who considered the following eight species to be valid:
<taxonomicName id="EA5EA5B8DC8EC03C412F3B8E6AC63F59" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="species" species="pelusius">M. pelusius (Solander, 1794)</taxonomicName>
,
<taxonomicName id="CBA65D630029ABAD02C979BD0EB25EBC" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="species" species="cavasius">M. cavasius (Hamilton, 1822)</taxonomicName>
,
<taxonomicName id="5A3C97008126EF6BC2812297B86F8093" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="species" species="singaringan">M. singaringan (Bleeker, 1846)</taxonomicName>
,
<taxonomicName id="C444D5147076BAD8A7A54480FC861D32" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="species" species="bocourti">M. bocourti (Bleeker, 1864)</taxonomicName>
,
<taxonomicName id="C6ACE4557B8A99B41E55E0FB9CBEC68B" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="species" species="bleekeri">M. bleekeri (Day, 1877)</taxonomicName>
,
<taxonomicName id="6DF87AAB4D9F454BB8E4421B322B64EB" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="species" species="rufescens">M. rufescens (Vinciguerra, 1890)</taxonomicName>
,
<taxonomicName id="CA7FDF5818CD6D15713FD9A3B37B73AD" LSID-ZBK="urn:lsid:zoobank.org:act:9FDCED76-C935-4508-9964-D553CC9D3421" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Mystus rhegma Fowler 1935:102" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="species" species="rhegma">M. rhegma Fowler, 1935</taxonomicName>
, and
<taxonomicName id="813D2D48BFDC17AF535B5ABE708EFEC5" LSID-ZBK="urn:lsid:zoobank.org:act:49E74E43-647D-4A23-B85C-3D013C64CB13" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Mystus albolineatus Roberts 1994:245" order="Siluriformes" pageNumber="1" phylum="Chordata" rank="species" species="albolineatus">M. albolineatus Roberts, 1994</taxonomicName>
.
</paragraph>
<paragraph id="6219007D59D57D3DD9A4FF655C4B6234" pageNumber="2">
A considerable amount of variation in the number of rakers on the first gill arch has been observed in
<taxonomicName id="6CBA9FA308617C2E846E2A64BDC10C60" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="2" phylum="Chordata" rank="species" species="cavasius">M. cavasius</taxonomicName>
(see Roberts, 1994). We investigated patterns in this variation and recognize three distinct species, one of which is undescribed. Redescriptions of two of these species,
<taxonomicName id="ECDEF2CC59E809697571D704A183C44C" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="2" phylum="Chordata" rank="species" species="cavasius">Mystus cavasius</taxonomicName>
and
<taxonomicName id="9730D849B06E3CD318408758A44F34BB" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageNumber="2" phylum="Chordata" rank="species" species="seengtee">M. seengtee</taxonomicName>
, from northern and southern India respectively, and the description of a new species,
<taxonomicName id="CFA7A8A86640DB4920CCA233EFF8BD31" LSID-ZBK="urn:lsid:zoobank.org:act:D8F1931C-5237-4A86-A537-20F04D7CEB04" family="Bagridae" genus="Mystus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Mystus falcarius Chakrabarty &amp; Ng 2005" order="Siluriformes" pageNumber="2" phylum="Chordata" rank="species" species="falcarius">Mystus falcarius</taxonomicName>
, from Myanmar form the basis of this study.
</paragraph>
</subSubSection>
</treatment>
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<subSubSection id="79F49212210C80FEFC2E0A2470A444FE" pageId="0" pageNumber="185" type="distribution">
<paragraph id="9350425A49134D103569DEDF635242B5" pageId="0" pageNumber="185">Distribution in adjacent reg.</paragraph>
<paragraph id="B41C525EC744D5952B86AF1FFC7EAA55" pageId="0" pageNumber="185">Russia (European Russia, Ural, Siberia), Kazakhstan (Altai, Western Tien Shan).</paragraph>
</subSubSection>
<subSubSection id="399CFE1430B5013CCE95F3ACE02ED375" pageId="0" pageNumber="185" type="materials_examined">
<paragraph id="195178F7C3572ED409E0E669BF0E141B" pageId="0" pageNumber="185">Specimens examined and literature records.</paragraph>
<paragraph id="CF2BAE364BCC3CF2E50E137976A21D86" pageId="0" pageNumber="185">
<materialsCitation id="80E21B45BE58F9A6408A60833BDC2CA0" accessionNumber="KUZ 00893" collectingDate="1896-07-02" collectingDateMax="2019-07-16" collectingDateMin="1896-07-02" county="Svetloe Lake" location="Kokchetav" municipality="Borovsky forest" specimenCount="1">
<location id="FF35620661CB8057DE71415C6A96EF39" LSID="urn:lsid:plazi:treatment:6629748874085EF8B08620DA7086C272:FF35620661CB8057DE71415C6A96EF39" county="Svetloe Lake" municipality="Borovsky forest" name="Kokchetav">Kokchetav</location>
:
<emphasis id="F48302CC06438C3949E41DB4F27E8802" bold="true" pageId="0" pageNumber="185">
<location id="264A62133516B30C80C8C47A31FF1FD6" LSID="urn:lsid:plazi:treatment:6629748874085EF8B08620DA7086C272:264A62133516B30C80C8C47A31FF1FD6" county="Svetloe Lake" municipality="Borovsky forest" name="Akmola Region">Akmola Region</location>
</emphasis>
:
<location id="A0C8AF12520F67B83606B9ED5CA76160" LSID="urn:lsid:plazi:treatment:6629748874085EF8B08620DA7086C272:A0C8AF12520F67B83606B9ED5CA76160" county="Svetloe Lake" municipality="Borovsky forest" name="Burabay District">Burabay District</location>
: near
<location id="2ED1A097BEB8F62E8CCD024786DF1BE1" LSID="urn:lsid:plazi:treatment:6629748874085EF8B08620DA7086C272:2ED1A097BEB8F62E8CCD024786DF1BE1" county="Svetloe Lake" municipality="Borovsky forest" name="Karas'e Lake">
<normalizedToken id="BB33AAB42763A2C4DAD7110BC80CFB43" originalValue="Karas’e">Karas'e</normalizedToken>
Lake
</location>
,
<collectingDate id="B75EB9BDDDBD37F3AF3004B89B52E548" value="1913-07-19">19 Jul 1913</collectingDate>
,
<emphasis id="BC1C0D9CC5811EF4ED0A922478CBF11C" italics="true" pageId="0" pageNumber="185">Semenov s.n..</emphasis>
(TK!); same loc.,
<collectingDate id="B9412EA4ECEDDEAAE047F877E453EA7F" value="1973-08-10">10 Aug 1973</collectingDate>
,
<emphasis id="15B5E2F23B5BE3E78271A38860816EBA" italics="true" pageId="0" pageNumber="185">Gorchakovskiy s.n..</emphasis>
(SVER 695775!); same loc.,
<collectingDate id="5AD9DFB15ED1FF3D9508A2584768413E" value="2011-06-12">12 Jun 2011</collectingDate>
,
<emphasis id="49DC21F696384D7A183F0C13CC0020E0" italics="true" pageId="0" pageNumber="185">Kupriyanov s.n..</emphasis>
(
<accessionNumber id="46DEB32C9884D346DDE1E26C7F9C4D96" httpUri="http://www.ncbi.nlm.nih.gov/protein/KUZ00893">KUZ 00893</accessionNumber>
!); same loc., small southern swamp,
<collectingDate id="2DAE6FBAF47CE7E59846248D99EE8845" value="1973-05-26">26 May 1973</collectingDate>
,
<emphasis id="751C6BF9C76C91BF2D6720EBE7125B2F" italics="true" pageId="0" pageNumber="185">Gorchakovskiy s.n..</emphasis>
(SVER 695772!); same loc., big swamp,
<collectingDate id="C9770C24CE5C0A152CD3D344C555604E" value="1974-06-29">29 Jun 1974</collectingDate>
,
<emphasis id="67B24EAAAE7B52DFDB42580E536B3668" italics="true" pageId="0" pageNumber="185">Gorchakovskiy s.n..</emphasis>
(SVER 695770!);
<location id="5EB29D1C980B0B007573B5805F2EF1DA" LSID="urn:lsid:plazi:treatment:6629748874085EF8B08620DA7086C272:5EB29D1C980B0B007573B5805F2EF1DA" county="Svetloe Lake" municipality="Borovsky forest" name="Sinyukha Mountain">Sinyukha Mountain</location>
,
<collectingDate id="234BCD2D30F96C739E24DDAE8FF019CD" value="1912-06-17">17 Jun 1912</collectingDate>
,
<emphasis id="155CF39415A1FAED33DC1EADE3315979" italics="true" pageId="0" pageNumber="185">Semenov s.n..</emphasis>
(TK!); same loc., north slope,
<collectingDate id="9FC95D0DD8DE4FB0B5250122175D5ECE" value="1929-06-24">24 Jun 1929</collectingDate>
,
<emphasis id="EEF5187FE41FC947C8CAE32B7D2E70E2" italics="true" pageId="0" pageNumber="185">Ilyin s.n..</emphasis>
(LE!); Zolotoborskiy forestry, sq. 24,
<collectingDate id="913517A61BAFF4DEC1168AC3D4DDA315" value="1972-06-18">18 Jun 1972</collectingDate>
,
<emphasis id="35EFED8D2B316AE228AFA3390AA81A7F" italics="true" pageId="0" pageNumber="185">Gorchakovskiy s.n..</emphasis>
(SVER 695773!);
<collectingMunicipality id="06145FB532C2D7791D273C6CE052D14C">Borovsky forest</collectingMunicipality>
area, near
<collectingCounty id="FB4CF30D3D5664E8ADB1AA890947AC3D">Svetloe Lake</collectingCounty>
,
<collectingDate id="7A50B7D1F65E9C30081727D9DF1C106B" value="1973-08-09">9 Aug 1973</collectingDate>
,
<emphasis id="97DF7726BAB2FA3C03D723D19B0802F4" italics="true" pageId="0" pageNumber="185">Gorchakovskiy s.n..</emphasis>
(SVER 695774!); same loc., near Shortankulskiy peat bog,
<collectingDate id="D753EF8DA8D9C09FC1B16A860917C414" value="1978-08-05">5 Aug 1978</collectingDate>
,
<emphasis id="26ABFB1CBAC96ECE928136027DD09154" italics="true" pageId="0" pageNumber="185">Gorchakovskiy s.n..</emphasis>
(SVER 695776!); Kokchetav Upland, upper reaches of the Imanayevsky Spring, 2 Jul [presumably 1895-1910],
<emphasis id="1009D191602BD738C9C010CEEF7EC4C8" italics="true" pageId="0" pageNumber="185">Gordiagin 593</emphasis>
(LE); near the top of Ush-Tas Mount,
<collectingDate id="E82AE00664BC6EEDFF57B84620BD1700" value="1896-07-02">2 Jul 1896</collectingDate>
,
<emphasis id="4D6EAB8530C43481E613D5881A61D73B" italics="true" pageId="0" pageNumber="185">Gordyagin 1105</emphasis>
(LE!); near Burabay,
<normalizedToken id="F81DB89BC9208F927CCE1B9CF09453DE" originalValue="“Burabay”">&quot;Burabay&quot;</normalizedToken>
State National Nature Park, Barmashinskoe forestry,
<collectingDate id="7DF897B7723420AC124E6FC4A98D8B20" value="2019-07-16">16 Jul 2019</collectingDate>
,
<emphasis id="E389022076E07025D51BC9819887CF5A" italics="true" pageId="0" pageNumber="185">Kubentayev s.n..</emphasis>
(NUR!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="2EF1EB12711CB1CA1206ACE5583C6B6F" pageId="0" pageNumber="185" type="habitat">
<paragraph id="29B1FFA485BB4CD69A99DC17C0C2B5EC" pageId="0" pageNumber="185">Habitat and ecology.</paragraph>
<paragraph id="8BB85D5940299945BD0095F839432395" pageId="0" pageNumber="185">Moist pine, birch and mixed forests.</paragraph>
</subSubSection>
<subSubSection id="2679082E27BD4E6FA0B5D6DDC02652B9" pageId="0" pageNumber="185" type="phenology">
<paragraph id="E22D1FDF8F1C61A19737B522153061E1" pageId="0" pageNumber="185">Phenology.</paragraph>
<paragraph id="51896ABF53B7FDC1735611A6153F5BD5" pageId="0" pageNumber="185">Flowering in Jul-Aug; fruiting in Sep-Oct.</paragraph>
</subSubSection>
<subSubSection id="E6DCB20B2546E452605CBB944A6E65F7" pageId="0" pageNumber="185" type="conservation status">
<paragraph id="73E34CC155536B9BF448DC465FEE3543" pageId="0" pageNumber="185">Conservation status.</paragraph>
<paragraph id="C3CFFF695F1A58C282EC9D321CFBCD79" pageId="0" pageNumber="185">
Not protected. It is necessary to strengthen security measures in the territory of the Shchuchinsko-Borovskaya resort area in the Kokchetav Upland as the habitats of
<taxonomicName id="3018C94AFBF9B0EB65E8C70C4C49994C" lsidName="G. repens" pageId="0" pageNumber="185" rank="species" species="repens">
<emphasis id="CD390857505D081509A47F1CC6A2B541" italics="true" pageId="0" pageNumber="185">G. repens</emphasis>
</taxonomicName>
in this area are exposed to strong recreational influences.
</paragraph>
</subSubSection>
<subSubSection id="52720822DE179DD88EF9B393F3ED5E2E" pageId="0" pageNumber="185" type="notes">
<paragraph id="921E41868D24B71F26A4787334B9FA49" pageId="0" pageNumber="185">Notes.</paragraph>
<paragraph id="9DFCB8B1D2470A6BF88C76CCFECD782E" pageId="0" pageNumber="185">
<taxonomicName id="6FF155F5614ECB6467BD8517726A12E1" class="Liliopsida" family="Orchidaceae" genus="Goodyera" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Goodyera repens" order="Asparagales" pageId="0" pageNumber="185" phylum="Tracheophyta" rank="species" species="repens">
<emphasis id="70B410542DB4A7EFEDC2EE4B914656C8" italics="true" pageId="0" pageNumber="185">Goodyera repens</emphasis>
</taxonomicName>
(Figs
<figureCitation id="D4D57656321194AE5AB7A99BD8767DE2" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Representative taxa of Orchidaceae in the northern part of Kazakhstan A Dactylorhiza incarnata B D. fuchsii C D. sibirica D Platanthera bifolia E Cypripedium calceolus F C. macranthos G Corallorhiza trifida H Epipogium aphyllum I Hemipilia cucullata J Gymnadenia conopsea K Goodyera repens L Epipactis palustris. (Photos: A-D, G, H, J, K by S. Kubentayev; E, F, I, L by A. Kupriyanov)." figureDoi="10.3897/phytokeys.229.105457.figure3" httpUri="https://binary.pensoft.net/fig/885068" pageId="0" pageNumber="185">3K</figureCitation>
,
<figureCitation id="9F8210A0B738BACD1B67DB259FA17FF7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Schematic map of the localities of orchids in the northern part of Kazakhstan A Dactylorhiza fuchsii B D. incarnata C D. maculata D D. sibirica E D. salina F G. repens G D. umbrosa." figureDoi="10.3897/phytokeys.229.105457.figure5" httpUri="https://binary.pensoft.net/fig/885070" pageId="0" pageNumber="185">5F</figureCitation>
) is found only in a limited area within the Kokchetav FR in the studied region.
</paragraph>
</subSubSection>
</treatment>
</document>