Add updates up until 2025-03-03 13:49:32
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03E715280A023C32E4FD285EFD4D02A4.xml03E715280A053C35E4FD2BC0FD9F03B4.xml03E715280A093C39E4FD286AFDA501E9.xml03E715280A0E3C3EE4FD2875FECA0228.xml
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<figureCitation id="94F860C3FF80E853FE8539C6FEE8E4DE" box="[272,309,1735,1761]" captionStart="FIGURE 2" captionStartId="3.[151,250,1749,1773]" captionTargetBox="[158,1430,186,1717]" captionTargetId="figure-19@3.[151,1436,179,1723]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum, lateral view. A, Heterogomphus chevrolatii Burmeister, 1847; B, Heterogomphus curvicornis Prell, 1912; C, Heterogomphus dilaticollis Burmeister, 1847; D, Heterogomphus mniszechi (Thomson, 1859); E, Heterogomphus rugicollis Prell, 1912; F, Heterogomphus schoenherri Burmeister, 1847; G, Heterogomphus telamon (Burmeister, 1847); H, Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953229" httpUri="https://zenodo.org/record/14953229/files/figure.png" pageId="16" pageNumber="361">2A</figureCitation>
|
||||
;
|
||||
<figureCitation id="94F860C3FF80E853FED439C6FE58E4DE" box="[321,389,1735,1761]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="16" pageNumber="361">3A–B</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="44D92FCDFF80E853FF62380EFB34E586" pageId="16" pageNumber="361" type="reference_group">
|
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<paragraph id="0C7C7C46FF80E853FF62380EFBE8E516" blockId="16.[246,1323,1807,1978]" box="[247,1077,1807,1833]" pageId="16" pageNumber="361">
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<treatmentCitationGroup id="2CD35B68FF80E853FF62380EFBE8E516" box="[247,1077,1807,1833]" pageId="16" pageNumber="361">
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<treatmentCitation id="8D625A57FF80E853FF62380EFCF4E516" author="Burmeister, H. C. C." box="[247,809,1807,1833]" page="233" pageId="16" pageNumber="361" year="1847">
|
||||
<taxonomicName id="CBC307C5FF80E853FF62380EFCF4E516" authority="Burmeister, 1847: 233" authorityName="Burmeister" authorityPageNumber="233" authorityYear="1847" box="[247,809,1807,1833]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="species" species="chevrolatii">
|
||||
<emphasis id="3EB7A054FF80E853FF62380EFDF5E516" box="[247,552,1807,1833]" italics="true" pageId="16" pageNumber="361">Heterogomphus chevrolatii</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FDBA380EFCF4E516" author="Burmeister, H. C. C." box="[559,809,1807,1833]" pageId="16" pageNumber="361" refId="ref10951" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie, Funfter Band. Coleoptera, Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1847">Burmeister, 1847: 233</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
(original combination).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF63383EFC63E566" blockId="16.[246,1323,1807,1978]" box="[246,958,1855,1882]" pageId="16" pageNumber="361">
|
||||
<treatmentCitationGroup id="2CD35B68FF80E853FF63383EFC63E566" box="[246,958,1855,1882]" pageId="16" pageNumber="361">
|
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<treatmentCitation id="8D625A57FF80E853FF63383EFCEBE566" author="Bates, H. W." box="[246,822,1855,1882]" page="327" pageId="16" pageNumber="361" year="1888">
|
||||
<taxonomicName id="CBC307C5FF80E853FF63383EFCEBE566" authority="Bates, 1888: 327" authorityName="Bates" authorityPageNumber="327" authorityYear="1888" box="[246,822,1855,1882]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="chevrolati" subSpecies="eurytis">
|
||||
<emphasis id="3EB7A054FF80E853FF63383EFDAFE566" box="[246,626,1855,1881]" italics="true" pageId="16" pageNumber="361">Heterogomphus chevrolati eurytis</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FDEC3841FCEBE566" author="Bates, H. W." box="[633,822,1855,1882]" pageId="16" pageNumber="361" pagination="296 - 342" refId="ref10764" refString="Bates, H. W. (1888) Pectinicornia and Lamellicornia, family Dynastinae. Biologia Centrali-Americana. Insecta, Coleoptera, 2 (2), 296-342." type="journal article" year="1888">Bates, 1888: 327</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
(synonym).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF63386EFAF6E5B6" blockId="16.[246,1323,1807,1978]" box="[246,1323,1903,1930]" pageId="16" pageNumber="361">
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<treatmentCitationGroup id="2CD35B68FF80E853FF63386EFAF6E5B6" box="[246,1323,1903,1930]" pageId="16" pageNumber="361">
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<treatmentCitation id="8D625A57FF80E853FF63386EFCA4E5B5" author="Prell, H." box="[246,889,1903,1930]" page="135" pageId="16" pageNumber="361" year="1912">
|
||||
<taxonomicName id="CBC307C5FF80E853FF63386EFCA4E5B5" authority="Prell, 1912: 135" authorityName="Prell" authorityPageNumber="135" authorityYear="1912" box="[246,889,1903,1930]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="chevrolati" subSpecies="puctatissimus">
|
||||
<emphasis id="3EB7A054FF80E853FF63386EFD63E5B6" box="[246,702,1903,1929]" italics="true" pageId="16" pageNumber="361">Heterogomphus chevrolati puctatissimus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FD50386EFCA4E5B5" author="Prell, H." box="[709,889,1903,1930]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 135</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
(synonym, described as a subspecies).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF63389EFB34E586" blockId="16.[246,1323,1807,1978]" box="[246,1257,1951,1978]" pageId="16" pageNumber="361">
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<treatmentCitationGroup id="2CD35B68FF80E853FF63389EFB34E586" box="[246,1257,1951,1978]" pageId="16" pageNumber="361">
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<treatmentCitation id="8D625A57FF80E853FF63389EFCE5E585" author="Prell, H." box="[246,824,1951,1978]" page="135" pageId="16" pageNumber="361" year="1912">
|
||||
<taxonomicName id="CBC307C5FF80E853FF63389EFCE5E585" authority="Prell, 1912: 135" authorityName="Prell" authorityPageNumber="135" authorityYear="1912" box="[246,824,1951,1978]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="subSpecies" species="chevrolati" subSpecies="insignis">
|
||||
<emphasis id="3EB7A054FF80E853FF63389EFDA0E586" box="[246,637,1951,1977]" italics="true" pageId="16" pageNumber="361">Heterogomphus chevrolati insignis</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FD11389EFCE5E585" author="Prell, H." box="[644,824,1951,1978]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 135</bibRefCitation>
|
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</taxonomicName>
|
||||
</treatmentCitation>
|
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(synonym, described as a subspecies).
|
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</treatmentCitationGroup>
|
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</paragraph>
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237
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<emphasis id="3EB7A054FF80E853FF023E92FC9EE392" bold="true" box="[151,835,403,430]" pageId="16" pageNumber="361">
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Checklist of the species of
|
||||
<taxonomicName id="CBC307C5FF80E853FE753E92FD7FE391" ID-CoL="636SS" authorityName="Burmeister" authorityYear="1847" box="[480,674,403,430]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
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<emphasis id="3EB7A054FF80E853FE753E92FD7FE391" bold="true" box="[480,674,403,430]" italics="true" pageId="16" pageNumber="361">Heterogomphus</emphasis>
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</taxonomicName>
|
||||
in
|
||||
<collectingCountry id="74D43CD6FF80E853FD5C3E92FC9EE392" box="[713,835,403,429]" name="Colombia" pageId="16" pageNumber="361">Colombia</collectingCountry>
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|
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||||
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||||
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<taxonomicName id="CBC307C5FF80E853FF023EDAFDFBE3CA" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[151,550,475,501]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF023EDAFE94E3CA" box="[151,329,475,501]" italics="true" pageId="16" pageNumber="361">Heterogomphus</emphasis>
|
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<bibRefCitation id="685201B7FF80E853FECC3EDAFDFBE3CA" author="Burmeister, H. C. C." box="[345,550,475,501]" pageId="16" pageNumber="361" refId="ref10951" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie, Funfter Band. Coleoptera, Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1847">Burmeister, 1847</bibRefCitation>
|
||||
</taxonomicName>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
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<subSubSection id="44D92FCDFF80E853FDAC3EDDFE11E026" pageId="16" pageNumber="361" type="type_taxon">
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<paragraph id="0C7C7C46FF80E853FDAC3EDDFE11E026" blockId="16.[151,1436,475,537]" pageId="16" pageNumber="361">
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species:
|
||||
<taxonomicName id="CBC307C5FF80E853FD7D3EDAFB3DE3CA" authority="Burmeister, 1847" authorityName="Burmeister" authorityYear="1847" box="[744,1248,475,501]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="species" species="eteocles">
|
||||
<emphasis id="3EB7A054FF80E853FD7D3EDAFBD9E3CA" box="[744,1028,475,501]" italics="true" pageId="16" pageNumber="361">Heterogomphus eteocles</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FB813EDAFB3DE3CA" author="Burmeister, H. C. C." box="[1044,1248,475,501]" pageId="16" pageNumber="361" refId="ref10951" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie, Funfter Band. Coleoptera, Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1847">Burmeister, 1847</bibRefCitation>
|
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</taxonomicName>
|
||||
, by subsequent designation of
|
||||
<bibRefCitation id="685201B7FF80E853FEAB3EFEFE1AE026" author="Prell, H." box="[318,455,511,537]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell (1912)</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="44D92FCDFF80E853FF633D39FCCDE6FC" pageId="16" pageNumber="361" type="reference_group">
|
||||
<paragraph id="0C7C7C46FF80E853FF633D39FD75E06C" blockId="16.[246,784,568,1219]" box="[246,680,568,595]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633D39FD75E06C" authority="Burmeister, 1847: 224" authorityName="Burmeister" authorityPageNumber="224" authorityYear="1847" box="[246,680,568,595]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633D39FE75E06D" box="[246,424,568,594]" italics="true" pageId="16" pageNumber="361">Heterogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE3B3D39FD75E06C" author="Burmeister, H. C. C." box="[430,680,568,595]" pageId="16" pageNumber="361" refId="ref10951" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie, Funfter Band. Coleoptera, Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1847">Burmeister, 1847: 224</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633D68FD26E0BC" blockId="16.[246,784,568,1219]" box="[246,763,616,643]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633D68FDA6E0BC" authority="Burmeister, 1847: 219" authorityName="Burmeister" authorityPageNumber="219" authorityYear="1847" box="[246,635,616,643]" class="Insecta" family="Scarabaeidae" genus="Stypotrupes" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633D68FEA7E0BC" box="[246,378,617,643]" italics="true" pageId="16" pageNumber="361">Stypotrupes</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE143D69FDA6E0BC" author="Burmeister, H. C. C." box="[385,635,616,643]" pageId="16" pageNumber="361" refId="ref10951" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie, Funfter Band. Coleoptera, Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1847">Burmeister, 1847: 219</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633D99FD29E08D" blockId="16.[246,784,568,1219]" box="[246,756,664,691]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633D99FDAFE08D" authority="Thomson, 1859: 68" authorityName="Thomson" authorityPageNumber="68" authorityYear="1859" box="[246,626,664,691]" class="Insecta" family="Scarabaeidae" genus="Daemonoplus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633D99FE4FE08D" box="[246,402,664,690]" italics="true" pageId="16" pageNumber="361">Daemonoplus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE0D3D99FDAFE08D" author="Thomson, J." box="[408,626,664,691]" pageId="16" pageNumber="361" pagination="3 - 22" refId="ref11881" refString="Thomson, J. (1859) Essai synoptique sur la sous-tribu des scarabaeitae vrais. Arcana Naturae, 1, 3-22." type="journal article" year="1859">Thomson, 1859: 68</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(subgenus)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633DC8FD7FE0DC" blockId="16.[246,784,568,1219]" box="[246,674,713,739]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633DC8FDFFE0DC" authority="Bates, 1891: 33" authorityName="Bates" authorityPageNumber="33" authorityYear="1891" box="[246,546,713,739]" class="Insecta" family="Scarabaeidae" genus="Baryxenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633DC8FEB1E0DD" box="[246,364,713,738]" italics="true" pageId="16" pageNumber="361">Baryxenus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FEE63DC8FDFFE0DC" author="Bates, H. W." box="[371,546,713,739]" pageId="16" pageNumber="361" pagination="7 - 88" refId="ref10796" refString="Bates, H. W. (1891) Coleoptera. In: Whymper, E. (Ed.), Supplementary Appendix to Travels Amongst the Great Andes of the Equador. John Murray, London, pp. 7-88." type="book chapter" year="1891">Bates, 1891: 33</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633DF9FD34E12C" blockId="16.[246,784,568,1219]" box="[246,745,760,787]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633DF9FDB5E12C" authority="Prell, 1912: 53" authorityName="Prell" authorityPageNumber="53" authorityYear="1912" box="[246,616,760,787]" class="Insecta" family="Scarabaeidae" genus="Syneterogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633DF9FE61E12D" box="[246,444,760,786]" italics="true" pageId="16" pageNumber="361">Syneterogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE563DF9FDB5E12C" author="Prell, H." box="[451,616,760,787]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 53</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633C29FD21E17C" blockId="16.[246,784,568,1219]" box="[246,764,808,835]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633C29FDA6E17C" authority="Prell, 1912: 103" authorityName="Prell" authorityPageNumber="103" authorityYear="1912" box="[246,635,808,835]" class="Insecta" family="Scarabaeidae" genus="Ortheterogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633C29FE14E17D" box="[246,457,808,834]" italics="true" pageId="16" pageNumber="361">Ortheterogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE5A3C29FDA6E17C" author="Prell, H." box="[463,635,808,835]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912:103</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633C59FCD7E14C" blockId="16.[246,784,568,1219]" box="[246,778,856,883]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633C59FD54E14C" authority="Prell, 1912: 105" authorityName="Prell" authorityPageNumber="105" authorityYear="1912" box="[246,649,856,883]" class="Insecta" family="Scarabaeidae" genus="Heterogomphidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633C59FE12E14D" box="[246,463,856,882]" italics="true" pageId="16" pageNumber="361">Heterogomphidium</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE403C59FD54E14C" author="Prell, H." box="[469,649,856,883]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 105</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633C89FD31E19C" blockId="16.[246,784,568,1219]" box="[246,748,904,931]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633C89FDB6E19C" authority="Prell, 1912: 123" authorityName="Prell" authorityPageNumber="123" authorityYear="1912" box="[246,619,904,931]" class="Insecta" family="Scarabaeidae" genus="Hoplitogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633C89FE6CE19D" box="[246,433,904,930]" italics="true" pageId="16" pageNumber="361">Hoplitogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE223C89FDB6E19C" author="Prell, H." box="[439,619,904,931]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 123</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633CB9FD25E1EC" blockId="16.[246,784,568,1219]" box="[246,760,952,979]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633CB9FDA5E1EC" authority="Prell, 1912: 129" authorityName="Prell" authorityPageNumber="129" authorityYear="1912" box="[246,632,952,979]" class="Insecta" family="Scarabaeidae" genus="Dineterogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633CB9FE63E1ED" box="[246,446,952,978]" italics="true" pageId="16" pageNumber="361">Dineterogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE513CB9FDA5E1EC" author="Prell, H." box="[452,632,952,979]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 129</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633CE9FD27E63C" blockId="16.[246,784,568,1219]" box="[246,762,1000,1027]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633CE9FDA4E63C" authority="Prell, 1912: 141" authorityName="Prell" authorityPageNumber="141" authorityYear="1912" box="[246,633,1000,1027]" class="Insecta" family="Scarabaeidae" genus="Psileterogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633CE9FE62E63D" box="[246,447,1000,1026]" italics="true" pageId="16" pageNumber="361">Psileterogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE533CE9FDA4E63C" author="Prell, H." box="[454,633,1000,1027]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 141</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633B19FD0CE60C" blockId="16.[246,784,568,1219]" box="[246,721,1048,1075]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633B19FD8DE60C" authority="Prell, 1912: 148" authorityName="Prell" authorityPageNumber="148" authorityYear="1912" box="[246,592,1048,1075]" class="Insecta" family="Scarabaeidae" genus="Neobaryxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633B19FE4BE60D" box="[246,406,1048,1074]" italics="true" pageId="16" pageNumber="361">Neobaryxenus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE083B19FD8DE60C" author="Prell, H." box="[413,592,1048,1075]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 148</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633B49FD3AE65C" blockId="16.[246,784,568,1219]" box="[246,743,1096,1123]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633B49FDBBE65C" authority="Prell, 1912: 149" authorityName="Prell" authorityPageNumber="149" authorityYear="1912" box="[246,614,1096,1123]" class="Insecta" family="Scarabaeidae" genus="Anoplogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633B49FE71E65D" box="[246,428,1096,1122]" italics="true" pageId="16" pageNumber="361">Anoplogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE263B49FDBBE65C" author="Prell, H." box="[435,614,1096,1123]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 149</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633B79FCD8E6AC" blockId="16.[246,784,568,1219]" box="[246,773,1144,1171]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633B79FD59E6AC" authority="Prell, 1912: 156" authorityName="Prell" authorityPageNumber="156" authorityYear="1912" box="[246,644,1144,1171]" class="Insecta" family="Scarabaeidae" genus="Notheterogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633B79FE17E6AD" box="[246,458,1144,1170]" italics="true" pageId="16" pageNumber="361">Notheterogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE443B79FD59E6AC" author="Prell, H." box="[465,644,1144,1171]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 156</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF80E853FF633BA9FCCDE6FC" blockId="16.[246,784,568,1219]" box="[246,784,1192,1219]" pageId="16" pageNumber="361">
|
||||
<taxonomicName id="CBC307C5FF80E853FF633BA9FD52E6FD" authority="Prell, 1912: 161" authorityName="Prell" authorityPageNumber="161" authorityYear="1912" box="[246,655,1192,1219]" class="Insecta" family="Scarabaeidae" genus="Tracheterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FF633BA9FE08E6FD" box="[246,469,1192,1218]" italics="true" pageId="16" pageNumber="361">Tracheterogomphus</emphasis>
|
||||
<bibRefCitation id="685201B7FF80E853FE493BA9FD52E6FD" author="Prell, H." box="[476,655,1192,1219]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 161</bibRefCitation>
|
||||
</taxonomicName>
|
||||
(synonym)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="44D92FCDFF80E853FF023BE6FE95E752" pageId="16" pageNumber="361" type="discussion">
|
||||
<paragraph id="0C7C7C46FF80E853FF023BE6FE95E752" blockId="16.[151,1436,1255,1389]" pageId="16" pageNumber="361">
|
||||
Subgenus note. The subgenus
|
||||
<taxonomicName id="CBC307C5FF80E853FE7F3BE6FD5BE73E" authorityName="Thomson" authorityYear="1859" box="[490,646,1255,1281]" class="Insecta" family="Scarabaeidae" genus="Daemonoplus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FE7F3BE6FD5BE73E" box="[490,646,1255,1281]" italics="true" pageId="16" pageNumber="361">Daemonoplus</emphasis>
|
||||
</taxonomicName>
|
||||
is valid and contains two species with males having three horns on the pronotum:
|
||||
<taxonomicName id="CBC307C5FF80E853FE873A0DFDB4E719" authority="(Thomson, 1859)" baseAuthorityName="Thomson" baseAuthorityYear="1859" box="[274,617,1291,1318]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="species" species="mniszechi">
|
||||
<emphasis id="3EB7A054FF80E853FE873A0DFE7CE71A" box="[274,417,1291,1317]" italics="true" pageId="16" pageNumber="361">H. mniszechi</emphasis>
|
||||
(
|
||||
<bibRefCitation id="685201B7FF80E853FE3A3A0AFDBCE719" author="Thomson, J." box="[431,609,1291,1318]" pageId="16" pageNumber="361" pagination="3 - 22" refId="ref11881" refString="Thomson, J. (1859) Essai synoptique sur la sous-tribu des scarabaeitae vrais. Arcana Naturae, 1, 3-22." type="journal article" year="1859">Thomson, 1859</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
and
|
||||
<taxonomicName id="CBC307C5FF80E853FD0B3A0DFC7EE71A" authority="(Kolbe, 1906)" baseAuthorityName="Kolbe" baseAuthorityYear="1906" box="[670,931,1291,1317]" class="Insecta" family="Scarabaeidae" genus="Hoplitogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="species" species="flohri">
|
||||
<emphasis id="3EB7A054FF80E853FD0B3A0DFD20E71A" box="[670,765,1291,1317]" italics="true" pageId="16" pageNumber="361">H. flohri</emphasis>
|
||||
(
|
||||
<bibRefCitation id="685201B7FF80E853FC993A0AFC46E71A" author="Kolbe, H." box="[780,923,1291,1317]" pageId="16" pageNumber="361" pagination="265 - 275" refId="ref11371" refString="Kolbe, H. (1906) Die Dynastiden-Gattung Daemonoplus. Stettiner Entomologische Zeitung, 67, 265-275." type="journal article" year="1906">Kolbe, 1906</bibRefCitation>
|
||||
)
|
||||
</taxonomicName>
|
||||
. The
|
||||
<typeStatus id="D378C2E4FF80E853FC4B3A0DFBD3E719" box="[990,1038,1292,1318]" pageId="16" pageNumber="361">type</typeStatus>
|
||||
species is
|
||||
<taxonomicName id="CBC307C5FF80E853FB113A0AFE94E775" authority="Thomson, 1858" authorityName="Thomson" authorityYear="1858" class="Insecta" family="Scarabaeidae" genus="Daemonoplus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="species" species="mniszechii">
|
||||
<emphasis id="3EB7A054FF80E853FB113A0AFA41E71A" box="[1156,1436,1291,1317]" italics="true" pageId="16" pageNumber="361">Daemonoplus mniszechii</emphasis>
|
||||
Thomson, 1858
|
||||
</taxonomicName>
|
||||
by monotypy (
|
||||
<bibRefCitation id="685201B7FF80E853FE6D3A2EFD72E775" author="Thomson, J." box="[504,687,1327,1354]" pageId="16" pageNumber="361" pagination="3 - 22" refId="ref11881" refString="Thomson, J. (1859) Essai synoptique sur la sous-tribu des scarabaeitae vrais. Arcana Naturae, 1, 3-22." type="journal article" year="1859">Thomson, 1859</bibRefCitation>
|
||||
).
|
||||
<taxonomicName id="CBC307C5FF80E853FD503A2EFCBCE776" authorityName="Thomson" authorityYear="1859" box="[709,865,1327,1353]" class="Insecta" family="Scarabaeidae" genus="Daemonoplus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FD503A2EFCBCE776" box="[709,865,1327,1353]" italics="true" pageId="16" pageNumber="361">Daemonoplus</emphasis>
|
||||
</taxonomicName>
|
||||
was reclassified to a subgenus of
|
||||
<taxonomicName id="CBC307C5FF80E853FB7F3A2EFA41E776" authorityName="Burmeister" authorityYear="1847" box="[1258,1436,1327,1353]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="361" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF80E853FB7F3A2EFA41E776" box="[1258,1436,1327,1353]" italics="true" pageId="16" pageNumber="361">Heterogomphus</emphasis>
|
||||
</taxonomicName>
|
||||
by
|
||||
<bibRefCitation id="685201B7FF80E853FF2F3A52FE99E752" author="Prell, H." box="[186,324,1363,1389]" pageId="16" pageNumber="361" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell (1912)</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
109
data/84/6A/CD/846ACD50FF81E852FF0239EAFCB4E5AC.xml
Normal file
109
data/84/6A/CD/846ACD50FF81E852FF0239EAFCB4E5AC.xml
Normal file
|
|
@ -0,0 +1,109 @@
|
|||
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<mods:title id="B11083F4D3DCAE93127050C7CC589FF0">A review of the genus Heterogomphus Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) from Colombia</mods:title>
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<mods:namePart id="970C8A20BB2536012000AFEA609B9FD3">Neita Moreno, Jhon C.</mods:namePart>
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<mods:affiliation id="565E70A63FEFF6E8AE3011B7C192FB31">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Claustro de San Agustín, Villa de Leyva, Boyacá, Colombia</mods:affiliation>
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</mods:role>
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<mods:namePart id="2C4252E78EF020248741B0A25E052000">Clavijo-Bustos, Julián</mods:namePart>
|
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<mods:affiliation id="38D9471640FB82561EE3D9A2A9AEAE6D">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Investigación de Recursos Biológicos Alexander von Humboldt, Claustro de San Agustín, Villa de Leyva, Boyacá, Colombia</mods:affiliation>
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<mods:namePart id="90942B40EE30C2C25BE9317737835E12">Ratcliffe, Brett C.</mods:namePart>
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<mods:affiliation id="CE48DB7071E6DCCE497028DDB6B7A10E">Systematics Research Collections, W- 436 Nebraska Hall, University of Nebraska State Museum, Lincoln, Nebraska, 68588 - 0514, United States of America</mods:affiliation>
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<mods:title id="A02E4C0BB43CE3AC5F4603C4FE92FD5A">Zootaxa</mods:title>
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<mods:part id="07C464CC06B664D784750D6B7F8556EB">
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<mods:date id="3D373E3CEBB5D52F34793ADD0577054F">2025</mods:date>
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<mods:number id="557E01EDEB6037F5DC0F5BC56B408513">2025-02-24</mods:number>
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|
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|
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<subSubSection id="44D92FCDFF81E852FF0239EAFD6CE53A" box="[151,689,1771,1798]" pageId="17" pageNumber="362" type="nomenclature">
|
||||
<paragraph id="0C7C7C46FF81E852FF0239EAFD6CE53A" blockId="17.[151,689,1771,1833]" box="[151,689,1771,1798]" pageId="17" pageNumber="362">
|
||||
<heading id="5734CB2AFF81E852FF0239EAFD6CE53A" bold="true" box="[151,689,1771,1798]" fontSize="11" level="1" pageId="17" pageNumber="362" reason="1">
|
||||
<taxonomicName id="CBC307C5FF81E852FF0239EAFD6CE53A" ID-CoL="7ZNJF" authority="(Burmeister, 1847)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[151,689,1771,1798]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="362" phylum="Arthropoda" rank="species" species="telamon">
|
||||
<emphasis id="3EB7A054FF81E852FF0239EAFD6CE53A" bold="true" box="[151,689,1771,1798]" pageId="17" pageNumber="362">
|
||||
<emphasis id="3EB7A054FF81E852FF0239EAFE1FE539" bold="true" box="[151,450,1771,1798]" italics="true" pageId="17" pageNumber="362">Heterogomphus telamon</emphasis>
|
||||
(
|
||||
<bibRefCitation id="685201B7FF81E852FE4439EAFD74E53A" author="Burmeister, H. C. C." box="[465,681,1771,1797]" pageId="17" pageNumber="362" refId="ref10951" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie, Funfter Band. Coleoptera, Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1847">Burmeister, 1847</bibRefCitation>
|
||||
)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</heading>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="44D92FCDFF81E852FF02380EFE52E516" box="[151,399,1807,1833]" pageId="17" pageNumber="362" type="description">
|
||||
<paragraph id="0C7C7C46FF81E852FF02380EFE52E516" blockId="17.[151,689,1771,1833]" box="[151,399,1807,1833]" pageId="17" pageNumber="362">
|
||||
(
|
||||
<figureCitation id="94F860C3FF81E852FF0A380EFED9E516" box="[159,260,1807,1833]" captionStart="FIGURE 1" captionStartId="2.[151,250,1767,1791]" captionTargetBox="[159,1423,559,1735]" captionTargetId="figure-140@2.[151,1436,552,1741]" captionTargetPageId="2" captionText="FIGURE 1. Habitus, dorsal view. A, Heterogomphus chevrolatii Burmeister, 1847; B, Heterogomphus curvicornis Prell, 1912; C, Heterogomphus dilaticollis Burmeister, 1847; D, Heterogomphus mniszechi (Thomson, 1859); E, Heterogomphus rugicollis Prell, 1912; F, Heterogomphus schoenherri Burmeister, 1847; G, Heterogomphus telamon (Burmeister, 1847); H, Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953225" httpUri="https://zenodo.org/record/14953225/files/figure.png" pageId="17" pageNumber="362">Figs. 1G</figureCitation>
|
||||
;
|
||||
<figureCitation id="94F860C3FF81E852FE85380EFEEBE516" box="[272,310,1807,1833]" captionStart="FIGURE 2" captionStartId="3.[151,250,1749,1773]" captionTargetBox="[158,1430,186,1717]" captionTargetId="figure-19@3.[151,1436,179,1723]" captionTargetPageId="3" captionText="FIGURE 2. Pronotum, lateral view. A, Heterogomphus chevrolatii Burmeister, 1847; B, Heterogomphus curvicornis Prell, 1912; C, Heterogomphus dilaticollis Burmeister, 1847; D, Heterogomphus mniszechi (Thomson, 1859); E, Heterogomphus rugicollis Prell, 1912; F, Heterogomphus schoenherri Burmeister, 1847; G, Heterogomphus telamon (Burmeister, 1847); H, Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953229" httpUri="https://zenodo.org/record/14953229/files/figure.png" pageId="17" pageNumber="362">2G</figureCitation>
|
||||
;
|
||||
<figureCitation id="94F860C3FF81E852FED4380EFE5BE516" box="[321,390,1807,1833]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="17" pageNumber="362">3N–O</figureCitation>
|
||||
)
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="44D92FCDFF81E852FF633848FCB4E5AC" pageId="17" pageNumber="362" type="reference_group">
|
||||
<paragraph id="0C7C7C46FF81E852FF633848FC35E55D" blockId="17.[246,1000,1864,1939]" box="[246,1000,1864,1891]" pageId="17" pageNumber="362">
|
||||
<treatmentCitationGroup id="2CD35B68FF81E852FF633848FC35E55D" box="[246,1000,1864,1891]" pageId="17" pageNumber="362">
|
||||
<treatmentCitation id="8D625A57FF81E852FF633848FD01E55C" author="Burmeister, H. C. C." box="[246,732,1864,1891]" page="210" pageId="17" pageNumber="362" year="1847">
|
||||
<taxonomicName id="CBC307C5FF81E852FF633848FD01E55C" ID-CoL="7Z55K" authority="Burmeister, 1847: 210" authorityName="Burmeister" authorityPageNumber="210" authorityYear="1847" box="[246,732,1864,1891]" class="Insecta" family="Scarabaeidae" genus="Stypotrupes" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="362" phylum="Arthropoda" rank="species" species="telamon">
|
||||
<emphasis id="3EB7A054FF81E852FF633848FE06E55D" box="[246,475,1864,1891]" italics="true" pageId="17" pageNumber="362">Stypotrupes telamon</emphasis>
|
||||
<bibRefCitation id="685201B7FF81E852FE773849FD01E55C" author="Burmeister, H. C. C." box="[482,732,1864,1891]" pageId="17" pageNumber="362" refId="ref10951" refString="Burmeister, H. C. C. (1847) Handbuch der Entomologie, Funfter Band. Coleoptera, Lamellicornia Xylophila et Pectinicornia. T. C. F. Enslin, Berlin, 584 pp." type="book" year="1847">Burmeister, 1847: 210</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
(original combination).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF81E852FF633879FCB4E5AC" blockId="17.[246,1000,1864,1939]" box="[246,873,1912,1939]" pageId="17" pageNumber="362">
|
||||
<treatmentCitationGroup id="2CD35B68FF81E852FF633879FCB4E5AC" box="[246,873,1912,1939]" pageId="17" pageNumber="362">
|
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<treatmentCitation id="8D625A57FF81E852FF633879FD3CE5AC" author="Prell, H." box="[246,737,1912,1939]" page="55" pageId="17" pageNumber="362" year="1912">
|
||||
<taxonomicName id="CBC307C5FF81E852FF633879FD3CE5AC" ID-CoL="7WLLJ" authority="Prell, 1912: 55" authorityName="Prell" authorityPageNumber="55" authorityYear="1912" box="[246,737,1912,1939]" class="Insecta" family="Scarabaeidae" genus="Syneterogomphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="362" phylum="Arthropoda" rank="species" species="sternbergi">
|
||||
<emphasis id="3EB7A054FF81E852FF633879FDE8E5AD" box="[246,565,1912,1938]" italics="true" pageId="17" pageNumber="362">Syneterogomphus sternbergi</emphasis>
|
||||
<bibRefCitation id="685201B7FF81E852FDA93879FD3CE5AC" author="Prell, H." box="[572,737,1912,1939]" pageId="17" pageNumber="362" pagination="93 - 176" refId="ref11705" refString="Prell, H. (1912) Revision des Dynastinen-Gattung Heterogomphus Burm. Memoirs de la Societe Entomologique de Belgique, 20, 93-176." type="journal article" year="1912">Prell, 1912: 55</bibRefCitation>
|
||||
</taxonomicName>
|
||||
</treatmentCitation>
|
||||
(synonym).
|
||||
</treatmentCitationGroup>
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,59 +1,59 @@
|
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Synonymy note. After reviewing the original description of
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Key to the adult females of
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species from
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Modified from
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<bibRefCitation id="685201B7FF97E844FED13BA9FDE4E6FC" author="Ratcliffe, B. C. & Cave, R. D. & Paucar-Cabrera, A." box="[324,569,1192,1219]" pageId="7" pageNumber="352" pagination="1 - 586" refId="ref11732" refString="Ratcliffe, B. C., Cave, R. D. & Paucar-Cabrera, A. (2020) The dynastine scarab beetles of Ecuador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 32, 1-586." type="journal article" year="2020">
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Ratcliffe
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. (2020)
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.
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</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023A29FA41E764" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023A29FA41E702" blockId="7.[151,1437,1261,1841]" box="[151,1436,1320,1342]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023A29FA41E702" box="[151,1436,1320,1342]" pageId="7" pageNumber="352">2. Elytra with long, moderately dense to dense setae............................................................ 3</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023A44FA41E764" blockId="7.[151,1437,1261,1841]" box="[151,1436,1349,1371]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023A44FA41E764" box="[151,1436,1349,1371]" pageId="7" pageNumber="352">2´. Elytra glabrous or, at most, with short, sparse setae confined apex............................................... 4</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023A62FA41E7EF" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023A62FA40E7A9" blockId="7.[151,1437,1261,1841]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023A62FA40E7A9" pageId="7" pageNumber="352">
|
||||
3. Elytra rugopunctate (
|
||||
<figureCitation id="94F860C3FF97E844FE3A3A62FE23E747" box="[431,510,1379,1401]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="7" pageNumber="352">Fig. 7H</figureCitation>
|
||||
); mesotibial apex truncate with 4–6 stout, short spinules (
|
||||
<figureCitation id="94F860C3FF97E844FBA43A62FBA3E747" box="[1073,1150,1379,1401]" captionStart="FIGURE 8" captionStartId="11.[151,250,1836,1860]" captionTargetBox="[286,1302,815,1805]" captionTargetId="figure-330@11.[281,1307,810,1810]" captionTargetPageId="11" captionText="FIGURE 8. Mesotibiae and metatibiae (A–H).A, Mesotibia of Heterogomphus carayoni Dechambre, 1986 (male); B, Metatibia of Heterogomphus carayoni (male); C, Mesotibia of Heterogomphus carayoni (female); D, Metatibia of Heterogomphus carayoni (female); E, Mesotibia of Heterogomphus kogui new species (male holotype); F, Metatibia of Heterogomphus kogui (male holotype); G, Mesotibia of Heterogomphus kogui (female paratype); H, Metatibia of Heterogomphus kogui (female paratype). Parameres of Heterogomphus species (I–L): I, Heterogomphus carayoni (dorsal view) and J, (lateral view); K, Heterogomphus kogui (holotype) (dorsal view) and L (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.14953247" httpUri="https://zenodo.org/record/14953247/files/figure.png" pageId="7" pageNumber="352">Fig. 8G</figureCitation>
|
||||
); metatibia truncate with 5–6 stout, short spinules (
|
||||
<figureCitation id="94F860C3FF97E844FE3B3A81FE24E7AA" box="[430,505,1408,1430]" captionStart="FIGURE 8" captionStartId="11.[151,250,1836,1860]" captionTargetBox="[286,1302,815,1805]" captionTargetId="figure-330@11.[281,1307,810,1810]" captionTargetPageId="11" captionText="FIGURE 8. Mesotibiae and metatibiae (A–H).A, Mesotibia of Heterogomphus carayoni Dechambre, 1986 (male); B, Metatibia of Heterogomphus carayoni (male); C, Mesotibia of Heterogomphus carayoni (female); D, Metatibia of Heterogomphus carayoni (female); E, Mesotibia of Heterogomphus kogui new species (male holotype); F, Metatibia of Heterogomphus kogui (male holotype); G, Mesotibia of Heterogomphus kogui (female paratype); H, Metatibia of Heterogomphus kogui (female paratype). Parameres of Heterogomphus species (I–L): I, Heterogomphus carayoni (dorsal view) and J, (lateral view); K, Heterogomphus kogui (holotype) (dorsal view) and L (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.14953247" httpUri="https://zenodo.org/record/14953247/files/figure.png" pageId="7" pageNumber="352">Fig. 8H</figureCitation>
|
||||
)......................
|
||||
<emphasis id="3EB7A054FF97E844FC9A3A81FA40E7A9" bold="true" box="[783,1437,1408,1430]" pageId="7" pageNumber="352">
|
||||
<taxonomicName id="CBC307C5FF97E844FC9A3A81FAC1E7A9" authority="Neita Moreno, Clavijo-Bustos, & Ratcliffe" authorityName="Neita Moreno, Clavijo-Bustos, & Ratcliffe" authorityYear="2025" box="[783,1308,1408,1430]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="kogui" status="new species">
|
||||
<emphasis id="3EB7A054FF97E844FC9A3A81FCBBE7A9" bold="true" box="[783,870,1408,1430]" italics="true" pageId="7" pageNumber="352">H. kogui</emphasis>
|
||||
Neita Moreno, Clavijo-Bustos, & Ratcliffe
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="25841D2FFF97E844FAB33A81FA40E7A9" box="[1318,1437,1408,1430]" pageId="7" pageNumber="352" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023A9CFA41E7EF" blockId="7.[151,1437,1261,1841]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023A9CFA41E7EF" pageId="7" pageNumber="352">
|
||||
3´. Elytral punctures ocellate (
|
||||
<figureCitation id="94F860C3FF97E844FE753A9CFDF1E78C" box="[480,556,1437,1459]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="7" pageNumber="352">Fig. 7G</figureCitation>
|
||||
); mesotibial apex with one large tooth (
|
||||
<figureCitation id="94F860C3FF97E844FC0B3A9CFC34E78C" box="[926,1001,1437,1459]" captionStart="FIGURE 8" captionStartId="11.[151,250,1836,1860]" captionTargetBox="[286,1302,815,1805]" captionTargetId="figure-330@11.[281,1307,810,1810]" captionTargetPageId="11" captionText="FIGURE 8. Mesotibiae and metatibiae (A–H).A, Mesotibia of Heterogomphus carayoni Dechambre, 1986 (male); B, Metatibia of Heterogomphus carayoni (male); C, Mesotibia of Heterogomphus carayoni (female); D, Metatibia of Heterogomphus carayoni (female); E, Mesotibia of Heterogomphus kogui new species (male holotype); F, Metatibia of Heterogomphus kogui (male holotype); G, Mesotibia of Heterogomphus kogui (female paratype); H, Metatibia of Heterogomphus kogui (female paratype). Parameres of Heterogomphus species (I–L): I, Heterogomphus carayoni (dorsal view) and J, (lateral view); K, Heterogomphus kogui (holotype) (dorsal view) and L (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.14953247" httpUri="https://zenodo.org/record/14953247/files/figure.png" pageId="7" pageNumber="352">Fig. 8C</figureCitation>
|
||||
; metatibial apex truncate with one moderately short tooth (
|
||||
<figureCitation id="94F860C3FF97E844FEC33ABAFE7CE7EF" box="[342,417,1467,1489]" captionStart="FIGURE 8" captionStartId="11.[151,250,1836,1860]" captionTargetBox="[286,1302,815,1805]" captionTargetId="figure-330@11.[281,1307,810,1810]" captionTargetPageId="11" captionText="FIGURE 8. Mesotibiae and metatibiae (A–H).A, Mesotibia of Heterogomphus carayoni Dechambre, 1986 (male); B, Metatibia of Heterogomphus carayoni (male); C, Mesotibia of Heterogomphus carayoni (female); D, Metatibia of Heterogomphus carayoni (female); E, Mesotibia of Heterogomphus kogui new species (male holotype); F, Metatibia of Heterogomphus kogui (male holotype); G, Mesotibia of Heterogomphus kogui (female paratype); H, Metatibia of Heterogomphus kogui (female paratype). Parameres of Heterogomphus species (I–L): I, Heterogomphus carayoni (dorsal view) and J, (lateral view); K, Heterogomphus kogui (holotype) (dorsal view) and L (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.14953247" httpUri="https://zenodo.org/record/14953247/files/figure.png" pageId="7" pageNumber="352">Fig. 8D</figureCitation>
|
||||
)................................................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB3E3ABAFA41E7EF" ID-CoL="7ZNH9" authority="Dechambre" authorityName="Dechambre" authorityYear="1986" box="[1195,1436,1467,1488]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="carayoni">
|
||||
<emphasis id="3EB7A054FF97E844FB3E3ABAFA41E7EF" bold="true" box="[1195,1436,1467,1488]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB3E3ABAFAFDE7EF" bold="true" box="[1195,1312,1467,1488]" italics="true" pageId="7" pageNumber="352">H. carayoni</emphasis>
|
||||
Dechambre
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023AD9FA41E434" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023AD9FA41E7D1" blockId="7.[151,1437,1261,1841]" box="[151,1436,1496,1518]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023AD9FA41E7D1" box="[151,1436,1496,1518]" pageId="7" pageNumber="352">
|
||||
4. Clypeal apex truncate or shallowly emarginate........................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB183AD9FA41E7D1" ID-CoL="7ZNJD" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1165,1436,1496,1518]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="schoenherri">
|
||||
<emphasis id="3EB7A054FF97E844FB183AD9FA41E7D1" bold="true" box="[1165,1436,1496,1518]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB183AD9FAFEE7D1" bold="true" box="[1165,1315,1496,1518]" italics="true" pageId="7" pageNumber="352">H. schoenherri</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023AF4FA41E434" blockId="7.[151,1437,1261,1841]" box="[151,1436,1525,1547]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023AF4FA41E434" box="[151,1436,1525,1547]" pageId="7" pageNumber="352">4´. Clypeal apex deeply emarginate.......................................................................... 5</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023912FA41E47A" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023912FA41E417" blockId="7.[151,1437,1261,1841]" box="[151,1436,1554,1577]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023912FA41E417" box="[151,1436,1554,1577]" pageId="7" pageNumber="352">
|
||||
5. Mesotibial apex with large tooth; elytral punctures large, dense.............................
|
||||
<taxonomicName id="CBC307C5FF97E844FB303912FA41E417" ID-CoL="3L9MW" authority="(Thomson)" baseAuthorityName="Thomson" baseAuthorityYear="1859" box="[1189,1436,1554,1576]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="mniszechi">
|
||||
<emphasis id="3EB7A054FF97E844FB303912FA41E417" bold="true" box="[1189,1436,1554,1576]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB303912FAFAE417" bold="true" box="[1189,1319,1554,1576]" italics="true" pageId="7" pageNumber="352">H. mniszechi</emphasis>
|
||||
(Thomson)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023931FA41E47A" blockId="7.[151,1437,1261,1841]" box="[151,1436,1584,1606]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023931FA41E47A" box="[151,1436,1584,1606]" pageId="7" pageNumber="352">
|
||||
5´. Mesotibial apex subtruncate, crenulate; elytral punctures sparse to moderate in density................
|
||||
<taxonomicName id="CBC307C5FF97E844FB7C3931FA41E47A" ID-CoL="7ZNJB" authority="Prell" authorityName="Prell" authorityYear="1912" box="[1257,1436,1584,1606]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="rugicollis">
|
||||
<emphasis id="3EB7A054FF97E844FB7C3931FA41E47A" bold="true" box="[1257,1436,1584,1606]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB7C3931FAB8E479" bold="true" box="[1257,1381,1584,1606]" italics="true" pageId="7" pageNumber="352">H. rugicollis</emphasis>
|
||||
Prell
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF02394CFA41E4BF" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF02394CFA41E45C" blockId="7.[151,1437,1261,1841]" box="[151,1436,1613,1635]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF02394CFA41E45C" box="[151,1436,1613,1635]" pageId="7" pageNumber="352">6. Pygidium on basal half transversely rugose or densely punctate; clypeal apex broad................................. 7</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF02396AFA41E4BF" blockId="7.[151,1437,1261,1841]" box="[151,1436,1643,1665]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF02396AFA41E4BF" box="[151,1436,1643,1665]" pageId="7" pageNumber="352">6´. Pygidium on basal half sparsely to moderately punctate; clypeal apex narrow...................................... 8</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023988FA41E484" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023988FA41E4A1" blockId="7.[151,1437,1261,1841]" box="[151,1436,1672,1694]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023988FA41E4A1" box="[151,1436,1672,1694]" pageId="7" pageNumber="352">
|
||||
7. Pronotal fovea and tubercle small, weak (
|
||||
<figureCitation id="94F860C3FF97E844FDE53989FD66E4A2" box="[624,699,1672,1694]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[232,1356,185,1911]" captionTargetId="figure-19@13.[226,1362,179,1917]" captionTargetPageId="13" captionText="FIGURE 9. Head and pronotum. A, Heterogomphus chevrolatii Burmeister, 1847 (male); B, Heterogomphus ulysses Burmeister, 1847 (male); C, Heterogomphus chevrolatii (female); D. Heterogomphus ulysses (female)." figureDoi="http://doi.org/10.5281/zenodo.14953249" httpUri="https://zenodo.org/record/14953249/files/figure.png" pageId="7" pageNumber="352">Fig. 9D</figureCitation>
|
||||
)..........................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB2B3989FA41E4A1" ID-CoL="7ZNJH" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1214,1436,1672,1694]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="ulysses">
|
||||
<emphasis id="3EB7A054FF97E844FB2B3989FA41E4A1" bold="true" box="[1214,1436,1672,1694]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB2B3989FAFFE4A1" bold="true" box="[1214,1314,1672,1694]" italics="true" pageId="7" pageNumber="352">H. ulysses</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF0239A7FA41E484" blockId="7.[151,1437,1261,1841]" box="[151,1436,1701,1723]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF0239A7FA41E484" box="[151,1436,1701,1723]" pageId="7" pageNumber="352">
|
||||
7’. Pronotal fovea and tubercle distinct (
|
||||
<figureCitation id="94F860C3FF97E844FDD139A4FD53E484" box="[580,654,1701,1723]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[232,1356,185,1911]" captionTargetId="figure-19@13.[226,1362,179,1917]" captionTargetPageId="13" captionText="FIGURE 9. Head and pronotum. A, Heterogomphus chevrolatii Burmeister, 1847 (male); B, Heterogomphus ulysses Burmeister, 1847 (male); C, Heterogomphus chevrolatii (female); D. Heterogomphus ulysses (female)." figureDoi="http://doi.org/10.5281/zenodo.14953249" httpUri="https://zenodo.org/record/14953249/files/figure.png" pageId="7" pageNumber="352">Fig. 9C</figureCitation>
|
||||
)...........................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB0F39A7FA41E484" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1178,1436,1701,1723]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="chevrolatii">
|
||||
<emphasis id="3EB7A054FF97E844FB0F39A7FA41E484" bold="true" box="[1178,1436,1701,1723]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB0F39A7FAFFE484" bold="true" box="[1178,1314,1701,1723]" italics="true" pageId="7" pageNumber="352">H. chevrolatii</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF0239C2FA41E4CA" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF0239C2FA41E4E7" blockId="7.[151,1437,1261,1841]" box="[151,1436,1731,1753]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF0239C2FA41E4E7" box="[151,1436,1731,1753]" pageId="7" pageNumber="352">8. Pronotum anteriorly rugose............................................................................. 9</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF0239E1FA41E4CA" blockId="7.[151,1437,1261,1841]" box="[151,1436,1760,1782]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF0239E1FA41E4CA" box="[151,1436,1760,1782]" pageId="7" pageNumber="352">
|
||||
8´. Pronotum anteriorly punctate.............................................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB4239E1FA41E4CA" ID-CoL="7ZNHJ" authority="Prell" authorityName="Prell" authorityYear="1912" box="[1239,1436,1760,1781]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="curvicornis">
|
||||
<emphasis id="3EB7A054FF97E844FB4239E1FA41E4CA" bold="true" box="[1239,1436,1760,1781]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB4239E1FAB8E4CA" bold="true" box="[1239,1381,1760,1781]" italics="true" pageId="7" pageNumber="352">H. curvicornis</emphasis>
|
||||
Prell
|
||||
</emphasis>
|
||||
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|
||||
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|
||||
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|
||||
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|
||||
<keyStep id="B73764E3FF97E844FF0239FFFA41E50F" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF0239FFFA41E52C" blockId="7.[151,1437,1261,1841]" box="[151,1436,1789,1811]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF0239FFFA41E52C" box="[151,1436,1789,1811]" pageId="7" pageNumber="352">
|
||||
9. Mesotibia and metatibia at apex truncate (
|
||||
<figureCitation id="94F860C3FF97E844FDE039FCFD16E52C" box="[629,715,1789,1811]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="7" pageNumber="352">Fig. 10C</figureCitation>
|
||||
)......................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB0839FFFA41E52C" ID-CoL="7ZNHL" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1181,1436,1789,1811]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="dilaticollis">
|
||||
<emphasis id="3EB7A054FF97E844FB0839FFFA41E52C" bold="true" box="[1181,1436,1789,1811]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB0839FFFAFEE52C" bold="true" box="[1181,1315,1789,1811]" italics="true" pageId="7" pageNumber="352">H. dilaticollis</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF02381AFA41E50F" blockId="7.[151,1437,1261,1841]" box="[151,1436,1818,1841]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF02381AFA41E50F" box="[151,1436,1818,1841]" pageId="7" pageNumber="352">
|
||||
9´. Mesotibia and metatibia at apex with two large teeth (
|
||||
<figureCitation id="94F860C3FF97E844FD73381AFCE0E50F" box="[742,829,1819,1841]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="7" pageNumber="352">Fig. 10D</figureCitation>
|
||||
)............................
|
||||
<taxonomicName id="CBC307C5FF97E844FB31381AFA41E50F" ID-CoL="7ZNJF" authority="(Burmeister)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1188,1436,1818,1840]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="telamon">
|
||||
<emphasis id="3EB7A054FF97E844FB31381AFA41E50F" bold="true" box="[1188,1436,1818,1840]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB31381AFACFE50F" bold="true" box="[1188,1298,1818,1840]" italics="true" pageId="7" pageNumber="352">H. telamon</emphasis>
|
||||
(Burmeister)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
267
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267
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<mods:title id="B9CC4A6875E14382B2498360C0F9DF03">A review of the genus Heterogomphus Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) from Colombia</mods:title>
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<mods:affiliation id="497FE87B3321C5AC06CCE9F5C3DC5D3C">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Investigación de Recursos Biológicos Alexander von Humboldt, Claustro de San Agustín, Villa de Leyva, Boyacá, Colombia</mods:affiliation>
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|
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Key to the adult males of
|
||||
<taxonomicName id="CBC307C5FF97E844FE423F96FD44E28D" ID-CoL="636SS" authorityName="Burmeister" authorityYear="1847" box="[471,665,151,178]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="3EB7A054FF97E844FE423F96FD44E28D" bold="true" box="[471,665,151,178]" italics="true" pageId="7" pageNumber="352">Heterogomphus</emphasis>
|
||||
</taxonomicName>
|
||||
species from
|
||||
<collectingCountry id="74D43CD6FF97E844FCD73F96FC61E28E" box="[834,956,151,177]" name="Colombia" pageId="7" pageNumber="352">Colombia</collectingCountry>
|
||||
</emphasis>
|
||||
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|
||||
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|
||||
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|
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Modified from
|
||||
<bibRefCitation id="685201B7FF97E844FED13FBAFDEBE2EA" author="Ratcliffe, B. C. & Cave, R. D. & Paucar-Cabrera, A." box="[324,566,187,214]" pageId="7" pageNumber="352" pagination="1 - 586" refId="ref11732" refString="Ratcliffe, B. C., Cave, R. D. & Paucar-Cabrera, A. (2020) The dynastine scarab beetles of Ecuador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 32, 1-586." type="journal article" year="2020">
|
||||
Ratcliffe
|
||||
<emphasis id="3EB7A054FF97E844FE393FBDFE00E2EA" box="[428,477,187,213]" italics="true" pageId="7" pageNumber="352">et al</emphasis>
|
||||
. (2020)
|
||||
</bibRefCitation>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023E02FE4BE3B6" blockId="7.[151,1437,259,393]" pageId="7" pageNumber="352">
|
||||
The absence of developed cephalic or pronotal structures is not a constant character that allows the differentiation of females and males because minor males may appear similar to females. However, males have the last abdominal ventrite medially emarginate (
|
||||
<figureCitation id="94F860C3FF97E844FE7A3E4AFD87E35A" box="[495,602,331,357]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="7" pageNumber="352">Fig. 10A</figureCitation>
|
||||
), while females have the last abdominal ventrite completely arcuate, not emarginate (
|
||||
<figureCitation id="94F860C3FF97E844FEB73E6EFE54E3B6" box="[290,393,367,393]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="7" pageNumber="352">Fig. 10B</figureCitation>
|
||||
).
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="44D92FCDFF97E844FF023EB5FA41E60D" pageId="7" pageNumber="352" type="key">
|
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<keyStep id="B73764E3FF97E844FF023EB5FA41E3D8" pageId="7" pageNumber="352">
|
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<paragraph id="0C7C7C46FF97E844FF023EB5FA41E3F6" blockId="7.[151,1437,436,1074]" box="[151,1436,436,458]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023EB5FA41E3F6" box="[151,1436,436,458]" pageId="7" pageNumber="352">1. Elytra completely, densely rugopunctate or with punctate rows................................................. 2</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023ED0FA41E3D8" blockId="7.[151,1437,436,1074]" box="[151,1436,465,487]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023ED0FA41E3D8" box="[151,1436,465,487]" pageId="7" pageNumber="352">1´. Elytra smooth........................................................................................ 7</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023EEEFA41E01E" pageId="7" pageNumber="352">
|
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<paragraph id="0C7C7C46FF97E844FF023EEEFA41E03B" blockId="7.[151,1437,436,1074]" box="[151,1436,495,517]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023EEEFA41E03B" box="[151,1436,495,517]" pageId="7" pageNumber="352">2. Elytra with long, moderately dense to dense setae............................................................ 3</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023D0DFA41E01E" blockId="7.[151,1437,436,1074]" box="[151,1436,524,546]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023D0DFA41E01E" box="[151,1436,524,546]" pageId="7" pageNumber="352">2´. Elytra glabrous or only with short, sparse setae on apex....................................................... 4</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
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|
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|
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<keyLead id="B732DF73FF97E844FF023D2BFA40E063" pageId="7" pageNumber="352">
|
||||
3. Clypeal anterior margin truncate (
|
||||
<figureCitation id="94F860C3FF97E844FDB23D28FDAEE000" box="[551,627,553,575]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="7" pageNumber="352">Fig. 7B</figureCitation>
|
||||
); pronotal punctures small (
|
||||
<figureCitation id="94F860C3FF97E844FCEA3D28FC1AE000" box="[895,967,553,575]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="7" pageNumber="352">Fig. 7F</figureCitation>
|
||||
)....................................................................................
|
||||
<emphasis id="3EB7A054FF97E844FC9A3D46FA40E063" bold="true" box="[783,1437,582,604]" pageId="7" pageNumber="352">
|
||||
<taxonomicName id="CBC307C5FF97E844FC9A3D46FAC1E063" authority="Neita Moreno, Clavijo-Bustos, & Ratcliffe" authorityName="Neita Moreno, Clavijo-Bustos, & Ratcliffe" authorityYear="2025" box="[783,1308,582,604]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="kogui" status="new species">
|
||||
<emphasis id="3EB7A054FF97E844FC9A3D46FCBBE063" bold="true" box="[783,870,582,604]" italics="true" pageId="7" pageNumber="352">H. kogui</emphasis>
|
||||
Neita Moreno, Clavijo-Bustos, & Ratcliffe
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="25841D2FFF97E844FAB33D46FA40E063" box="[1318,1437,582,604]" pageId="7" pageNumber="352" rank="species">new species</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023D65FA41E0A8" blockId="7.[151,1437,436,1074]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023D65FA41E0A8" pageId="7" pageNumber="352">
|
||||
3´. Clypeal anterior margin broad, deeply and triangularly emarginate (
|
||||
<figureCitation id="94F860C3FF97E844FCFE3D65FC64E046" box="[875,953,612,634]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="7" pageNumber="352">Fig. 7A</figureCitation>
|
||||
); pronotal punctures large (
|
||||
<figureCitation id="94F860C3FF97E844FB553D65FAD4E046" box="[1216,1289,612,634]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="7" pageNumber="352">Fig. 7E</figureCitation>
|
||||
)...........................................................................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB3E3D83FA41E0A8" ID-CoL="7ZNH9" authority="Dechambre" authorityName="Dechambre" authorityYear="1986" box="[1195,1436,641,663]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="carayoni">
|
||||
<emphasis id="3EB7A054FF97E844FB3E3D83FA41E0A8" bold="true" box="[1195,1436,641,663]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB3E3D83FAFDE0A9" bold="true" box="[1195,1312,641,663]" italics="true" pageId="7" pageNumber="352">H. carayoni</emphasis>
|
||||
Dechambre
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023D9EFA41E0ED" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023D9EFA41E08B" blockId="7.[151,1437,436,1074]" box="[151,1436,671,693]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023D9EFA41E08B" box="[151,1436,671,693]" pageId="7" pageNumber="352">
|
||||
4. Clypeal apex truncate...................................................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB423D9EFA41E08B" ID-CoL="7ZNHJ" authority="Prell" authorityName="Prell" authorityYear="1912" box="[1239,1436,671,692]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="curvicornis">
|
||||
<emphasis id="3EB7A054FF97E844FB423D9EFA41E08B" bold="true" box="[1239,1436,671,692]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB423D9EFAB8E08B" bold="true" box="[1239,1381,671,692]" italics="true" pageId="7" pageNumber="352">H. curvicornis</emphasis>
|
||||
Prell
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023DBDFA41E0ED" blockId="7.[151,1437,436,1074]" box="[151,1436,700,722]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023DBDFA41E0ED" box="[151,1436,700,722]" pageId="7" pageNumber="352">4´. Clypeal apex broad, deeply and triangularly emarginate....................................................... 5</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023DD8FA41E132" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023DD8FA41E0D0" blockId="7.[151,1437,436,1074]" box="[151,1436,729,751]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023DD8FA41E0D0" box="[151,1436,729,751]" pageId="7" pageNumber="352">
|
||||
5. Color dark reddish brown; elytra finely punctate...............................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB7C3DDBFA41E0D0" ID-CoL="7ZNJB" authority="Prell" authorityName="Prell" authorityYear="1912" box="[1257,1436,729,751]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="rugicollis">
|
||||
<emphasis id="3EB7A054FF97E844FB7C3DDBFA41E0D0" bold="true" box="[1257,1436,729,751]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB7C3DDBFAB8E0D0" bold="true" box="[1257,1381,729,751]" italics="true" pageId="7" pageNumber="352">H. rugicollis</emphasis>
|
||||
Prell
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023DF6FA41E132" blockId="7.[151,1437,436,1074]" box="[151,1436,759,781]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023DF6FA41E132" box="[151,1436,759,781]" pageId="7" pageNumber="352">5´. Color black; elytra densely punctate or rugopunctate......................................................... 6</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023C15FA41E178" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023C15FA41E115" blockId="7.[151,1437,436,1074]" box="[151,1436,788,810]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023C15FA41E115" box="[151,1436,788,810]" pageId="7" pageNumber="352">
|
||||
6. Mesotibial apex with 2–3 large teeth; parameres as in
|
||||
<figureCitation id="94F860C3FF97E844FD7C3C15FB7FE116" box="[745,1186,788,810]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="7" pageNumber="352">
|
||||
Fig. 3L–M.........................
|
||||
<taxonomicName id="CBC307C5FF97E844FB183C15FB7FE116" ID-CoL="7ZNJD" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1165,1186,788,809]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="schoenherri">
|
||||
<emphasis id="3EB7A054FF97E844FB183C15FB7FE116" bold="true" box="[1165,1186,788,809]" italics="true" pageId="7" pageNumber="352">H</emphasis>
|
||||
</taxonomicName>
|
||||
</figureCitation>
|
||||
. schoenherri Burmeister
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023C30FA41E178" blockId="7.[151,1437,436,1074]" box="[151,1436,817,839]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023C30FA41E178" box="[151,1436,817,839]" pageId="7" pageNumber="352">
|
||||
6´. Mesotibial apex with 1 large tooth; parameres as in
|
||||
<figureCitation id="94F860C3FF97E844FD463C30FB67E178" box="[723,1210,817,839]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="7" pageNumber="352">
|
||||
Fig. 3H–I..............................
|
||||
<taxonomicName id="CBC307C5FF97E844FB303C33FB67E178" ID-CoL="3L9MW" authority="(Thomson)" baseAuthorityName="Thomson" baseAuthorityYear="1859" box="[1189,1210,818,839]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="mniszechi">
|
||||
<emphasis id="3EB7A054FF97E844FB303C33FB67E178" bold="true" box="[1189,1210,818,839]" italics="true" pageId="7" pageNumber="352">H</emphasis>
|
||||
</taxonomicName>
|
||||
</figureCitation>
|
||||
. mniszechi (Thomson)
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023C4EFA41E1BE" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023C4EFA41E15B" blockId="7.[151,1437,436,1074]" box="[151,1436,847,869]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023C4EFA41E15B" box="[151,1436,847,869]" pageId="7" pageNumber="352">7. Clypeal apex truncate.................................................................................. 8</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023C6DFA41E1BE" blockId="7.[151,1437,436,1074]" box="[151,1436,876,898]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023C6DFA41E1BE" box="[151,1436,876,898]" pageId="7" pageNumber="352">7´. Clypeal apex broadly emarginate......................................................................... 9</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023C8BFA41E183" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023C8BFA41E1A0" blockId="7.[151,1437,436,1074]" box="[151,1436,905,927]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023C8BFA41E1A0" box="[151,1436,905,927]" pageId="7" pageNumber="352">
|
||||
8. Mesotibia and metatibia with truncate apex (
|
||||
<figureCitation id="94F860C3FF97E844FD1B3C88FD39E1A0" box="[654,740,905,927]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="7" pageNumber="352">Fig. 10C</figureCitation>
|
||||
)....................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB083C8BFA41E1A0" ID-CoL="7ZNHL" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1181,1436,905,927]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="dilaticollis">
|
||||
<emphasis id="3EB7A054FF97E844FB083C8BFA41E1A0" bold="true" box="[1181,1436,905,927]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB083C8BFAFEE1A0" bold="true" box="[1181,1315,905,927]" italics="true" pageId="7" pageNumber="352">H. dilaticollis</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023CA6FA41E183" blockId="7.[151,1437,436,1074]" box="[151,1436,934,957]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023CA6FA41E183" box="[151,1436,934,957]" pageId="7" pageNumber="352">
|
||||
8´. Mesotibia and metatibia at apex with two large teeth (
|
||||
<figureCitation id="94F860C3FF97E844FD733CA6FCE0E183" box="[742,829,935,957]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="7" pageNumber="352">Fig. 10D</figureCitation>
|
||||
)............................
|
||||
<taxonomicName id="CBC307C5FF97E844FB313CA6FA41E183" ID-CoL="7ZNJF" authority="(Burmeister)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1188,1436,934,956]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="telamon">
|
||||
<emphasis id="3EB7A054FF97E844FB313CA6FA41E183" bold="true" box="[1188,1436,934,956]" pageId="7" pageNumber="352">
|
||||
<emphasis id="3EB7A054FF97E844FB313CA6FACFE183" bold="true" box="[1188,1298,934,956]" italics="true" pageId="7" pageNumber="352">H. telamon</emphasis>
|
||||
(Burmeister)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF97E844FF023CC5FA41E60D" pageId="7" pageNumber="352">
|
||||
<paragraph id="0C7C7C46FF97E844FF023CC5FA41E1C8" blockId="7.[151,1437,436,1074]" pageId="7" pageNumber="352">
|
||||
<keyLead id="B732DF73FF97E844FF023CC5FA41E1C8" pageId="7" pageNumber="352">
|
||||
9. Apex of the medial trough in the elevated section of the pronotum broad, wider than width of head, broadly and deeply emarginate at middle (
|
||||
<figureCitation id="94F860C3FF97E844FE2C3CE0FDD8E1C8" box="[441,517,993,1015]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[232,1356,185,1911]" captionTargetId="figure-19@13.[226,1362,179,1917]" captionTargetPageId="13" captionText="FIGURE 9. Head and pronotum. A, Heterogomphus chevrolatii Burmeister, 1847 (male); B, Heterogomphus ulysses Burmeister, 1847 (male); C, Heterogomphus chevrolatii (female); D. Heterogomphus ulysses (female)." figureDoi="http://doi.org/10.5281/zenodo.14953249" httpUri="https://zenodo.org/record/14953249/files/figure.png" pageId="7" pageNumber="352">Fig. 9B</figureCitation>
|
||||
); parameres as in
|
||||
<figureCitation id="94F860C3FF97E844FD273CE0FB0EE1C8" box="[690,1235,993,1015]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="7" pageNumber="352">
|
||||
Fig. 3P–Q...................................
|
||||
<taxonomicName id="CBC307C5FF97E844FB2B3CE3FB0EE1C8" ID-CoL="7ZNJH" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1214,1235,994,1015]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="ulysses">
|
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<emphasis id="3EB7A054FF97E844FB2B3CE3FB0EE1C8" bold="true" box="[1214,1235,994,1015]" italics="true" pageId="7" pageNumber="352">H</emphasis>
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</taxonomicName>
|
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|
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. ulysses Burmeister
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF97E844FF023CFEFA41E60D" blockId="7.[151,1437,436,1074]" pageId="7" pageNumber="352">
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<keyLead id="B732DF73FF97E844FF023CFEFA41E60D" pageId="7" pageNumber="352">
|
||||
9´. Apex of the medial trough in the elevated section of the pronotum narrower, subequal in width to head in major males or narrower in minor males, weakly emarginate at middle (
|
||||
<figureCitation id="94F860C3FF97E844FD683B1DFC97E60E" box="[765,842,1052,1074]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[232,1356,185,1911]" captionTargetId="figure-19@13.[226,1362,179,1917]" captionTargetPageId="13" captionText="FIGURE 9. Head and pronotum. A, Heterogomphus chevrolatii Burmeister, 1847 (male); B, Heterogomphus ulysses Burmeister, 1847 (male); C, Heterogomphus chevrolatii (female); D. Heterogomphus ulysses (female)." figureDoi="http://doi.org/10.5281/zenodo.14953249" httpUri="https://zenodo.org/record/14953249/files/figure.png" pageId="7" pageNumber="352">Fig. 9A</figureCitation>
|
||||
); parameres as in
|
||||
<figureCitation id="94F860C3FF97E844FC623B1DFB72E60E" box="[1015,1199,1052,1074]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="7" pageNumber="352">
|
||||
Fig. 3A–B.....
|
||||
<taxonomicName id="CBC307C5FF97E844FB0F3B1DFB72E60E" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1178,1199,1052,1073]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="chevrolatii">
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<emphasis id="3EB7A054FF97E844FB0F3B1DFB72E60E" bold="true" box="[1178,1199,1052,1073]" italics="true" pageId="7" pageNumber="352">H</emphasis>
|
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</taxonomicName>
|
||||
</figureCitation>
|
||||
. chevrolatii Burmeister
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</subSubSection>
|
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</treatment>
|
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</document>
|
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263
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<mods:title id="CEF0BA428CD57FA947113E328D98A77B">A review of the genus Heterogomphus Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) from Colombia</mods:title>
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<mods:affiliation id="28F22E0C9A74166B764251682ADAFE43">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Investigación de Recursos Biológicos Alexander von Humboldt, Claustro de San Agustín, Villa de Leyva, Boyacá, Colombia</mods:affiliation>
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Clave para las hembras adultas de las especies de
|
||||
<taxonomicName id="CBC307C5FF98E84BFC923CE2FC14E1C1" ID-CoL="636SS" authorityName="Burmeister" authorityYear="1847" box="[775,969,995,1022]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="genus">
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<emphasis id="3EB7A054FF98E84BFC923CE2FC14E1C1" bold="true" box="[775,969,995,1022]" italics="true" pageId="8" pageNumber="353">Heterogomphus</emphasis>
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</taxonomicName>
|
||||
de
|
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<collectingCountry id="74D43CD6FF98E84BFC613CE2FBB3E1C2" box="[1012,1134,995,1021]" name="Colombia" pageId="8" pageNumber="353">Colombia</collectingCountry>
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Modificado de
|
||||
<bibRefCitation id="685201B7FF98E84BFED73B06FDE5E61E" author="Ratcliffe, B. C. & Cave, R. D. & Paucar-Cabrera, A." box="[322,568,1031,1058]" pageId="8" pageNumber="353" pagination="1 - 586" refId="ref11732" refString="Ratcliffe, B. C., Cave, R. D. & Paucar-Cabrera, A. (2020) The dynastine scarab beetles of Ecuador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 32, 1-586." type="journal article" year="2020">
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Ratcliffe
|
||||
<emphasis id="3EB7A054FF98E84BFE3E3B09FE01E61E" box="[427,476,1031,1057]" italics="true" pageId="8" pageNumber="353">et al</emphasis>
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||||
. (2020)
|
||||
</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
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</subSubSection>
|
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<subSubSection id="44D92FCDFF98E84BFF023B4DFA41E4B0" pageId="8" pageNumber="353" type="key">
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<key id="78522B77FF98E84BFF023B4DFA41E4B0" pageId="8" pageNumber="353">
|
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<keyStep id="B73764E3FF98E84BFF023B4DFA41E640" pageId="8" pageNumber="353">
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<paragraph id="0C7C7C46FF98E84BFF023B4DFA41E65E" blockId="8.[151,1437,1100,1679]" box="[151,1436,1100,1123]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023B4DFA41E65E" box="[151,1436,1100,1123]" pageId="8" pageNumber="353">1. Élitros completamente, densamente rugopunteados o con filas punteadas......................................... 2</keyLead>
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||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023B68FA41E640" blockId="8.[151,1437,1100,1679]" box="[151,1436,1129,1153]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023B68FA41E640" box="[151,1436,1129,1153]" pageId="8" pageNumber="353">1´. Élitros lisos, brillantes; puntos grandes, si están presentes, solamente cerca de los húmeros........................... 6</keyLead>
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||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023B86FA41E686" pageId="8" pageNumber="353">
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<paragraph id="0C7C7C46FF98E84BFF023B86FA41E6A3" blockId="8.[151,1437,1100,1679]" box="[151,1436,1159,1182]" pageId="8" pageNumber="353">
|
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<keyLead id="B732DF73FF98E84BFF023B86FA41E6A3" box="[151,1436,1159,1182]" pageId="8" pageNumber="353">2. Élitros con setas largas, moderadamente densas o densas...................................................... 3</keyLead>
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</paragraph>
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<paragraph id="0C7C7C46FF98E84BFF023BA5FA41E686" blockId="8.[151,1437,1100,1679]" box="[151,1436,1188,1211]" pageId="8" pageNumber="353">
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<keyLead id="B732DF73FF98E84BFF023BA5FA41E686" box="[151,1436,1188,1211]" pageId="8" pageNumber="353">2´. Élitros glabros o con no más de unas pocas setas cortas en sus ápices............................................ 4</keyLead>
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</paragraph>
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|
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<keyStep id="B73764E3FF98E84BFF023BC3FA41E710" pageId="8" pageNumber="353">
|
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<paragraph id="0C7C7C46FF98E84BFF023BC3FA41E6CB" blockId="8.[151,1437,1100,1679]" pageId="8" pageNumber="353">
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3. Élitros rugopunteados (
|
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<figureCitation id="94F860C3FF98E84BFE553BC0FDD0E6E8" box="[448,525,1217,1239]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="8" pageNumber="353">Fig. 7G</figureCitation>
|
||||
); mesotibia con el ápice truncado, con 4–6 setas gruesas, cortas (
|
||||
<figureCitation id="94F860C3FF98E84BFBE43BC0FB61E6E8" box="[1137,1212,1217,1239]" captionStart="FIGURE 8" captionStartId="11.[151,250,1836,1860]" captionTargetBox="[286,1302,815,1805]" captionTargetId="figure-330@11.[281,1307,810,1810]" captionTargetPageId="11" captionText="FIGURE 8. Mesotibiae and metatibiae (A–H).A, Mesotibia of Heterogomphus carayoni Dechambre, 1986 (male); B, Metatibia of Heterogomphus carayoni (male); C, Mesotibia of Heterogomphus carayoni (female); D, Metatibia of Heterogomphus carayoni (female); E, Mesotibia of Heterogomphus kogui new species (male holotype); F, Metatibia of Heterogomphus kogui (male holotype); G, Mesotibia of Heterogomphus kogui (female paratype); H, Metatibia of Heterogomphus kogui (female paratype). Parameres of Heterogomphus species (I–L): I, Heterogomphus carayoni (dorsal view) and J, (lateral view); K, Heterogomphus kogui (holotype) (dorsal view) and L (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.14953247" httpUri="https://zenodo.org/record/14953247/files/figure.png" pageId="8" pageNumber="353">Fig. 8G</figureCitation>
|
||||
); metatibia con el ápice truncado, con 5–6 setas gruesas, cortas (
|
||||
<figureCitation id="94F860C3FF98E84BFDF13BDEFD73E6CB" box="[612,686,1247,1269]" captionStart="FIGURE 8" captionStartId="11.[151,250,1836,1860]" captionTargetBox="[286,1302,815,1805]" captionTargetId="figure-330@11.[281,1307,810,1810]" captionTargetPageId="11" captionText="FIGURE 8. Mesotibiae and metatibiae (A–H).A, Mesotibia of Heterogomphus carayoni Dechambre, 1986 (male); B, Metatibia of Heterogomphus carayoni (male); C, Mesotibia of Heterogomphus carayoni (female); D, Metatibia of Heterogomphus carayoni (female); E, Mesotibia of Heterogomphus kogui new species (male holotype); F, Metatibia of Heterogomphus kogui (male holotype); G, Mesotibia of Heterogomphus kogui (female paratype); H, Metatibia of Heterogomphus kogui (female paratype). Parameres of Heterogomphus species (I–L): I, Heterogomphus carayoni (dorsal view) and J, (lateral view); K, Heterogomphus kogui (holotype) (dorsal view) and L (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.14953247" httpUri="https://zenodo.org/record/14953247/files/figure.png" pageId="8" pageNumber="353">Fig. 8H</figureCitation>
|
||||
).....
|
||||
<emphasis id="3EB7A054FF98E84BFD6F3BDEFA41E6CB" bold="true" box="[762,1436,1246,1268]" pageId="8" pageNumber="353">
|
||||
<taxonomicName id="CBC307C5FF98E84BFD6F3BDEFADAE6CB" authority="Neita Moreno, Clavijo-Bustos, & Ratcliffe" authorityName="Neita Moreno, Clavijo-Bustos, & Ratcliffe" authorityYear="2025" box="[762,1287,1246,1268]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="kogui" status="nueva especie">
|
||||
<emphasis id="3EB7A054FF98E84BFD6F3BDEFC8CE6CB" bold="true" box="[762,849,1246,1268]" italics="true" pageId="8" pageNumber="353">H. kogui</emphasis>
|
||||
Neita Moreno, Clavijo-Bustos, & Ratcliffe
|
||||
</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="25841D2FFF98E84BFA843BDEFA41E6CB" box="[1297,1436,1246,1268]" pageId="8" pageNumber="353" rank="species">nueva especie</taxonomicNameLabel>
|
||||
</emphasis>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023BFDFA41E710" blockId="8.[151,1437,1100,1679]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023BFDFA41E710" pageId="8" pageNumber="353">
|
||||
3´. Élitros con puntos oceladas (
|
||||
<figureCitation id="94F860C3FF98E84BFE6F3BFDFD95E72E" box="[506,584,1276,1298]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="8" pageNumber="353">Fig. 7H</figureCitation>
|
||||
); mesotibia con un diente apical largo (
|
||||
<figureCitation id="94F860C3FF98E84BFC5D3BFDFBC8E72E" box="[968,1045,1276,1298]" captionStart="FIGURE 8" captionStartId="11.[151,250,1836,1860]" captionTargetBox="[286,1302,815,1805]" captionTargetId="figure-330@11.[281,1307,810,1810]" captionTargetPageId="11" captionText="FIGURE 8. Mesotibiae and metatibiae (A–H).A, Mesotibia of Heterogomphus carayoni Dechambre, 1986 (male); B, Metatibia of Heterogomphus carayoni (male); C, Mesotibia of Heterogomphus carayoni (female); D, Metatibia of Heterogomphus carayoni (female); E, Mesotibia of Heterogomphus kogui new species (male holotype); F, Metatibia of Heterogomphus kogui (male holotype); G, Mesotibia of Heterogomphus kogui (female paratype); H, Metatibia of Heterogomphus kogui (female paratype). Parameres of Heterogomphus species (I–L): I, Heterogomphus carayoni (dorsal view) and J, (lateral view); K, Heterogomphus kogui (holotype) (dorsal view) and L (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.14953247" httpUri="https://zenodo.org/record/14953247/files/figure.png" pageId="8" pageNumber="353">Fig. 8C</figureCitation>
|
||||
); metatibia truncada con un diente corto apical (
|
||||
<figureCitation id="94F860C3FF98E84BFEBD3A18FEAEE710" box="[296,371,1305,1327]" captionStart="FIGURE 8" captionStartId="11.[151,250,1836,1860]" captionTargetBox="[286,1302,815,1805]" captionTargetId="figure-330@11.[281,1307,810,1810]" captionTargetPageId="11" captionText="FIGURE 8. Mesotibiae and metatibiae (A–H).A, Mesotibia of Heterogomphus carayoni Dechambre, 1986 (male); B, Metatibia of Heterogomphus carayoni (male); C, Mesotibia of Heterogomphus carayoni (female); D, Metatibia of Heterogomphus carayoni (female); E, Mesotibia of Heterogomphus kogui new species (male holotype); F, Metatibia of Heterogomphus kogui (male holotype); G, Mesotibia of Heterogomphus kogui (female paratype); H, Metatibia of Heterogomphus kogui (female paratype). Parameres of Heterogomphus species (I–L): I, Heterogomphus carayoni (dorsal view) and J, (lateral view); K, Heterogomphus kogui (holotype) (dorsal view) and L (lateral view)." figureDoi="http://doi.org/10.5281/zenodo.14953247" httpUri="https://zenodo.org/record/14953247/files/figure.png" pageId="8" pageNumber="353">Fig. 8D</figureCitation>
|
||||
)....................................................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB3E3A1BFA41E710" authority="Dechambre" authorityName="Dechambre" authorityYear="1986" box="[1195,1436,1305,1327]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="carayoni">
|
||||
<emphasis id="3EB7A054FF98E84BFB3E3A1BFA41E710" bold="true" box="[1195,1436,1305,1327]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB3E3A1BFAFDE711" bold="true" box="[1195,1312,1305,1327]" italics="true" pageId="8" pageNumber="353">H. carayoni</emphasis>
|
||||
Dechambre
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023A36FA41E755" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023A36FA41E773" blockId="8.[151,1437,1100,1679]" box="[151,1436,1334,1358]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023A36FA41E773" box="[151,1436,1334,1358]" pageId="8" pageNumber="353">
|
||||
4. Ápice del clípeo truncado o ligeramente emarginado...................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB183A36FA41E773" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1165,1436,1334,1356]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="schoenherri">
|
||||
<emphasis id="3EB7A054FF98E84BFB183A36FA41E773" bold="true" box="[1165,1436,1334,1356]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB183A36FAFEE773" bold="true" box="[1165,1315,1334,1356]" italics="true" pageId="8" pageNumber="353">H. schoenherri</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023A55FA41E755" blockId="8.[151,1437,1100,1679]" box="[151,1436,1364,1387]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023A55FA41E755" box="[151,1436,1364,1387]" pageId="8" pageNumber="353">4´. Ápice del clípeo profundamente emarginado................................................................ 5</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023A70FA41E79B" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023A70FA41E7B8" blockId="8.[151,1437,1100,1679]" box="[151,1436,1393,1417]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023A70FA41E7B8" box="[151,1436,1393,1417]" pageId="8" pageNumber="353">
|
||||
5. Ápice de la metatibia con un diente grande; élitros con puntos grandes y densos................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB303A73FA41E7B8" authority="(Thomson)" baseAuthorityName="Thomson" baseAuthorityYear="1859" box="[1189,1436,1393,1415]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="mniszechi">
|
||||
<emphasis id="3EB7A054FF98E84BFB303A73FA41E7B8" bold="true" box="[1189,1436,1393,1415]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB303A73FAFAE7B8" bold="true" box="[1189,1319,1393,1415]" italics="true" pageId="8" pageNumber="353">H. mniszechi</emphasis>
|
||||
(Thomson)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023A8EFA41E79B" blockId="8.[151,1437,1100,1679]" box="[151,1436,1422,1446]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023A8EFA41E79B" box="[151,1436,1422,1446]" pageId="8" pageNumber="353">
|
||||
5´. Ápice de la mesotibia subtruncado, crenulado; élitros con puntos dispersos a moderadamente densos.....
|
||||
<taxonomicName id="CBC307C5FF98E84BFB7C3A8EFA41E79B" authority="Prell" authorityName="Prell" authorityYear="1912" box="[1257,1436,1422,1444]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="rugicollis">
|
||||
<emphasis id="3EB7A054FF98E84BFB7C3A8EFA41E79B" bold="true" box="[1257,1436,1422,1444]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB7C3A8EFAB8E79B" bold="true" box="[1257,1381,1422,1444]" italics="true" pageId="8" pageNumber="353">H. rugicollis</emphasis>
|
||||
Prell
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023AADFA41E7E0" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023AADFA41E7FD" blockId="8.[151,1437,1100,1679]" box="[151,1436,1452,1474]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023AADFA41E7FD" box="[151,1436,1452,1474]" pageId="8" pageNumber="353">6. Pigidio en su mitad basal transversalmente rugoso o densamente punteado; ápice del clípeo ancho..................... 7</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023AC8FA41E7E0" blockId="8.[151,1437,1100,1679]" box="[151,1436,1481,1503]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023AC8FA41E7E0" box="[151,1436,1481,1503]" pageId="8" pageNumber="353">6´. Pigidio en su mitad basal escasa a moderadamente punteado; ápice del clípeo estrecho.............................. 8</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023AE6FA41E425" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023AE6FA41E7C3" blockId="8.[151,1437,1100,1679]" box="[151,1436,1510,1533]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023AE6FA41E7C3" box="[151,1436,1510,1533]" pageId="8" pageNumber="353">
|
||||
7. Pronoto con fóvea y tubérculos poco desarrollados (
|
||||
<figureCitation id="94F860C3FF98E84BFD433AE6FCFDE7C3" box="[726,800,1511,1533]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[232,1356,185,1911]" captionTargetId="figure-19@13.[226,1362,179,1917]" captionTargetPageId="13" captionText="FIGURE 9. Head and pronotum. A, Heterogomphus chevrolatii Burmeister, 1847 (male); B, Heterogomphus ulysses Burmeister, 1847 (male); C, Heterogomphus chevrolatii (female); D. Heterogomphus ulysses (female)." figureDoi="http://doi.org/10.5281/zenodo.14953249" httpUri="https://zenodo.org/record/14953249/files/figure.png" pageId="8" pageNumber="353">Fig. 9D</figureCitation>
|
||||
)..................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB2B3AE6FA41E7C3" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1214,1436,1510,1532]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="ulysses">
|
||||
<emphasis id="3EB7A054FF98E84BFB2B3AE6FA41E7C3" bold="true" box="[1214,1436,1510,1532]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB2B3AE6FAFFE7C3" bold="true" box="[1214,1314,1510,1532]" italics="true" pageId="8" pageNumber="353">H. ulysses</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023905FA41E425" blockId="8.[151,1437,1100,1679]" box="[151,1436,1540,1562]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023905FA41E425" box="[151,1436,1540,1562]" pageId="8" pageNumber="353">
|
||||
7´. Pronoto con fóvea y tubérculos desarrollados (
|
||||
<figureCitation id="94F860C3FF98E84BFD343905FD36E426" box="[673,747,1540,1562]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[232,1356,185,1911]" captionTargetId="figure-19@13.[226,1362,179,1917]" captionTargetPageId="13" captionText="FIGURE 9. Head and pronotum. A, Heterogomphus chevrolatii Burmeister, 1847 (male); B, Heterogomphus ulysses Burmeister, 1847 (male); C, Heterogomphus chevrolatii (female); D. Heterogomphus ulysses (female)." figureDoi="http://doi.org/10.5281/zenodo.14953249" httpUri="https://zenodo.org/record/14953249/files/figure.png" pageId="8" pageNumber="353">Fig. 9C</figureCitation>
|
||||
)...................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB0F3905FA41E425" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1178,1436,1540,1562]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="chevrolatii">
|
||||
<emphasis id="3EB7A054FF98E84BFB0F3905FA41E425" bold="true" box="[1178,1436,1540,1562]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB0F3905FAFFE425" bold="true" box="[1178,1314,1540,1562]" italics="true" pageId="8" pageNumber="353">H. chevrolatii</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023923FA41E46B" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023923FA41E408" blockId="8.[151,1437,1100,1679]" box="[151,1436,1569,1591]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023923FA41E408" box="[151,1436,1569,1591]" pageId="8" pageNumber="353">8. Pronoto anteriormente rugoso........................................................................... 9</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF02393EFA41E46B" blockId="8.[151,1437,1100,1679]" box="[151,1436,1599,1621]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF02393EFA41E46B" box="[151,1436,1599,1621]" pageId="8" pageNumber="353">
|
||||
8´. Pronoto anteriormente punteado...........................................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB42393EFA41E46B" authority="Prell" authorityName="Prell" authorityYear="1912" box="[1239,1436,1599,1620]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="curvicornis">
|
||||
<emphasis id="3EB7A054FF98E84BFB42393EFA41E46B" bold="true" box="[1239,1436,1599,1620]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB42393EFAB8E46B" bold="true" box="[1239,1381,1599,1620]" italics="true" pageId="8" pageNumber="353">H. curvicornis</emphasis>
|
||||
Prell
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF02395DFA41E4B0" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF02395DFA41E44D" blockId="8.[151,1437,1100,1679]" box="[151,1436,1628,1650]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF02395DFA41E44D" box="[151,1436,1628,1650]" pageId="8" pageNumber="353">
|
||||
9. Mesotibia y metatibia con el ápice truncado (
|
||||
<figureCitation id="94F860C3FF98E84BFD06395DFD34E44E" box="[659,745,1628,1650]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="8" pageNumber="353">Fig. 10C</figureCitation>
|
||||
)...................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB08395DFA41E44D" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1181,1436,1628,1650]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="dilaticollis">
|
||||
<emphasis id="3EB7A054FF98E84BFB08395DFA41E44D" bold="true" box="[1181,1436,1628,1650]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB08395DFAFEE44D" bold="true" box="[1181,1315,1628,1650]" italics="true" pageId="8" pageNumber="353">H. dilaticollis</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023978FA41E4B0" blockId="8.[151,1437,1100,1679]" box="[151,1436,1657,1679]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023978FA41E4B0" box="[151,1436,1657,1679]" pageId="8" pageNumber="353">
|
||||
9´. Mesotibia y metatibia con dos dientes grandes en el ápice (
|
||||
<figureCitation id="94F860C3FF98E84BFC8D3978FCB3E4B0" box="[792,878,1657,1679]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="8" pageNumber="353">Fig. 10D</figureCitation>
|
||||
)........................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB31397BFA41E4B0" authority="(Burmeister)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1188,1436,1657,1679]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="telamon">
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<emphasis id="3EB7A054FF98E84BFB31397BFA41E4B0" bold="true" box="[1188,1436,1657,1679]" pageId="8" pageNumber="353">
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<emphasis id="3EB7A054FF98E84BFB31397BFACFE4B0" bold="true" box="[1188,1298,1657,1679]" italics="true" pageId="8" pageNumber="353">H. telamon</emphasis>
|
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(Burmeister)
|
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</emphasis>
|
||||
</taxonomicName>
|
||||
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|
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263
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<mods:title id="B76003B5C458E9565C43A39B2A59E1DF">A review of the genus Heterogomphus Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) from Colombia</mods:title>
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<mods:affiliation id="1D987207558B60A09A592DA5B6FB0018">Sección Entomología, Colecciones Biológicas, Centro Colecciones y Gestión de Especies, Investigación de Recursos Biológicos Alexander von Humboldt, Claustro de San Agustín, Villa de Leyva, Boyacá, Colombia</mods:affiliation>
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Clave para los machos adultos de las especies de
|
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<taxonomicName id="CBC307C5FF98E84BFD6D3F96FC67E28D" ID-CoL="636SS" authorityName="Burmeister" authorityYear="1847" box="[760,954,151,178]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="genus">
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<emphasis id="3EB7A054FF98E84BFD6D3F96FC67E28D" bold="true" box="[760,954,151,178]" italics="true" pageId="8" pageNumber="353">Heterogomphus</emphasis>
|
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</taxonomicName>
|
||||
de
|
||||
<collectingCountry id="74D43CD6FF98E84BFC733F96FBBDE28E" box="[998,1120,151,177]" name="Colombia" pageId="8" pageNumber="353">Colombia</collectingCountry>
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</emphasis>
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|
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|
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<paragraph id="0C7C7C46FF98E84BFF023FBAFDE1E2EA" blockId="8.[151,1120,151,214]" box="[151,572,187,214]" pageId="8" pageNumber="353">
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Modificado de
|
||||
<bibRefCitation id="685201B7FF98E84BFED73FBAFDE5E2EA" author="Ratcliffe, B. C. & Cave, R. D. & Paucar-Cabrera, A." box="[322,568,187,214]" pageId="8" pageNumber="353" pagination="1 - 586" refId="ref11732" refString="Ratcliffe, B. C., Cave, R. D. & Paucar-Cabrera, A. (2020) The dynastine scarab beetles of Ecuador (Coleoptera: Scarabaeidae: Dynastinae). Bulletin of the University of Nebraska State Museum, 32, 1-586." type="journal article" year="2020">
|
||||
Ratcliffe
|
||||
<emphasis id="3EB7A054FF98E84BFE3E3FBDFE01E2EA" box="[427,476,187,213]" italics="true" pageId="8" pageNumber="353">et al</emphasis>
|
||||
. (2020)
|
||||
</bibRefCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="44D92FCDFF98E84BFF023E01FA41E1A4" pageId="8" pageNumber="353" type="key">
|
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<key id="78522B77FF98E84BFF023E01FA41E1A4" pageId="8" pageNumber="353">
|
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<paragraph id="0C7C7C46FF98E84BFF023E01FA41E32A" blockId="8.[151,1436,256,923]" box="[151,1436,256,279]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023E01FA41E32A" box="[151,1436,256,279]" pageId="8" pageNumber="353">1. Élitros completamente, densamente rugopunteados o con filas punteadas......................................... 2</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023E1CFA41E30C" blockId="8.[151,1436,256,923]" box="[151,1436,285,309]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023E1CFA41E30C" box="[151,1436,285,309]" pageId="8" pageNumber="353">1´. Élitros lisos.......................................................................................... 7</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023E3AFA41E352" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023E3AFA41E36F" blockId="8.[151,1436,256,923]" box="[151,1436,315,338]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023E3AFA41E36F" box="[151,1436,315,338]" pageId="8" pageNumber="353">2. Élitros con setas largas y moderadamente densas a densas..................................................... 3</keyLead>
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||||
</paragraph>
|
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<paragraph id="0C7C7C46FF98E84BFF023E59FA41E352" blockId="8.[151,1436,256,923]" box="[151,1436,344,367]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023E59FA41E352" box="[151,1436,344,367]" pageId="8" pageNumber="353">2´. Élitros glabros o con setas cortas y dispersas en los ápices..................................................... 4</keyLead>
|
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</paragraph>
|
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</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023E77FA41E3DC" pageId="8" pageNumber="353">
|
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<paragraph id="0C7C7C46FF98E84BFF023E77FA41E397" blockId="8.[151,1436,256,923]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023E77FA41E397" pageId="8" pageNumber="353">
|
||||
3. Ápice del clípeo truncado (
|
||||
<figureCitation id="94F860C3FF98E84BFE733E74FDEFE3B4" box="[486,562,373,395]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="8" pageNumber="353">Fig. 7B</figureCitation>
|
||||
); puntos en el pronoto densos y pequeños (
|
||||
<figureCitation id="94F860C3FF98E84BFC453E74FBCAE3B4" box="[976,1047,373,395]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="8" pageNumber="353">Fig. 7F</figureCitation>
|
||||
)...........................................................................
|
||||
<emphasis id="3EB7A054FF98E84BFD6F3E92FA41E397" bold="true" box="[762,1436,402,424]" pageId="8" pageNumber="353">
|
||||
<taxonomicName id="CBC307C5FF98E84BFD6F3E92FADAE397" authority="Neita Moreno, Clavijo-Bustos, & Ratcliffe" authorityName="Neita Moreno, Clavijo-Bustos, & Ratcliffe" authorityYear="2025" box="[762,1287,402,424]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="kogui" status="nueva especie">
|
||||
<emphasis id="3EB7A054FF98E84BFD6F3E92FC8CE397" bold="true" box="[762,849,402,424]" italics="true" pageId="8" pageNumber="353">H. kogui</emphasis>
|
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Neita Moreno, Clavijo-Bustos, & Ratcliffe
|
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</taxonomicName>
|
||||
,
|
||||
<taxonomicNameLabel id="25841D2FFF98E84BFA843E92FA41E397" box="[1297,1436,402,424]" pageId="8" pageNumber="353" rank="species">nueva especie</taxonomicNameLabel>
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</emphasis>
|
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</keyLead>
|
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</paragraph>
|
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<paragraph id="0C7C7C46FF98E84BFF023EB1FA41E3DC" blockId="8.[151,1436,256,923]" pageId="8" pageNumber="353">
|
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<keyLead id="B732DF73FF98E84BFF023EB1FA41E3DC" pageId="8" pageNumber="353">
|
||||
3´. Ápice del clípeo ancho, profunda y triangularmente emarginado (
|
||||
<figureCitation id="94F860C3FF98E84BFCC03EB1FC7FE3FA" box="[853,930,432,454]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="8" pageNumber="353">Fig. 7A</figureCitation>
|
||||
); puntos en el pronoto densos y grandes (
|
||||
<figureCitation id="94F860C3FF98E84BFAA53EB1FAA5E3FA" box="[1328,1400,432,454]" captionStart="FIGURE 7" captionStartId="10.[151,250,1485,1509]" captionTargetBox="[231,1356,564,1453]" captionTargetId="figure-214@10.[225,1362,558,1459]" captionTargetPageId="10" captionText="FIGURE 7. Head (dorsal view). A, Heterogomphus carayoni Dechambre, 1986 (male); B, Heterogomphus kogui new species (male holotype); C, Heterogomphus carayoni (female); D, Heterogomphus kogui (female paratype). Pronotum (male) (E, F): E; Heterogomphus carayoni; F; Heterogomphus kogui (holotype). Elytra (female) (G, H): G, Heterogomphus carayoni; H, Heterogomphus kogui (paratype)." figureDoi="http://doi.org/10.5281/zenodo.14953245" httpUri="https://zenodo.org/record/14953245/files/figure.png" pageId="8" pageNumber="353">Fig. 7E</figureCitation>
|
||||
)..................................................................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB3E3ECFFA41E3DC" authority="Dechambre" authorityName="Dechambre" authorityYear="1986" box="[1195,1436,461,483]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="carayoni">
|
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<emphasis id="3EB7A054FF98E84BFB3E3ECFFA41E3DC" bold="true" box="[1195,1436,461,483]" pageId="8" pageNumber="353">
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<emphasis id="3EB7A054FF98E84BFB3E3ECFFAFDE3DD" bold="true" box="[1195,1312,461,483]" italics="true" pageId="8" pageNumber="353">H. carayoni</emphasis>
|
||||
Dechambre
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
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|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023EEAFA41E021" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023EEAFA41E03F" blockId="8.[151,1436,256,923]" box="[151,1436,491,514]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023EEAFA41E03F" box="[151,1436,491,514]" pageId="8" pageNumber="353">
|
||||
4. Ápice del clípeo truncado................................................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB423EEAFA41E03F" authority="Prell" authorityName="Prell" authorityYear="1912" box="[1239,1436,491,512]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="curvicornis">
|
||||
<emphasis id="3EB7A054FF98E84BFB423EEAFA41E03F" bold="true" box="[1239,1436,491,512]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB423EEAFAB8E03F" bold="true" box="[1239,1381,491,512]" italics="true" pageId="8" pageNumber="353">H. curvicornis</emphasis>
|
||||
Prell
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023D09FA41E021" blockId="8.[151,1436,256,923]" box="[151,1436,520,543]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023D09FA41E021" box="[151,1436,520,543]" pageId="8" pageNumber="353">4´. Ápice del clípeo ancho, profunda y triangularmente emarginado................................................ 5</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023D24FA41E066" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023D24FA41E004" blockId="8.[151,1436,256,923]" box="[151,1436,549,571]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023D24FA41E004" box="[151,1436,549,571]" pageId="8" pageNumber="353">
|
||||
5. Color castaño rojizo oscuro; élitros finamente punteados........................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB7C3D27FA41E004" authority="Prell" authorityName="Prell" authorityYear="1912" box="[1257,1436,549,571]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="rugicollis">
|
||||
<emphasis id="3EB7A054FF98E84BFB7C3D27FA41E004" bold="true" box="[1257,1436,549,571]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB7C3D27FAB8E004" bold="true" box="[1257,1381,549,571]" italics="true" pageId="8" pageNumber="353">H. rugicollis</emphasis>
|
||||
Prell
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023D42FA41E066" blockId="8.[151,1436,256,923]" box="[151,1436,579,601]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023D42FA41E066" box="[151,1436,579,601]" pageId="8" pageNumber="353">5´. Color negro; élitros fuertemente punteados o rugopunteados................................................... 6</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023D61FA41E0AC" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023D61FA41E049" blockId="8.[151,1436,256,923]" box="[151,1436,608,631]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023D61FA41E049" box="[151,1436,608,631]" pageId="8" pageNumber="353">
|
||||
6. Ápice de la mesotibia con 2–3 dientes largos; parámeros como en la
|
||||
<figureCitation id="94F860C3FF98E84BFCE03D61FB7FE04A" box="[885,1186,608,630]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="8" pageNumber="353">
|
||||
Fig. 3L–M..............
|
||||
<taxonomicName id="CBC307C5FF98E84BFB183D61FB7FE04A" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1165,1186,608,629]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="schoenherri">
|
||||
<emphasis id="3EB7A054FF98E84BFB183D61FB7FE04A" bold="true" box="[1165,1186,608,629]" italics="true" pageId="8" pageNumber="353">H</emphasis>
|
||||
</taxonomicName>
|
||||
</figureCitation>
|
||||
. schoenherri Burmeister
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023D7CFA41E0AC" blockId="8.[151,1436,256,923]" box="[151,1436,637,661]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023D7CFA41E0AC" box="[151,1436,637,661]" pageId="8" pageNumber="353">
|
||||
6´. Ápice de la mesotibia con un diente largo; parámeros como en la
|
||||
<figureCitation id="94F860C3FF98E84BFCC23D7CFB67E0AC" box="[855,1210,637,659]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="8" pageNumber="353">
|
||||
Fig. 3H–I...................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB303D7FFB67E0AC" authority="(Thomson)" baseAuthorityName="Thomson" baseAuthorityYear="1859" box="[1189,1210,638,659]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="mniszechi">
|
||||
<emphasis id="3EB7A054FF98E84BFB303D7FFB67E0AC" bold="true" box="[1189,1210,638,659]" italics="true" pageId="8" pageNumber="353">H</emphasis>
|
||||
</taxonomicName>
|
||||
</figureCitation>
|
||||
. mniszechi (Thomson)
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023D9AFA41E0F2" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023D9AFA41E08F" blockId="8.[151,1436,256,923]" box="[151,1436,667,690]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023D9AFA41E08F" box="[151,1436,667,690]" pageId="8" pageNumber="353">7. Ápice clipeal truncado................................................................................. 8</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023DB9FA41E0F2" blockId="8.[151,1436,256,923]" box="[151,1436,696,719]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023DB9FA41E0F2" box="[151,1436,696,719]" pageId="8" pageNumber="353">7´. Ápice clipeal ampliamente emarginado.................................................................... 9</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023DD7FA41E137" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023DD7FA41E0D4" blockId="8.[151,1436,256,923]" box="[151,1436,725,747]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023DD7FA41E0D4" box="[151,1436,725,747]" pageId="8" pageNumber="353">
|
||||
8. Mesotibia y metatibia con el ápice truncado (
|
||||
<figureCitation id="94F860C3FF98E84BFD063DD4FD34E0D4" box="[659,745,725,747]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="8" pageNumber="353">Fig. 10C</figureCitation>
|
||||
)...................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB083DD7FA41E0D4" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1181,1436,725,747]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="dilaticollis">
|
||||
<emphasis id="3EB7A054FF98E84BFB083DD7FA41E0D4" bold="true" box="[1181,1436,725,747]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB083DD7FAFEE0D4" bold="true" box="[1181,1315,725,747]" italics="true" pageId="8" pageNumber="353">H. dilaticollis</emphasis>
|
||||
Burmeister
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023DF2FA41E137" blockId="8.[151,1436,256,923]" box="[151,1436,754,777]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023DF2FA41E137" box="[151,1436,754,777]" pageId="8" pageNumber="353">
|
||||
8´. Mesotibia y metatibia con dos dientes largos (
|
||||
<figureCitation id="94F860C3FF98E84BFD0F3DF2FD2CE137" box="[666,753,755,777]" captionStart="FIGURE 10" captionStartId="14.[151,250,1769,1793]" captionTargetBox="[158,1429,186,1736]" captionTargetId="figure-29@14.[151,1436,179,1743]" captionTargetPageId="14" captionText="FIGURE 10. Ventrite and metatibia diagnostic characters. A, Last abdominal ventrite emarginate of Heterogomphus chevrolatii Burmeister, 1847 (male); B, last abdominal ventrite entire Heterogomphus chevrolatii (male); C, metatibiae of Heterogomphus dilaticollis Burmeister, 1847; D, metatibiae of Heterogomphus telamon (Burmeister, 1847)." figureDoi="http://doi.org/10.5281/zenodo.14953252" httpUri="https://zenodo.org/record/14953252/files/figure.png" pageId="8" pageNumber="353">Fig. 10D</figureCitation>
|
||||
)...................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB313DF2FA41E137" authority="(Burmeister)" baseAuthorityName="Burmeister" baseAuthorityYear="1847" box="[1188,1436,754,776]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="telamon">
|
||||
<emphasis id="3EB7A054FF98E84BFB313DF2FA41E137" bold="true" box="[1188,1436,754,776]" pageId="8" pageNumber="353">
|
||||
<emphasis id="3EB7A054FF98E84BFB313DF2FACFE137" bold="true" box="[1188,1298,754,776]" italics="true" pageId="8" pageNumber="353">H. telamon</emphasis>
|
||||
(Burmeister)
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
<keyStep id="B73764E3FF98E84BFF023C11FA41E1A4" pageId="8" pageNumber="353">
|
||||
<paragraph id="0C7C7C46FF98E84BFF023C11FA41E17C" blockId="8.[151,1436,256,923]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023C11FA41E17C" pageId="8" pageNumber="353">
|
||||
9. Ápice de la escotadura media de la sección elevada del pronoto ancha, más amplio que el ancho de la cabeza, ampliamente y profundamente emarginado en el medio (
|
||||
<figureCitation id="94F860C3FF98E84BFDF83C2CFD64E17C" box="[621,697,813,835]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[232,1356,185,1911]" captionTargetId="figure-19@13.[226,1362,179,1917]" captionTargetPageId="13" captionText="FIGURE 9. Head and pronotum. A, Heterogomphus chevrolatii Burmeister, 1847 (male); B, Heterogomphus ulysses Burmeister, 1847 (male); C, Heterogomphus chevrolatii (female); D. Heterogomphus ulysses (female)." figureDoi="http://doi.org/10.5281/zenodo.14953249" httpUri="https://zenodo.org/record/14953249/files/figure.png" pageId="8" pageNumber="353">Fig. 9B</figureCitation>
|
||||
); parameros como en la
|
||||
<figureCitation id="94F860C3FF98E84BFC313C2CFB0EE17C" box="[932,1235,813,835]" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="8" pageNumber="353">
|
||||
Fig. 3P–Q...............
|
||||
<taxonomicName id="CBC307C5FF98E84BFB2B3C2FFB0EE17C" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1214,1235,814,835]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="ulysses">
|
||||
<emphasis id="3EB7A054FF98E84BFB2B3C2FFB0EE17C" bold="true" box="[1214,1235,814,835]" italics="true" pageId="8" pageNumber="353">H</emphasis>
|
||||
</taxonomicName>
|
||||
</figureCitation>
|
||||
. ulysses Burmeister
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
<paragraph id="0C7C7C46FF98E84BFF023C4AFA41E1A4" blockId="8.[151,1436,256,923]" pageId="8" pageNumber="353">
|
||||
<keyLead id="B732DF73FF98E84BFF023C4AFA41E1A4" pageId="8" pageNumber="353">
|
||||
9´. Ápice de la escotadura media de la sección elevada del pronoto angosta, subigual al ancho de la cabeza en los machos mayores o más angosto en los machos menores, ligeramente emarginado en el medio (
|
||||
<figureCitation id="94F860C3FF98E84BFC5D3C69FBC8E142" box="[968,1045,872,894]" captionStart="FIGURE 9" captionStartId="13.[151,250,1943,1967]" captionTargetBox="[232,1356,185,1911]" captionTargetId="figure-19@13.[226,1362,179,1917]" captionTargetPageId="13" captionText="FIGURE 9. Head and pronotum. A, Heterogomphus chevrolatii Burmeister, 1847 (male); B, Heterogomphus ulysses Burmeister, 1847 (male); C, Heterogomphus chevrolatii (female); D. Heterogomphus ulysses (female)." figureDoi="http://doi.org/10.5281/zenodo.14953249" httpUri="https://zenodo.org/record/14953249/files/figure.png" pageId="8" pageNumber="353">Fig. 9A</figureCitation>
|
||||
); parameros como en la
|
||||
<figureCitation id="94F860C3FF98E84BFA953C69FB72E1A4" captionStart="FIGURE 3" captionStartId="4.[151,250,1529,1553]" captionTargetBox="[158,1430,186,1497]" captionTargetId="figure-29@4.[151,1436,179,1504]" captionTargetPageId="4" captionText="FIGURE 3. Parameres. A (dorsal view) and B (lateral view) of Heterogomphus chevrolatii Burmeister, 1847; C (dorsal view) and D (lateral view) of Heterogomphus curvicornis Prell, 1912; F (dorsal view) and G (lateral view) of Heterogomphus dilaticollis Burmeister, 1847; H (dorsal view) and I (lateral view) of Heterogomphus mniszechi (Thomson, 1859); J (dorsal view) and K (lateral view) of Heterogomphus rugicollis Prell, 1912; L (dorsal view) and M (lateral view) of Heterogomphus schoenherri Burmeister, 1847; N (dorsal view) and O (lateral view) of Heterogomphus telamon (Burmeister, 1847); P (dorsal view) and Q (lateral view) of Heterogomphus ulysses Burmeister, 1847." figureDoi="http://doi.org/10.5281/zenodo.14953231" httpUri="https://zenodo.org/record/14953231/files/figure.png" pageId="8" pageNumber="353">
|
||||
Fig. 3A–B...................................................................................
|
||||
<taxonomicName id="CBC307C5FF98E84BFB0F3C87FB72E1A4" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[1178,1199,902,923]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="chevrolatii">
|
||||
<emphasis id="3EB7A054FF98E84BFB0F3C87FB72E1A4" bold="true" box="[1178,1199,902,923]" italics="true" pageId="8" pageNumber="353">H</emphasis>
|
||||
</taxonomicName>
|
||||
</figureCitation>
|
||||
. chevrolatii Burmeister
|
||||
</keyLead>
|
||||
</paragraph>
|
||||
</keyStep>
|
||||
</key>
|
||||
</subSubSection>
|
||||
</treatment>
|
||||
</document>
|
||||
|
|
@ -1,59 +1,59 @@
|
|||
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deposited in IAvH-E: “
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| Bosque intervenido, Bosque | Montano 11°06´19.4´´ N; 74°03´24.4´´ | W. WGS84.
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intervenido,
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; 74°03´24.4´´ | W. WGS
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Trampa
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|
||||
<collectorName id="A1361990FF98E84BFF7D3897FE9DE58F" box="[232,320,1942,1968]" pageId="8" pageNumber="353">Manual</collectorName>
|
||||
<collectingMethod id="D5820451FF98E84BFB7D3873FE9DE58F" pageId="8" pageNumber="353">Trampa de Luz | Negra, Manual</collectingMethod>
|
||||
,
|
||||
<date id="787D5A86FF98E84BFEDF3897FE65E58F" box="[330,440,1942,1968]" pageId="8" pageNumber="353" value="2019-06-02">
|
||||
<collectingDate id="6839A36EFF98E84BFEDF3897FE65E58F" box="[330,440,1942,1968]" pageId="8" pageNumber="353" value="2019-06-02">2.vi.2019</collectingDate>
|
||||
</date>
|
||||
.
|
||||
<collectingMunicipality id="EC18E63CFF98E84BFE543897FE2BE58F" box="[449,502,1942,1968]" pageId="8" pageNumber="353">Jhon</collectingMunicipality>
|
||||
|
|
||||
<collectingRegion id="CE07B2A4FF98E84BFD923897FD9AE58F" box="[519,583,1942,1968]" country="Colombia" name="Cesar" pageId="8" pageNumber="353">César</collectingRegion>
|
||||
<location id="091C2A9DFF98E84BFDD93897FD30E58F" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF98E84BFDD93897FD30E58F" box="[588,749,1942,1968]" country="Colombia" county="Trampa de Luz" latitude="11.10539" municipality="Jhon" name="Neita-Moreno" pageId="8" pageNumber="353" stateProvince="Magdalena">Neita-Moreno</location>
|
||||
” [white printed label] // “
|
||||
</materialsCitation>
|
||||
<materialsCitation id="BCAB761BFF98E849FB913897FB61E2C6" collectingDate="2019-06-02" collectionCode="UNSM" collectorName="Clavijo-Bustos & Ratcliffe & Vda. El Ramo & de Luz" country="Colombia" county="Neita-Moreno" elevation="84" lastPageId="10" lastPageNumber="355" latitude="11.10539" location="Santa" municipality="Instituto Humboldt" pageId="8" pageNumber="353" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="holotype">
|
||||
<typeStatus id="D378C2E4FF98E84BFB913897FBA6E58F" box="[1028,1147,1942,1968]" pageId="8" pageNumber="353" type="holotype">Holotypus</typeStatus>
|
||||
<specimenCount id="1AC5B7CFFF98E84BFB143897FB4BE58F" box="[1153,1174,1942,1968]" count="1" pageId="8" pageNumber="353" type="male">♂</specimenCount>
|
||||
<collectorName id="A1361990FF98E84BFE543897FE2BE58F" box="[449,502,1942,1968]" pageId="8" pageNumber="353">Jhon</collectorName>
|
||||
|
|
||||
<collectorName id="A1361990FF98E84BFD923897FD30E58F" box="[519,749,1942,1968]" pageId="8" pageNumber="353">César Neita-Moreno</collectorName>
|
||||
” [white printed label] // “Holotypus ♂ |
|
||||
<taxonomicName id="CBC307C5FF98E84BFB323897FA41E58F" authorityName="Neita Moreno & Clavijo-Bustos & Ratcliffe" authorityYear="2025" box="[1191,1436,1942,1968]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="353" phylum="Arthropoda" rank="species" species="kogui" status="sp. nov.">
|
||||
<emphasis id="3EB7A054FF98E84BFB323897FA41E58F" box="[1191,1436,1942,1968]" italics="true" pageId="8" pageNumber="353">Heterogomphus kogui</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="25841D2FFF98E84BFF0238BBFF08E5EB" box="[151,213,1978,2004]" pageId="8" pageNumber="353" rank="species">n. sp.</taxonomicNameLabel>
|
||||
|
|
||||
<collectingCounty id="E51D04CAFF98E84BFF7938BBFE53E5EB" box="[236,398,1978,2004]" pageId="8" pageNumber="353">Neita-Moreno</collectingCounty>
|
||||
,
|
||||
<collectorName id="A1361990FF98E84BFE0F38BBFD9FE5EB" box="[410,578,1978,2004]" pageId="8" pageNumber="353">Clavijo-Bustos</collectorName>
|
||||
&
|
||||
<collectorName id="A1361990FF98E84BFDFC38BBFD10E5EB" box="[617,717,1978,2004]" pageId="8" pageNumber="353">Ratcliffe</collectorName>
|
||||
, | 2024” [red printed label] // “
|
||||
<collectingMunicipality id="EC18E63CFF98E84BFBBB38BBFAD4E5EB" box="[1070,1289,1978,2004]" pageId="8" pageNumber="353">Instituto Humboldt</collectingMunicipality>
|
||||
|
|
||||
<collectingCountry id="74D43CD6FF98E84BFA8A38BBFA50E5EB" box="[1311,1421,1978,2004]" name="Colombia" pageId="8" pageNumber="353">Colombia</collectingCountry>
|
||||
| IAvH-E-240550” [white printed label with QR code].
|
||||
| Neita-Moreno, Clavijo-Bustos & Ratcliffe, | 2024” [red printed label] // “Instituto Humboldt | Colombia |
|
||||
<specimenCode id="5C65D43DFF98E84BFF0238DFFE92E5C6" box="[151,335,2014,2041]" pageId="8" pageNumber="353">IAvH-E-240550</specimenCode>
|
||||
” [white printed label with QR code]
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="BCAB761BFF98E849FD7138DCFB61E2C6" collectingDate="2019-06-02" collectingMethod="Trampa de Luz | Negra, Manual" collectionCode="UNSM" collectorName="Jhon & Cesar Neita-Moreno" country="Colombia" elevation="0" lastPageId="10" lastPageNumber="355" location="Santa | Marta, Corrg. Minca, Vda. El Ramo" pageId="8" pageNumber="353" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena" typeStatus="allotype">
|
||||
<emphasis id="3EB7A054FF98E84BFD7138DCFC8DE5C8" bold="true" box="[740,848,2013,2039]" pageId="8" pageNumber="353">
|
||||
<typeStatus id="D378C2E4FF98E84BFD7138DCFC96E5C8" box="[740,843,2013,2039]" pageId="8" pageNumber="353" type="allotype">Allotype</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
Female deposited in
|
||||
<specimenCount id="1AC5B7CFFF98E84BFCC138DFFC7BE5C7" box="[852,934,2014,2040]" pageId="8" pageNumber="353" type="female">Female</specimenCount>
|
||||
deposited in
|
||||
<collectionCode id="6AD2E483FF98E84BFBA038DFFB57E5C7" box="[1077,1162,2014,2040]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34868" lsid="urn:lsid:biocol.org:col:34868" name="University of Nebraska State Museum" pageId="8" pageNumber="353">UNSM</collectionCode>
|
||||
:“
|
||||
<collectingCountry id="74D43CD6FF98E84BFB0938DFFACDE5C7" box="[1180,1296,2014,2040]" name="Colombia" pageId="8" pageNumber="353">Colombia</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="CE07B2A4FF98E84BFA8C38DFFA45E5C7" box="[1305,1432,2014,2040]" country="Colombia" name="Magdalena" pageId="8" pageNumber="353">Magdalena</collectingRegion>
|
||||
,
|
||||
<location id="091C2A9DFF9AE849FF023F96FF08E28E" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FF023F96FF08E28E" box="[151,213,151,177]" country="Colombia" county="Neita-Moreno" latitude="11.10539" municipality="Instituto Humboldt" name="Santa" pageId="10" pageNumber="355" stateProvince="Magdalena">Santa</location>
|
||||
|
|
||||
<location id="091C2A9DFF9AE849FF7D3F96FEF3E28E" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FF7D3F96FEF3E28E" box="[232,302,151,177]" country="Colombia" county="Neita-Moreno" latitude="11.10539" municipality="Instituto Humboldt" name="Marta" pageId="10" pageNumber="355" stateProvince="Magdalena">Marta</location>
|
||||
, Corrg. Minca,
|
||||
<collectorName id="A1361990FF9AE849FE493F96FDA6E28E" box="[476,635,151,177]" pageId="10" pageNumber="355">Vda. El Ramo</collectorName>
|
||||
|
|
||||
<location id="091C2A9DFF9AE849FD183F96FD3CE28E" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FD183F96FD3CE28E" box="[653,737,151,177]" country="Colombia" county="Neita-Moreno" latitude="11.10539" municipality="Instituto Humboldt" name="Bosque" pageId="10" pageNumber="355" stateProvince="Magdalena">Bosque</location>
|
||||
intervenido,
|
||||
<location id="091C2A9DFF9AE849FCE03F96FC14E28E" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FCE03F96FC14E28E" box="[885,969,151,177]" country="Colombia" county="Neita-Moreno" latitude="11.10539" municipality="Instituto Humboldt" name="Bosque" pageId="10" pageNumber="355" stateProvince="Magdalena">Bosque</location>
|
||||
|
|
||||
<location id="091C2A9DFF9AE849FC493F96FB9CE28E" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FC493F96FB9CE28E" box="[988,1089,151,177]" country="Colombia" county="Neita-Moreno" latitude="11.10539" municipality="Instituto Humboldt" name="Montano" pageId="10" pageNumber="355" stateProvince="Magdalena">Montano</location>
|
||||
<geoCoordinate id="69F71A81FF9AE849FBD23F96FB29E28E" box="[1095,1268,151,177]" degrees="11" direction="north" minutes="06" orientation="latitude" pageId="10" pageNumber="355" precision="1" seconds="19.4" value="11.10539">11°06´19.4´´ N</geoCoordinate>
|
||||
; 74°03´24.4´´ | W. WGS
|
||||
<quantity id="CB3BD1A3FF9AE849FF683FBAFEA4E2EA" box="[253,377,186,213]" metricMagnitude="1" metricUnit="m" metricValue="8.42561" pageId="10" pageNumber="355" unit="m" value="84.2561">
|
||||
<elevation id="87EE9B75FF9AE849FF683FBAFEA4E2EA" box="[253,377,186,213]" metricMagnitude="1" metricUnit="m" metricValue="8.42561" pageId="10" pageNumber="355" unit="m" value="84.2561">84. 2561 m</elevation>
|
||||
</quantity>
|
||||
<location id="091C2A9DFF9AE849FF023F96FDA6E28E" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FF023F96FDA6E28E" box="[151,635,151,177]" country="Colombia" name="Santa | Marta, Corrg. Minca, Vda. El Ramo" pageId="10" pageNumber="355" stateProvince="Magdalena">Santa | Marta, Corrg. Minca, Vda. El Ramo</location>
|
||||
| Bosque intervenido, Bosque | Montano 11°06´19.4´´ N; 74°03´24.4´´ | W. WGS84.
|
||||
<elevation id="87EE9B75FF9AE849FEB63FBBFEA4E2EA" box="[291,377,186,213]" pageId="10" pageNumber="355">
|
||||
<quantity id="CB3BD1A3FF9AE849FEB63FBBFEA4E2EA" box="[291,377,186,213]" metricMagnitude="1" metricUnit="m" metricValue="8.42561" pageId="10" pageNumber="355" unit="m" value="84.2561">2561 m</quantity>
|
||||
</elevation>
|
||||
,
|
||||
<location id="091C2A9DFF9AE849FE163FBAFDF6E2EA" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FE163FBAFDF6E2EA" box="[387,555,187,213]" country="Colombia" county="Neita-Moreno" latitude="11.10539" municipality="Instituto Humboldt" name="Trampa de Luz" pageId="10" pageNumber="355" stateProvince="Magdalena">
|
||||
Trampa
|
||||
<collectorName id="A1361990FF9AE849FE4A3FBAFDF6E2EA" box="[479,555,187,213]" pageId="10" pageNumber="355">de Luz</collectorName>
|
||||
</location>
|
||||
|
|
||||
<location id="091C2A9DFF9AE849FDA93FBAFD5EE2EA" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FDA93FBAFD5EE2EA" box="[572,643,187,213]" country="Colombia" county="Neita-Moreno" latitude="11.10539" municipality="Instituto Humboldt" name="Negra" pageId="10" pageNumber="355" stateProvince="Magdalena">Negra</location>
|
||||
,
|
||||
<location id="091C2A9DFF9AE849FD1B3FBAFD3BE2EA" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FD1B3FBAFD3BE2EA" box="[654,742,187,213]" country="Colombia" county="Neita-Moreno" latitude="11.10539" municipality="Instituto Humboldt" name="Manual" pageId="10" pageNumber="355" stateProvince="Magdalena">Manual</location>
|
||||
<collectingMethod id="D5820451FF9AE849FE163FBAFD3BE2EA" box="[387,742,187,213]" pageId="10" pageNumber="355">Trampa de Luz | Negra, Manual</collectingMethod>
|
||||
,
|
||||
<date id="787D5A86FF9AE849FD653FBAFC83E2EA" box="[752,862,187,213]" pageId="10" pageNumber="355" value="2019-06-02">
|
||||
<collectingDate id="6839A36EFF9AE849FD653FBAFC83E2EA" box="[752,862,187,213]" pageId="10" pageNumber="355" value="2019-06-02">2.vi.2019</collectingDate>
|
||||
</date>
|
||||
. Jhon |
|
||||
<collectingRegion id="CE07B2A4FF9AE849FC383FBAFC30E2EA" box="[941,1005,187,213]" country="Colombia" name="Cesar" pageId="10" pageNumber="355">César</collectingRegion>
|
||||
Neita-Moreno” [white printed label] // “
|
||||
<typeStatus id="D378C2E4FF9AE849FF343FDEFECFE2C6" box="[161,274,223,249]" pageId="10" pageNumber="355" type="allotype">Allotypus</typeStatus>
|
||||
<specimenCount id="1AC5B7CFFF9AE849FE8D3FDEFEF0E2C6" box="[280,301,223,249]" count="1" pageId="10" pageNumber="355" type="female">♀</specimenCount>
|
||||
.
|
||||
<collectorName id="A1361990FF9AE849FCF23FBAFC41E2EA" box="[871,924,187,213]" pageId="10" pageNumber="355">Jhon</collectorName>
|
||||
|
|
||||
<collectorName id="A1361990FF9AE849FC383FBAFB49E2EA" box="[941,1172,187,213]" pageId="10" pageNumber="355">César Neita-Moreno</collectorName>
|
||||
” [white printed label] // “Allotypus ♀ |
|
||||
<taxonomicName id="CBC307C5FF9AE849FEAB3FDEFDE9E2C6" authorityName="Neita Moreno & Clavijo-Bustos & Ratcliffe" authorityYear="2025" box="[318,564,223,249]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="355" phylum="Arthropoda" rank="species" species="kogui" status="sp. nov.">
|
||||
<emphasis id="3EB7A054FF9AE849FEAB3FDEFDE9E2C6" box="[318,564,223,249]" italics="true" pageId="10" pageNumber="355">Heterogomphus kogui</emphasis>
|
||||
</taxonomicName>
|
||||
|
|
@ -206,67 +157,68 @@ Neita-Moreno” [white printed label] // “
|
|||
| Neita-Moreno, Clavijo-Bustos & Ratcliffe, | 2024”
|
||||
</materialsCitation>
|
||||
.
|
||||
<materialsCitation id="BCAB761BFF9AE849FB503FDEFBD5E3EE" collectingDate="1976-04-25" collectingDateMax="2019-06-02" collectingDateMin="1976-04-25" collectionCode="R" collectorName="C. Kugler & Det. J. C. Neita" country="Colombia" determinerName="Det. J. C. Neita" elevation="84" latitude="11.10539" location="Trampa de Luz" municipality="Female" pageId="10" pageNumber="355" specimenCount="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
|
||||
<materialsCitation id="BCAB761BFF9AE849FB503FDEFD3BE3B6" collectingDate="1976-04-25" collectionCode="IAvH-E" collectorName="R. C. Kugler" country="Colombia" determinerName="J. C. Neita" location="Santa Marta" municipality="Santa Marta" pageId="10" pageNumber="355" specimenCode="IAvH-E-88638" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Magdalena" typeStatus="paratype">
|
||||
<emphasis id="3EB7A054FF9AE849FB503FDEFA99E2C6" bold="true" box="[1221,1348,223,249]" pageId="10" pageNumber="355">
|
||||
<typeStatus id="D378C2E4FF9AE849FB503FDEFAE2E2C6" box="[1221,1343,223,249]" pageId="10" pageNumber="355" type="paratype">Paratypes</typeStatus>
|
||||
.
|
||||
</emphasis>
|
||||
<collectingMunicipality id="EC18E63CFF9AE849FADF3FDEFA41E2C6" box="[1354,1436,223,249]" pageId="10" pageNumber="355">Female</collectingMunicipality>
|
||||
and male deposited in IAvH-E: “
|
||||
<specimenCount id="1AC5B7CFFF9AE849FADF3FDEFA41E2C6" box="[1354,1436,223,249]" pageId="10" pageNumber="355" type="female">Female</specimenCount>
|
||||
and
|
||||
<specimenCount id="1AC5B7CFFF9AE849FF523E02FF20E322" box="[199,253,259,285]" pageId="10" pageNumber="355" type="male">male</specimenCount>
|
||||
deposited in
|
||||
<collectionCode id="6AD2E483FF9AE849FE033E02FE2EE322" box="[406,499,259,285]" pageId="10" pageNumber="355">IAvH-E</collectionCode>
|
||||
: “
|
||||
<collectingCountry id="74D43CD6FF9AE849FD923E02FDA5E322" box="[519,632,259,285]" name="Colombia" pageId="10" pageNumber="355">Colombia</collectingCountry>
|
||||
| Santa Marta |
|
||||
|
|
||||
<collectingMunicipality id="EC18E63CFF9AE849FD183E02FCCBE322" box="[653,790,259,285]" pageId="10" pageNumber="355">Santa Marta</collectingMunicipality>
|
||||
|
|
||||
<date id="787D5A86FF9AE849FCBE3E03FC15E322" box="[811,968,258,285]" pageId="10" pageNumber="355" value="1976-04-25">
|
||||
<collectingDate id="6839A36EFF9AE849FCBE3E03FC15E322" box="[811,968,258,285]" pageId="10" pageNumber="355" value="1976-04-25">25 April 1976</collectingDate>
|
||||
</date>
|
||||
//
|
||||
<collectingCountry id="74D43CD6FF9AE849FC723E02FB88E322" box="[999,1109,259,285]" name="Colombia" pageId="10" pageNumber="355">Colombia</collectingCountry>
|
||||
|
|
||||
// Colombia |
|
||||
<collectingRegion id="CE07B2A4FF9AE849FBFE3E02FB3AE322" box="[1131,1255,259,285]" country="Colombia" name="Magdalena" pageId="10" pageNumber="355">Magdalena</collectingRegion>
|
||||
|
|
||||
<collectionCode id="6AD2E483FF9AE849FB693E02FACFE322" box="[1276,1298,259,285]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="355">R</collectionCode>
|
||||
.
|
||||
<collectorName id="A1361990FF9AE849FA833E02FA58E322" box="[1302,1413,259,285]" pageId="10" pageNumber="355">C. Kugler</collectorName>
|
||||
<collectorName id="A1361990FF9AE849FB693E02FA58E322" box="[1276,1413,259,285]" pageId="10" pageNumber="355">R.C. Kugler</collectorName>
|
||||
//
|
||||
<taxonomicName id="CBC307C5FF9AE849FF023E26FE6BE37F" authorityName="Dechambre" authorityYear="1986" box="[151,438,295,321]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="355" phylum="Arthropoda" rank="species" species="carayoni">
|
||||
<emphasis id="3EB7A054FF9AE849FF023E26FE6BE37F" box="[151,438,295,321]" italics="true" pageId="10" pageNumber="355">Heterogomphus carayoni</emphasis>
|
||||
</taxonomicName>
|
||||
|
|
||||
<bibRefCitation id="685201B7FF9AE849FE583E26FD4BE37E" author="Dechambre, R. P." box="[461,662,295,321]" pageId="10" pageNumber="355" pagination="306 - 307" refId="ref11062" refString="Dechambre, R. P. (1986) Heterogomphus carayoni, une nouvelle espece de coleoptere Dynastidae. Annales de la Societe Entomologique de France, 22, 306-307. https://doi.org/10.1080/21686351.1986.12278441" type="journal article" year="1986">Dechambre, 1986</bibRefCitation>
|
||||
|
|
||||
<determinerName id="F9A09082FF9AE849FD383E26FCBEE37E" box="[685,867,295,321]" pageId="10" pageNumber="355">
|
||||
<collectorName id="A1361990FF9AE849FD383E26FCBEE37E" box="[685,867,295,321]" pageId="10" pageNumber="355">Det. J. C. Neita</collectorName>
|
||||
</determinerName>
|
||||
, 2006” // “
|
||||
<typeStatus id="D378C2E4FF9AE849FC4A3E26FB8DE37E" box="[991,1104,295,321]" pageId="10" pageNumber="355" type="paratype">Paratypus</typeStatus>
|
||||
<specimenCount id="1AC5B7CFFF9AE849FBCC3E26FBB3E37E" box="[1113,1134,295,321]" count="1" pageId="10" pageNumber="355" type="male">♂</specimenCount>
|
||||
|
|
||||
| Det.
|
||||
<determinerName id="F9A09082FF9AE849FD733E26FCBEE37E" box="[742,867,295,321]" pageId="10" pageNumber="355">J. C. Neita</determinerName>
|
||||
, 2006” // “Paratypus ♂ |
|
||||
<taxonomicName id="CBC307C5FF9AE849FB103E26FAA3E37E" authorityName="Neita Moreno & Clavijo-Bustos & Ratcliffe" authorityYear="2025" box="[1157,1406,295,321]" class="Insecta" family="Scarabaeidae" genus="Heterogomphus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="355" phylum="Arthropoda" rank="species" species="kogui" status="sp. nov.">
|
||||
<emphasis id="3EB7A054FF9AE849FB103E26FAA3E37E" box="[1157,1406,295,321]" italics="true" pageId="10" pageNumber="355">Heterogomphus kogui</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="25841D2FFF9AE849FA123E26FF6AE35A" pageId="10" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
|
||||
| Neita-Moreno, Clavijo-Bustos & Ratcliffe, | 2024” [yellow printed label] // “Instituto Humboldt |
|
||||
<collectingCountry id="74D43CD6FF9AE849FAB53E4AFA53E35A" box="[1312,1422,331,357]" name="Colombia" pageId="10" pageNumber="355">Colombia</collectingCountry>
|
||||
| “IAvH-E-88638” [white printed label with QR code]; “
|
||||
| Neita-Moreno, Clavijo-Bustos & Ratcliffe, | 2024” [yellow printed label] // “Instituto Humboldt | Colombia | “
|
||||
<specimenCode id="5C65D43DFF9AE849FF343E6EFE93E3B6" box="[161,334,367,393]" pageId="10" pageNumber="355">IAvH-E-88638</specimenCode>
|
||||
” [white printed label with QR code]
|
||||
</materialsCitation>
|
||||
;
|
||||
<materialsCitation id="BCAB761BFF9AE849FD643E6EFD2EE026" collectingDate="2019-06-02" collectingMethod="Trampa de Luz | Negra, Manual" collectorName="Jhon & Cesar Neita-Moreno" country="Colombia" elevation="84.2561" location="Santa | Marta, Corrg. Minca, Vda. El Ramo" pageId="10" pageNumber="355" specimenCode="IAvH-E-240549" specimenCount="1" specimenCount-female="1" stateProvince="Magdalena" typeStatus="paratype">
|
||||
“
|
||||
<collectingCountry id="74D43CD6FF9AE849FD6E3E6EFCB2E3B6" box="[763,879,367,393]" name="Colombia" pageId="10" pageNumber="355">Colombia</collectingCountry>
|
||||
,
|
||||
<collectingRegion id="CE07B2A4FF9AE849FCEC3E6EFC25E3B6" box="[889,1016,367,393]" country="Colombia" name="Magdalena" pageId="10" pageNumber="355">Magdalena</collectingRegion>
|
||||
, Santa | Marta, Corrg. Minca, Vda. El Ramo | Bosque intervenido, Bosque | Montano
|
||||
<geoCoordinate id="69F71A81FF9AE849FD363E92FC8DE392" box="[675,848,403,429]" degrees="11" direction="north" minutes="06" orientation="latitude" pageId="10" pageNumber="355" precision="1" seconds="19.4" value="11.10539">11°06´19.4´´ N</geoCoordinate>
|
||||
; 74°03´24.4´´ | W. WGS
|
||||
<quantity id="CB3BD1A3FF9AE849FBF73E92FB03E392" box="[1122,1246,402,429]" metricMagnitude="1" metricUnit="m" metricValue="8.42561" pageId="10" pageNumber="355" unit="m" value="84.2561">
|
||||
<elevation id="87EE9B75FF9AE849FBF73E92FB03E392" box="[1122,1246,402,429]" metricMagnitude="1" metricUnit="m" metricValue="8.42561" pageId="10" pageNumber="355" unit="m" value="84.2561">84. 2561 m</elevation>
|
||||
</quantity>
|
||||
,
|
||||
<location id="091C2A9DFF9AE849FB7D3E92FA4CE392" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FB7D3E92FA4CE392" box="[1256,1425,403,429]" country="Colombia" latitude="11.10539" municipality="Female" name="Trampa de Luz" pageId="10" pageNumber="355" stateProvince="Magdalena">Trampa de Luz</location>
|
||||
| Negra, Manual,
|
||||
<location id="091C2A9DFF9AE849FB963E6EFF07E392" LSID="urn:lsid:plazi:treatment:846ACD50FF98E84FFF0239E0FCD5E7DA:091C2A9DFF9AE849FB963E6EFF07E392" country="Colombia" name="Santa | Marta, Corrg. Minca, Vda. El Ramo" pageId="10" pageNumber="355" stateProvince="Magdalena">Santa | Marta, Corrg. Minca, Vda. El Ramo</location>
|
||||
| Bosque intervenido, Bosque | Montano 11°06´19.4´´ N; 74°03´24.4´´ | W. WGS84.
|
||||
<elevation id="87EE9B75FF9AE849FB1D3E93FB03E392" box="[1160,1246,402,429]" pageId="10" pageNumber="355">
|
||||
<quantity id="CB3BD1A3FF9AE849FB1D3E93FB03E392" box="[1160,1246,402,429]" metricMagnitude="1" metricUnit="m" metricValue="8.42561" pageId="10" pageNumber="355" unit="m" value="84.2561">2561 m</quantity>
|
||||
</elevation>
|
||||
,
|
||||
<collectingMethod id="D5820451FF9AE849FB7D3E92FE9CE3EE" pageId="10" pageNumber="355">Trampa de Luz | Negra, Manual</collectingMethod>
|
||||
,
|
||||
<date id="787D5A86FF9AE849FEDE3EB6FE64E3EE" box="[331,441,439,465]" pageId="10" pageNumber="355" value="2019-06-02">
|
||||
<collectingDate id="6839A36EFF9AE849FEDE3EB6FE64E3EE" box="[331,441,439,465]" pageId="10" pageNumber="355" value="2019-06-02">2.vi.2019</collectingDate>
|
||||
</date>
|
||||
. Jhon |
|
||||
<collectingRegion id="CE07B2A4FF9AE849FD9C3EB6FD94E3EE" box="[521,585,439,465]" country="Colombia" name="Cesar" pageId="10" pageNumber="355">César</collectingRegion>
|
||||
Neita-Moreno” [white printed label] // “
|
||||
</materialsCitation>
|
||||
<materialsCitation id="BCAB761BFF9AE849FB9D3EB6FD32E026" collectorName="Clavijo-Bustos & Ratcliffe" country="Colombia" location="Colombia" pageId="10" pageNumber="355" specimenCount="1" specimenCount-female="1" typeStatus="allotype">
|
||||
.
|
||||
<collectorName id="A1361990FF9AE849FE573EB6FE2AE3EE" box="[450,503,439,465]" pageId="10" pageNumber="355">Jhon</collectorName>
|
||||
|
|
||||
<collectorName id="A1361990FF9AE849FD9C3EB6FD2DE3EE" box="[521,752,439,465]" pageId="10" pageNumber="355">César Neita-Moreno</collectorName>
|
||||
” [white printed label] // “
|
||||
<typeStatus id="D378C2E4FF9AE849FB9D3EB6FBA4E3EE" box="[1032,1145,439,465]" pageId="10" pageNumber="355" type="allotype">Allotypus</typeStatus>
|
||||
<specimenCount id="1AC5B7CFFF9AE849FBEA3EB6FB49E3EE" box="[1151,1172,439,465]" count="1" pageId="10" pageNumber="355" type="female">♀</specimenCount>
|
||||
|
|
||||
|
|
@ -274,15 +226,11 @@ Neita-Moreno” [white printed label] // “
|
|||
<emphasis id="3EB7A054FF9AE849FB333EB6FA41E3EE" box="[1190,1436,439,465]" italics="true" pageId="10" pageNumber="355">Heterogomphus kogui</emphasis>
|
||||
</taxonomicName>
|
||||
<taxonomicNameLabel id="25841D2FFF9AE849FF023EDAFF08E3CA" box="[151,213,475,501]" pageId="10" pageNumber="355" rank="species">n. sp.</taxonomicNameLabel>
|
||||
| Neita-Moreno,
|
||||
<collectorName id="A1361990FF9AE849FE0F3EDAFD9FE3CA" box="[410,578,475,501]" pageId="10" pageNumber="355">Clavijo-Bustos</collectorName>
|
||||
&
|
||||
<collectorName id="A1361990FF9AE849FDFC3EDAFD10E3CA" box="[617,717,475,501]" pageId="10" pageNumber="355">Ratcliffe</collectorName>
|
||||
, | 2024” [red printed label] // “Instituto Humboldt |
|
||||
<collectingCountry id="74D43CD6FF9AE849FA8A3EDAFA50E3CA" box="[1311,1421,475,501]" name="Colombia" pageId="10" pageNumber="355">Colombia</collectingCountry>
|
||||
| IAvH-E-240549” [white printed label with QR code]
|
||||
| Neita-Moreno, Clavijo-Bustos & Ratcliffe, | 2024” [red printed label] // “Instituto Humboldt | Colombia |
|
||||
<specimenCode id="5C65D43DFF9AE849FF023EFFFE8DE026" box="[151,336,510,537]" pageId="10" pageNumber="355">IAvH-E-240549</specimenCode>
|
||||
” [white printed label with QR code].
|
||||
</materialsCitation>
|
||||
.’
|
||||
’
|
||||
</paragraph>
|
||||
<caption id="58BC2CCEFF99E84AFF023896FD32E5EB" ID-DOI="http://doi.org/10.5281/zenodo.14953243" ID-Zenodo-Dep="14953243" httpUri="https://zenodo.org/record/14953243/files/figure.png" pageId="9" pageNumber="354" startId="9.[151,250,1943,1967]" targetBox="[209,1378,196,1911]" targetPageId="9" targetType="figure">
|
||||
<paragraph id="0C7C7C46FF99E84AFF023896FD32E5EB" blockId="9.[151,1436,1943,2004]" pageId="9" pageNumber="354">
|
||||
|
|
|
|||
|
|
@ -1,77 +1,77 @@
|
|||
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|
||||
<mods:mods id="183CCB132AF512B079B0054056232E5C" xmlns:mods="http://www.loc.gov/mods/v3">
|
||||
<mods:titleInfo id="03D1AA4A348EECC282DE6045A0AF8331">
|
||||
<mods:title id="1EE70AA1B8F09B09202C3B674BC3592A">Barkeria wixarika (Laeliinae, Epidendroideae, Orchidaceae), a new species from Nayarit, Mexico</mods:title>
|
||||
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<mods:title id="65AA561AC2D50A1E5A09799A4E6B7117">Barkeria wixarika (Laeliinae, Epidendroideae, Orchidaceae), a new species from Nayarit, Mexico</mods:title>
|
||||
</mods:titleInfo>
|
||||
<mods:name id="4910D0D5F18F7151398C966CFE602AA0" type="personal">
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<mods:role id="E6990F1402A2B23A33AADF49FE04B4A5">
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||||
<mods:role id="67BE85D5D8ABD5821FF3F16B4C21457B">
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<mods:roleTerm id="C1CB6A1AE8DCFD209C59A5CB422EBC90">Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart id="28C001F1B58679BC0A848C37788E4639">León-Peralta, Cekouat E.</mods:namePart>
|
||||
<mods:nameIdentifier id="0BA0E7ED094BA11808F431A47D3189A4" type="ORCID">0000-0003-4496-2044</mods:nameIdentifier>
|
||||
<mods:affiliation id="6056BB48CE3ADABF22246A8371140380">Departamento de Botánica, Instituto de Biología, Universidad Nacional Autónoma de México, Apartado Postal 70 - 367, C. P. 04510, Ciudad de México, México & Departamento de Ecología y Recursos Naturales, Facultad de Ciencias, Universidad Nacional Autónoma de México, Ciudad Universitaria, Coyoacán, 04510, Ciudad de México, México & Posgrado en Ciencias Biológicas, Unidad de Posgrado, Edificio D, 1 ° Piso, Circuito de Posgrados, Ciudad Universitaria, Coyoacán, C. P. 04510, Ciudad de México, México</mods:affiliation>
|
||||
<mods:nameIdentifier id="98D5E44B09331AB1874EC57465AFEBBD" type="email">cekouat@gmail.com</mods:nameIdentifier>
|
||||
<mods:namePart id="12416FA45DD1D2BFA4E281F90B14E291">León-Peralta, Cekouat E.</mods:namePart>
|
||||
<mods:nameIdentifier id="675ADFCC16FC251DA9F65B9E1D529361" type="ORCID">0000-0003-4496-2044</mods:nameIdentifier>
|
||||
<mods:affiliation id="974D65513956F926EA371638466600B7">Departamento de Botánica, Instituto de Biología, Universidad Nacional Autónoma de México, Apartado Postal 70 - 367, C. P. 04510, Ciudad de México, México & Departamento de Ecología y Recursos Naturales, Facultad de Ciencias, Universidad Nacional Autónoma de México, Ciudad Universitaria, Coyoacán, 04510, Ciudad de México, México & Posgrado en Ciencias Biológicas, Unidad de Posgrado, Edificio D, 1 ° Piso, Circuito de Posgrados, Ciudad Universitaria, Coyoacán, C. P. 04510, Ciudad de México, México</mods:affiliation>
|
||||
<mods:nameIdentifier id="2BF21FEDC8125C4D6378D42AA3E2B46B" type="email">cekouat@gmail.com</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="0A39C5C6BB4ED3FD9BC842414B3C3A4F" type="personal">
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<mods:role id="50D9338C48F8722282A021D7F72FA398">
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<mods:name id="6759F7EA2BAE6A8D3A14B6F748305B4A" type="personal">
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<mods:role id="E24B84BFFEB0FB1FF4C86835D47AA32B">
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<mods:roleTerm id="B1EA71005D79F04ADB55703B5B435440">Author</mods:roleTerm>
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||||
</mods:role>
|
||||
<mods:namePart id="E5ACF62DD2721547C79EE8B95CA2EC88">Jones, Jason</mods:namePart>
|
||||
<mods:nameIdentifier id="9E4FE44275143E69C9D9D674294EFBE4" type="ORCID">0000-0001-6460-8427</mods:nameIdentifier>
|
||||
<mods:affiliation id="D203C374D0F9B87D314331746B789424">Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, México</mods:affiliation>
|
||||
<mods:nameIdentifier id="075FEF12D10B74A1DABC5026B3A98514" type="email">jasonjones@crotalus.com</mods:nameIdentifier>
|
||||
<mods:namePart id="B5DD852E650297E87E8152A2F1A612E3">Jones, Jason</mods:namePart>
|
||||
<mods:nameIdentifier id="ACBF6A27827FE375F04B840E6F4EE449" type="ORCID">0000-0001-6460-8427</mods:nameIdentifier>
|
||||
<mods:affiliation id="F0BAC34CB42C7FBB7B658FE0FF96A5EC">Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, México</mods:affiliation>
|
||||
<mods:nameIdentifier id="4A8D77A1E1C4FD8FB93A2FC55067248D" type="email">jasonjones@crotalus.com</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="259D15D664D3CBB8C264734503FFF800" type="personal">
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<mods:roleTerm id="8DA2EC1DA176003DA06198503D4D1C4A">Author</mods:roleTerm>
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</mods:role>
|
||||
<mods:namePart id="AEBB3B82A736E0E9230FA3ADAED4BD06">Reynaud, Sergio</mods:namePart>
|
||||
<mods:nameIdentifier id="23192E7EC0E2CEE3C39F38B6E5B609DC" type="ORCID">0009-0009-6455-9544</mods:nameIdentifier>
|
||||
<mods:affiliation id="E89EFA7AA4E899FE1D97CB80B041B511">Departamento de Ecología y Recursos Naturales, Facultad de Ciencias, Universidad Nacional Autónoma de México, Ciudad Universitaria, Coyoacán, 04510, Ciudad de México, México</mods:affiliation>
|
||||
<mods:nameIdentifier id="45863E08F138631F03B6530380C05C49" type="email">orquidiero@gmail.com</mods:nameIdentifier>
|
||||
<mods:namePart id="C77D748487D8D4914F223552950FD23A">Reynaud, Sergio</mods:namePart>
|
||||
<mods:nameIdentifier id="8F6C8285945F21CD3BF01214F59908C4" type="ORCID">0009-0009-6455-9544</mods:nameIdentifier>
|
||||
<mods:affiliation id="B65E62D521B174A4318A07F136131602">Departamento de Ecología y Recursos Naturales, Facultad de Ciencias, Universidad Nacional Autónoma de México, Ciudad Universitaria, Coyoacán, 04510, Ciudad de México, México</mods:affiliation>
|
||||
<mods:nameIdentifier id="D8C5EFDBA8FCEEA3C7710249775F870A" type="email">orquidiero@gmail.com</mods:nameIdentifier>
|
||||
</mods:name>
|
||||
<mods:name id="90FA7B6C533139B209F1A1782CB7DCF1" type="personal">
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<mods:role id="0F8326722C02267042C8EBE26BAA3443">
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</mods:role>
|
||||
<mods:namePart id="6983243589AFEA98A696F528817C33FF">Pérez-García, Eduardo A.</mods:namePart>
|
||||
<mods:nameIdentifier id="4BDB26FE443BB3FEE210BB8ADDAABF78" type="ORCID">0000-0002-4136-4500</mods:nameIdentifier>
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<mods:affiliation id="81EB171DB01320E23B16837F975168F9">Departamento de Ecología y Recursos Naturales, Facultad de Ciencias, Universidad Nacional Autónoma de México, Ciudad Universitaria, Coyoacán, 04510, Ciudad de México, México</mods:affiliation>
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<mods:nameIdentifier id="98C3725510185B24CD17C289D6C4AE30" type="email">eapg@ciencias.unam.mx</mods:nameIdentifier>
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<mods:title id="A6A661037D111A6CCC4B1828885B4529">Phytotaxa</mods:title>
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<mods:title id="0238663DDB4AF12CE46976871BF9A4FD">Phytotaxa</mods:title>
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<mods:part id="AB95016AA61BD58BFF2254B1D61C7082">
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<mods:date id="85D381428BC3C81B2F259227459260B4">2024</mods:date>
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<mods:number id="079CDB28B7C96DF79A9E576AC1D97845">2024-11-28</mods:number>
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<mods:date id="818909F1BEBEEA888192811124A3F440">2024</mods:date>
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<mods:number id="2F180F2E5FBA80FA16336CB50EF7DDB8">2024-11-28</mods:number>
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<mods:detail id="F7609BD7045C9EBC02E2F1CE7208BC9E" type="volume">
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<mods:number id="BB23567201CCDBFE65E10A31DA74D673">674</mods:number>
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<mods:detail id="5F5E20AD156366E9EAAC38BD7C318A40" type="volume">
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<mods:number id="53A707564F7F06CC403492CDF6D1BE4B">674</mods:number>
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<mods:detail id="75EC214444A5BD9D2923D5E9A0791725" type="issue">
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<mods:number id="F4657CD7276EC92157B7097785D2B747">3</mods:number>
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<mods:number id="30B64ACF4BE53DFD1B958436D04EF98D">3</mods:number>
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<mods:start id="9D8FD48A00A209D889EC8F4BF0952325">265</mods:start>
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<mods:start id="E087E75705FB78C1FAB9A4B6BE5EE9EB">265</mods:start>
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<mods:end id="9193C7A18163B88D4A626E571C2144EB">274</mods:end>
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<mods:url id="898BAAECFD65A4B0B7A6524A0111F8B5">https://doi.org/10.11646/phytotaxa.674.3.3</mods:url>
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<mods:location id="7099108E10C82961FD6FB7E06FB6615B">
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<mods:url id="E9B94D718FD65722FE13A2572FC70D1C">https://doi.org/10.11646/phytotaxa.674.3.3</mods:url>
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</mods:location>
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<mods:classification id="2AF2378B1C4338481353B869E2E0EB21">journal article</mods:classification>
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<mods:identifier id="E9D77318D4628C557256C497799E986B" type="DOI">10.11646/phytotaxa.674.3.3</mods:identifier>
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<mods:identifier id="6449D6DFE9291B15F97771F4DF03DF05" type="ISSN">1179-3163</mods:identifier>
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<mods:identifier id="798895CE69FB98013C98C56305105924" type="Zenodo-Dep">14520837</mods:identifier>
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<mods:classification id="DFB507B849375E67BFBFA467E32D07B1">journal article</mods:classification>
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<mods:identifier id="F6D7E36AD20E51A2BD452C53BF7789F2" type="Zenodo-Dep">14520837</mods:identifier>
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</mods:mods>
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<treatment id="8774C44F6567FFAAFF7E8663FC8FF885" ID-DOI="http://doi.org/10.5281/zenodo.14960706" ID-Zenodo-Dep="14960706" LSID="urn:lsid:plazi:treatment:8774C44F6567FFAAFF7E8663FC8FF885" httpUri="http://treatment.plazi.org/id/8774C44F6567FFAAFF7E8663FC8FF885" lastPageId="6" lastPageNumber="271" pageId="1" pageNumber="266" scope_family="Orchidaceae">
|
||||
<treatment id="8774C44F6567FFAAFF7E8663FC8FF885" ID-DOI="http://doi.org/10.5281/zenodo.14960706" ID-GBIF-Taxon="254900170" ID-Zenodo-Dep="14960706" LSID="urn:lsid:plazi:treatment:8774C44F6567FFAAFF7E8663FC8FF885" httpUri="http://treatment.plazi.org/id/8774C44F6567FFAAFF7E8663FC8FF885" lastPageId="6" lastPageNumber="271" pageId="1" pageNumber="266" scope_family="Orchidaceae">
|
||||
<subSubSection id="47C726D26567FFADFF7E8663FCE7FB7D" box="[136,1007,1140,1166]" pageId="1" pageNumber="266" type="nomenclature">
|
||||
<paragraph id="0F6275596567FFADFF7E8663FCE7FB7D" blockId="1.[136,1007,1140,1166]" box="[136,1007,1140,1166]" pageId="1" pageNumber="266">
|
||||
<taxonomicName id="C8DD0EDA6567FFADFF7E8663FC10FB7D" authority="C.E. Leon-Peralta & E.A. Perez-Garcia" authorityName="C. E. Leon-Peralta & E. A. Perez-Garcia. A. Flowering" authorityYear="2024" box="[136,792,1140,1166]" class="Liliopsida" family="Orchidaceae" genus="Barkeria" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="266" phylum="Tracheophyta" rank="species" species="wixarika" status="sp. nov.">
|
||||
|
|
@ -90,7 +90,7 @@ C.E. León-Peralta & E.A. Pérez-García
|
|||
<subSubSection id="47C726D26567FFADFF7E86D7FDBCFB0A" pageId="1" pageNumber="266" type="materials_examined">
|
||||
<paragraph id="0F6275596567FFADFF7E86D7FDBCFB0A" blockId="1.[136,1452,1216,1382]" pageId="1" pageNumber="266">
|
||||
Type:—
|
||||
<materialsCitation id="BFB57F046567FFADFF2186D7FDB8FB0A" collectedFrom="on Crescentia alata in tropical deciduous forest" collectingDate="2024-03-17" collectionCode="MEXU" collectorName="J. Jones sub C. Leon-Peralta" country="Mexico" elevation="800" location="Santa Maria del Oro Municipality" municipality="Santa Maria del Oro Municipality" pageId="1" pageNumber="266" specimenCount="1" stateProvince="Nayarit" typeStatus="holotype">
|
||||
<materialsCitation id="BFB57F046567FFADFF2186D7FDB8FB0A" ID-GBIF-Occurrence="5074195301" collectedFrom="on Crescentia alata in tropical deciduous forest" collectingDate="2024-03-17" collectionCode="MEXU" collectorName="J. Jones sub C. Leon-Peralta" country="Mexico" elevation="800" location="Santa Maria del Oro Municipality" municipality="Santa Maria del Oro Municipality" pageId="1" pageNumber="266" specimenCount="1" stateProvince="Nayarit" typeStatus="holotype">
|
||||
<collectingCountry id="77CA35C96567FFADFF2186D7FE31FB26" box="[215,313,1216,1237]" name="Mexico" pageId="1" pageNumber="266">MEXICO</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="CD19BBBB6567FFADFEB786D7FE85FB25" box="[321,397,1216,1238]" country="Mexico" name="Nayarit" pageId="1" pageNumber="266">Nayarit</collectingRegion>
|
||||
|
|
@ -365,7 +365,7 @@ and
|
|||
Additional specimens examined (
|
||||
<typeStatus id="D066CBFB6563FFA9FDC4838DFDAAFE47" box="[562,674,410,436]" pageId="5" pageNumber="270" type="paratype">paratypes</typeStatus>
|
||||
):—
|
||||
<materialsCitation id="BFB57F046563FFA9FD3D838DFE68FE2B" collectingDate="1993-08-19" collectionCode="MEXU" collectorName="G. Flores-Franco" country="Mexico" location="Nayarit" pageId="5" pageNumber="270" specimenCode="602891" specimenCount="1" stateProvince="Nayarit" typeStatus="paratype">
|
||||
<materialsCitation id="BFB57F046563FFA9FD3D838DFE68FE2B" ID-GBIF-Occurrence="5074195304" collectingDate="1993-08-19" collectionCode="MEXU" collectorName="G. Flores-Franco" country="Mexico" location="Nayarit" pageId="5" pageNumber="270" specimenCode="602891" specimenCount="1" stateProvince="Nayarit" typeStatus="paratype">
|
||||
<collectingCountry id="77CA35C96563FFA9FD3D838DFC37FE47" box="[715,831,410,436]" name="Mexico" pageId="5" pageNumber="270">MEXICO</collectingCountry>
|
||||
.
|
||||
<collectingRegion id="CD19BBBB6563FFA9FCBE838DFCA8FE47" box="[840,928,410,436]" country="Mexico" name="Nayarit" pageId="5" pageNumber="270">Nayarit</collectingRegion>
|
||||
|
|
@ -385,7 +385,7 @@ et al. 2736
|
|||
]!)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="BFB57F046563FFA9FE9A83A9FBC3FE2A" box="[364,1227,446,473]" collectingDate="1960-09-15" collectingDateMax="1960-09-16" collectingDateMin="1960-09-15" collectionCode="F, MICH" collectorName="R. McVaugh" pageId="5" pageNumber="270" specimenCode="1575058, 1646952" specimenCount="1" typeStatus="paratype">
|
||||
<materialsCitation id="BFB57F046563FFA9FE9A83A9FBC3FE2A" ID-GBIF-Occurrence="5074195303" box="[364,1227,446,473]" collectingDate="1960-09-15" collectingDateMax="1960-09-16" collectingDateMin="1960-09-15" collectionCode="F, MICH" collectorName="R. McVaugh" pageId="5" pageNumber="270" specimenCode="1575058, 1646952" specimenCount="1" typeStatus="paratype">
|
||||
<date id="7B6353996563FFA9FE9A83A9FD7DFE2B" box="[364,629,446,473]" pageId="5" pageNumber="270" value="1960-09-15" valueMax="1960-09-16" valueMin="1960-09-15">
|
||||
<collectingDate id="6B27AA716563FFA9FE9A83A9FD7DFE2B" box="[364,629,446,473]" pageId="5" pageNumber="270" value="1960-09-15" valueMax="1960-09-16" valueMin="1960-09-15">15–16 September 1960</collectingDate>
|
||||
</date>
|
||||
|
|
@ -405,7 +405,7 @@ et al. 2736
|
|||
])
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="BFB57F046563FFA9FB2183A9FD33FE0F" collectingDate="2024-07-20" collectionCode="MEXU" collectorName="C. E. Leon-Peralta" pageId="5" pageNumber="270" specimenCount="1" typeStatus="paratype">
|
||||
<materialsCitation id="BFB57F046563FFA9FB2183A9FD33FE0F" ID-GBIF-Occurrence="5074195302" collectingDate="2024-07-20" collectionCode="MEXU" collectorName="C. E. Leon-Peralta" pageId="5" pageNumber="270" specimenCount="1" typeStatus="paratype">
|
||||
<date id="7B6353996563FFA9FB2183A9FA65FE2B" box="[1239,1389,446,472]" pageId="5" pageNumber="270" value="2024-07-20">
|
||||
<collectingDate id="6B27AA716563FFA9FB2183A9FA65FE2B" box="[1239,1389,446,472]" pageId="5" pageNumber="270" value="2024-07-20">20 July 2024</collectingDate>
|
||||
</date>
|
||||
|
|
@ -422,7 +422,7 @@ et al. 370–373
|
|||
</paragraph>
|
||||
<paragraph id="0F6275596563FFA9FF4B8011FBEFFDD3" blockId="5.[136,1452,158,1048]" box="[189,1255,518,544]" pageId="5" pageNumber="270">
|
||||
Other records:—
|
||||
<materialsCitation id="BFB57F046563FFA9FE8D8011FBEBFDD3" box="[379,1251,518,544]" collectingDate="2023-09-06" collectionCode="AMO" collectorName="W. Medina" country="Mexico" location="Nayarit" pageId="5" pageNumber="270" specimenCount="1" stateProvince="Nayarit">
|
||||
<materialsCitation id="BFB57F046563FFA9FE8D8011FBEBFDD3" ID-GBIF-Occurrence="5074195305" box="[379,1251,518,544]" collectingDate="2023-09-06" collectionCode="AMO" collectorName="W. Medina" country="Mexico" location="Nayarit" pageId="5" pageNumber="270" specimenCount="1" stateProvince="Nayarit">
|
||||
<collectingCountry id="77CA35C96563FFA9FE8D8011FEE6FDD3" box="[379,494,518,544]" name="Mexico" pageId="5" pageNumber="270">MEXICO</collectingCountry>
|
||||
:
|
||||
<collectingRegion id="CD19BBBB6563FFA9FE0E8011FD58FDD3" box="[504,592,518,544]" country="Mexico" name="Nayarit" pageId="5" pageNumber="270">Nayarit</collectingRegion>
|
||||
|
|
|
|||
Loading…
Reference in a new issue