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<mods:title id="1D14639246DAD8FEB105BCB65904B1FD">Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="4F59BBD34EBE70AA70726817D2C9BE2F">Génier, François</mods:namePart>
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<mods:nameIdentifier id="5E0D9367862106F39D0A37B36BDDD076" type="email">enier@nature.ca</mods:nameIdentifier>
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<mods:namePart id="B20E21FC94F8A3BF54D8ACE948E509C1">Gunter, Nicole L.</mods:namePart>
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<mods:affiliation id="D56077E92C25AF0FB32189C2EE6A3806">Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD, 4101 Australia.</mods:affiliation>
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<mods:namePart id="5B261971F77CA4EC9B29F88CE0D65738">Gunter, Nicole L.</mods:namePart>
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<mods:affiliation id="E30D673A9247D35C66A5877F94DC9B1B">Biodiversity and Geosciences Program, Queensland Museum, South Brisbane, QLD, 4101 Australia.</mods:affiliation>
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<mods:namePart id="E225ECFBAF2789F94A54CD1AABAB2325">Saxton, Natalie A.</mods:namePart>
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<mods:affiliation id="E1BB550C9FC64BD2593B89367E23BE56">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. & Corresponding author</mods:affiliation>
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<mods:namePart id="AEAF62F5716D1A53F4629CC70EDBEFF3">Saxton, Natalie A.</mods:namePart>
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<mods:affiliation id="A0AC3605B6B06F0468F8F759A716A09C">Department of Biology, Case Western Reserve University, 2080 Adelbert Road, Cleveland, OH, 44106, USA. & Corresponding author</mods:affiliation>
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<taxonomicName id="4C204D7DFFFE6E12FF4613CBFD96FC59" authority="Saxton & Genier" authorityName="Saxton & Genier" authorityYear="2025" box="[151,597,974,1001]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="fervidus" status="sp. nov.">
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@ -69,13 +69,15 @@ Saxton & Génier
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<figureCitation id="131B2A7BFFFE6E12FF461412FF3BFB81" box="[151,248,1047,1073]" captionStart="FIGURE 4" captionStartId="7.[152,255,934,958]" captionTargetBox="[151,1435,182,906]" captionTargetId="figure-17@7.[151,1435,179,909]" captionTargetPageId="7" captionText="FIGURE 4. Diagnostic characters of Rhytomus. A, dorsal head showing emargination and interocular space (R. howdeni); B, dorsal head showing lack of emargination and interocular space (R. fervidus) C, ventral view of male foreleg (R. latidentatus); D, lateral view of elytra (R. vanus); E, dorsal view of male protibia (R. latidentatus)." figureDoi="http://doi.org/10.5281/zenodo.14746450" httpUri="https://zenodo.org/record/14746450/files/figure.png" pageId="9" pageNumber="96">Figs. 4B</figureCitation>
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@ -89,31 +91,49 @@ urn:lsid:zoobank.org:act:
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<figureCitation id="131B2A7BFFFE6E12FD991412FDB4FB81" box="[584,631,1047,1073]" captionStart="FIGURE 12" captionStartId="18.[152,255,1623,1647]" captionTargetBox="[151,1435,179,1597]" captionTargetId="figure-22@18.[151,1435,179,1597]" captionTargetPageId="18" captionText="FIGURE 12. Characters used for species identification. A, elytral surface with microsculpture (R. latidentatus); B, elytral surface glossy (R. fervidus); C, male metatibial apicointernal tooth in line with posterior tibial edge (R. sodalis); D, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (R. larseni); E, lateral view of strongly convex elytra (R. setulosus); F, lateral view of moderately convex elytra (R. vanus)." figureDoi="http://doi.org/10.5281/zenodo.14746472" httpUri="https://zenodo.org/record/14746472/files/figure.png" pageId="9" pageNumber="96">12B</figureCitation>
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<emphasis id="B954EAECFFFE6E12FF46145FFE7DFBC5" bold="true" box="[151,446,1114,1141]" pageId="9" pageNumber="96">Name-bearing type data.</emphasis>
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<materialsCitation id="3B483CA3FFFE6E12FE15145EFED8FB51" collectedFrom="oak forest dung" collectingDate="1974-07-05" collectingDateMax="1974-07-08" collectingDateMin="1974-07-05" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" determinerName=". N. Saxton & F. Genier" elevation="0" location="Canadian Museum" longitude="144.23334" pageId="9" pageNumber="96" specimenCode="CMNEN 00049283" specimenCount="1" specimenCount-male="1" stateProvince="Western Highlands" typeStatus="holotype">
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<specimenCount id="9D26FD77FFFE6E12FDE2145EFD8BFBC5" box="[563,584,1115,1141]" count="1" pageId="9" pageNumber="96" type="male">♂</specimenCount>
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(CMNC): ||
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<collectingCountry id="F337766EFFFE6E12FD02145EFC23FBC5" box="[723,992,1115,1141]" name="Papua New Guinea" pageId="9" pageNumber="96">PAPUA NEW GUINEA</collectingCountry>
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(
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<collectionCode id="ED31AE3BFFFE6E12FD87145EFD73FBC5" box="[598,688,1115,1141]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="9" pageNumber="96">CMNC</collectionCode>
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): || PAPUA NEW GUINEA | Western highlands, Mt. | Hagen,
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<elevation id="000DD1CDFFFE6E12FA8C145EFA5FFBC5" box="[1373,1436,1115,1141]" pageId="9" pageNumber="96">6000’</elevation>
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<collectingRegion id="49E4F81CFFFE6E12FC20145EFB00FBC5" box="[1009,1219,1115,1141]" country="Papua New Guinea" name="Western Highlands" pageId="9" pageNumber="96">Western highlands</collectingRegion>
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, Mt. | Hagen, 6000’ |
|
||||
<date id="FF9E103EFFFE6E12FF74147BFEF9FB29" box="[165,314,1150,1177]" pageId="9" pageNumber="96" value="1974-07-05" valueMax="1974-07-08" valueMin="1974-07-05">5-8.VII.1974</date>
|
||||
, S.Peck | oak forest dung tps. 24-25 ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049283 || [QR code label]; ||
|
||||
<typeStatus id="549B885CFFFE6E12FDB714A6FD39FB0D" box="[614,762,1187,1213]" pageId="9" pageNumber="96" type="holotype">HOLOTYPE</typeStatus>
|
||||
<specimenCount id="9D26FD77FFFE6E12FCD014A6FCD5FB0D" box="[769,790,1187,1213]" count="1" pageId="9" pageNumber="96" type="male">♂</specimenCount>
|
||||
<date id="FF9E103EFFFE6E12FF74147BFEF9FB29" box="[165,314,1150,1177]" pageId="9" pageNumber="96" value="1974-07-05" valueMax="1974-07-08" valueMin="1974-07-05">
|
||||
<collectingDate id="EFDAE9D6FFFE6E12FF74147BFEF9FB29" box="[165,314,1150,1177]" pageId="9" pageNumber="96" value="1974-07-05" valueMax="1974-07-08" valueMin="1974-07-05">5-8.VII.1974</collectingDate>
|
||||
</date>
|
||||
,
|
||||
<collectorName id="26D55328FFFE6E12FE97147AFE57FB29" box="[326,404,1151,1177]" pageId="9" pageNumber="96">S.Peck</collectorName>
|
||||
|
|
||||
<collectedFrom id="383C7503FFFE6E12FE7B147AFD99FB29" box="[426,602,1151,1177]" pageId="9" pageNumber="96">oak forest dung</collectedFrom>
|
||||
tps. 24-25 ||; ||
|
||||
<location id="8EFF6025FFFE6E12FCDA147AFC1CFB29" LSID="urn:lsid:plazi:treatment:038987E8FFFE6E12FF4613CBFBADF801:8EFF6025FFFE6E12FCDA147AFC1CFB29" box="[779,991,1151,1177]" name="Canadian Museum" pageId="9" pageNumber="96">Canadian Museum</location>
|
||||
of |
|
||||
<location id="8EFF6025FFFE6E12FBC5147AFB9CFB29" LSID="urn:lsid:plazi:treatment:038987E8FFFE6E12FF4613CBFBADF801:8EFF6025FFFE6E12FBC5147AFB9CFB29" box="[1044,1119,1151,1177]" name="Musee" pageId="9" pageNumber="96">Musée</location>
|
||||
canadien de la | NATURE |
|
||||
<specimenCode id="DB869E85FFFE6E12FF4614A6FEB7FB0D" box="[151,372,1187,1213]" pageId="9" pageNumber="96">CMNEN 00049283</specimenCode>
|
||||
|| [QR code label]; || HOLOTYPE ♂ |
|
||||
<taxonomicName id="4C204D7DFFFE6E12FCFA14A6FC5AFB0D" box="[811,921,1187,1213]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="genus">
|
||||
<emphasis id="B954EAECFFFE6E12FCFA14A6FC5AFB0D" box="[811,921,1187,1213]" italics="true" pageId="9" pageNumber="96">Rhytomus</emphasis>
|
||||
<emphasis id="B954EAECFFFE6E12FCFA14A6FC5AFB0D" box="[811,921,1187,1213]" italics="true" pageId="9" pageNumber="96">
|
||||
<location id="8EFF6025FFFE6E12FCFA14A6FC5AFB0D" LSID="urn:lsid:plazi:treatment:038987E8FFFE6E12FF4613CBFBADF801:8EFF6025FFFE6E12FCFA14A6FC5AFB0D" box="[811,921,1187,1213]" name="Rhytomus" pageId="9" pageNumber="96">Rhytomus</location>
|
||||
</emphasis>
|
||||
</taxonomicName>
|
||||
|
|
||||
<taxonomicName id="4C204D7DFFFE6E12FC7E14A6FBCAFB0D" authorityName="Saxton & Genier" authorityYear="2025" box="[943,1033,1187,1213]" class="Insecta" family="Scarabaeidae" genus="Rhytomus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="96" phylum="Arthropoda" rank="species" species="fervidus">
|
||||
<emphasis id="B954EAECFFFE6E12FC7E14A6FBCAFB0D" box="[943,1033,1187,1213]" italics="true" pageId="9" pageNumber="96">fervidus</emphasis>
|
||||
</taxonomicName>
|
||||
| Des. N. Saxton & F. Génier, 2024 || [red card].
|
||||
| Des
|
||||
<determinerName id="7E43DA3AFFFE6E12FB9C14A6FB0AFB0D" box="[1101,1225,1187,1213]" pageId="9" pageNumber="96">. N. Saxton</determinerName>
|
||||
&
|
||||
<determinerName id="7E43DA3AFFFE6E12FB3F14A6FA9BFB0D" box="[1262,1368,1187,1213]" pageId="9" pageNumber="96">F. Génier</determinerName>
|
||||
, 2024 || [red card]
|
||||
</materialsCitation>
|
||||
.
|
||||
</paragraph>
|
||||
</subSubSection>
|
||||
<subSubSection id="C33A6575FFFE6E12FF1614EFFA58FAFD" pageId="9" pageNumber="96" type="materials_examined">
|
||||
<paragraph id="8B9F36FEFFFE6E12FF1614EFFA58FAFD" blockId="9.[151,1437,1114,1969]" pageId="9" pageNumber="96">
|
||||
<emphasis id="B954EAECFFFE6E12FF1614EFFE6CFAB4" bold="true" box="[199,431,1258,1284]" pageId="9" pageNumber="96">Material examined</emphasis>
|
||||
(
|
||||
|
|
@ -121,7 +141,6 @@ urn:lsid:zoobank.org:act:
|
|||
,
|
||||
<specimenCount id="9D26FD77FFFE6E12FDFD14EEFDADFAB5" box="[556,622,1259,1285]" count="9" pageId="9" pageNumber="96" type="female">9 ♀♀</specimenCount>
|
||||
).
|
||||
<materialsCitation id="3B483CA3FFFE6E12FD5C14EFFE38FA99" country="Papua New Guinea" elevation="1829" location="Mount Hagen" pageId="9" pageNumber="96" specimenCount="1" stateProvince="Western Highlands">
|
||||
<emphasis id="B954EAECFFFE6E12FD5C14EFFC77FAB4" bold="true" box="[653,948,1258,1285]" pageId="9" pageNumber="96">
|
||||
<collectingCountry id="F337766EFFFE6E12FD5C14EFFC77FAB4" box="[653,948,1258,1285]" name="Papua New Guinea" pageId="9" pageNumber="96">PAPUA NEW GUINEA</collectingCountry>
|
||||
</emphasis>
|
||||
|
|
@ -135,8 +154,6 @@ urn:lsid:zoobank.org:act:
|
|||
</quantity>
|
||||
, [
|
||||
<geoCoordinate id="EE145039FFFE6E12FE7A150BFE38FA99" box="[427,507,1294,1321]" degrees="05" direction="south" minutes="52" orientation="latitude" pageId="9" pageNumber="96" precision="925" value="-5.866667">05°52’</geoCoordinate>
|
||||
</materialsCitation>
|
||||
<materialsCitation id="3B483CA3FFFE6E12FE2A150BFAA9FA99" box="[507,1386,1294,1321]" collectingDate="1974-07-05" collectingDateMax="1974-07-08" collectingDateMin="1974-07-05" collectorName="S. Peck" country="Papua New Guinea" location="S" longitude="144.23334" pageId="9" pageNumber="96" specimenCount="1" stateProvince="S">
|
||||
<collectingRegion id="49E4F81CFFFE6E12FE2A150BFDC8FA99" box="[507,523,1294,1321]" country="Papua New Guinea" name="S" pageId="9" pageNumber="96">S</collectingRegion>
|
||||
,
|
||||
<geoCoordinate id="EE145039FFFE6E12FDC6150AFD44FA99" box="[535,647,1295,1321]" degrees="144" direction="east" minutes="14" orientation="longitude" pageId="9" pageNumber="96" precision="925" value="144.23334">144°14’E</geoCoordinate>
|
||||
|
|
@ -150,42 +167,40 @@ urn:lsid:zoobank.org:act:
|
|||
<collectorName id="26D55328FFFE6E12FBE9150AFB4CFA99" box="[1080,1167,1295,1321]" pageId="9" pageNumber="96">S. Peck</collectorName>
|
||||
, (
|
||||
<collectingMethod id="52614EE9FFFE6E12FB73150AFADAFA99" box="[1186,1305,1295,1321]" pageId="9" pageNumber="96">dung traps</collectingMethod>
|
||||
24-25)
|
||||
</materialsCitation>
|
||||
—
|
||||
24-25)—
|
||||
<specimenCount id="9D26FD77FFFE6E12FA58150BFA5FFA99" box="[1417,1436,1294,1321]" count="1" pageId="9" pageNumber="96" type="male">♂</specimenCount>
|
||||
<typeStatus id="549B885CFFFE6E12FF461536FF3FFAFD" box="[151,252,1331,1357]" pageId="9" pageNumber="96" type="holotype">holotype</typeStatus>
|
||||
,
|
||||
<specimenCount id="9D26FD77FFFE6E12FED61536FEDFFAFD" box="[263,284,1331,1357]" count="1" pageId="9" pageNumber="96" type="female">♀</specimenCount>
|
||||
<typeStatus id="549B885CFFFE6E12FEF21536FE43FAFD" box="[291,384,1331,1357]" pageId="9" pageNumber="96" type="allotype">allotype</typeStatus>
|
||||
,
|
||||
<materialsCitation id="3B483CA3FFFE6E12FE5A1536FCC4FAFD" box="[395,775,1331,1357]" collectionCode="CMNC" country="Papua New Guinea" pageId="9" pageNumber="96" specimenCount="18" specimenCount-female="6" specimenCount-male="12" stateProvince="S" typeStatus="paratype">
|
||||
<materialsCitation id="3B483CA3FFFE6E12FE5A1536FCC4FAFD" box="[395,775,1331,1357]" collectingDate="1974-07-05" collectingDateMax="1974-07-08" collectingDateMin="1974-07-05" collectionCode="CMNC" collectorName="S. Peck" country="Papua New Guinea" elevation="1829" location="Mount Hagen" longitude="144.23334" pageId="9" pageNumber="96" specimenCount="18" specimenCount-female="6" specimenCount-male="12" stateProvince="S" typeStatus="paratype">
|
||||
<specimenCount id="9D26FD77FFFE6E12FE5A1536FE19FAFD" box="[395,474,1331,1357]" count="12" pageId="9" pageNumber="96" type="male">12 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D26FD77FFFE6E12FE371536FDE6FAFD" box="[486,549,1331,1357]" count="6" pageId="9" pageNumber="96" type="female">6 ♀♀</specimenCount>
|
||||
<typeStatus id="549B885CFFFE6E12FDFD1536FD54FAFD" box="[556,663,1331,1357]" pageId="9" pageNumber="96" type="paratype">paratypes</typeStatus>
|
||||
(
|
||||
<collectionCode id="ED31AE3BFFFE6E12FD761536FD3CFAFD" box="[679,767,1331,1357]" pageId="9" pageNumber="96">CMNC</collectionCode>
|
||||
<collectionCode id="ED31AE3BFFFE6E12FD761536FD3CFAFD" box="[679,767,1331,1357]" collectionName="Canada, Ottawa, Canadian Museum of Nature" pageId="9" pageNumber="96">CMNC</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B483CA3FFFE6E12FCC31536FBAEFAFD" box="[786,1133,1331,1357]" collectionCode="MNHN" country="Papua New Guinea" pageId="9" pageNumber="96" specimenCount="7" specimenCount-female="1" specimenCount-male="6" stateProvince="S" typeStatus="paratype">
|
||||
<materialsCitation id="3B483CA3FFFE6E12FCC31536FBAEFAFD" box="[786,1133,1331,1357]" collectingDate="1974-07-05" collectingDateMax="1974-07-08" collectingDateMin="1974-07-05" collectionCode="MNHN" collectorName="S. Peck" country="Papua New Guinea" elevation="1829" location="Mount Hagen" longitude="144.23334" pageId="9" pageNumber="96" specimenCount="7" specimenCount-female="1" specimenCount-male="6" stateProvince="S" typeStatus="paratype">
|
||||
<specimenCount id="9D26FD77FFFE6E12FCC31536FC90FAFD" box="[786,851,1331,1357]" count="6" pageId="9" pageNumber="96" type="male">6 ♂♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D26FD77FFFE6E12FC8E1536FC4AFAFD" box="[863,905,1331,1357]" count="1" pageId="9" pageNumber="96" type="female">1 ♀</specimenCount>
|
||||
<typeStatus id="549B885CFFFE6E12FC411536FC38FAFD" box="[912,1019,1331,1357]" pageId="9" pageNumber="96" type="paratype">paratypes</typeStatus>
|
||||
(
|
||||
<collectionCode id="ED31AE3BFFFE6E12FBDB1536FBA6FAFD" box="[1034,1125,1331,1357]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="96">MNHN</collectionCode>
|
||||
<collectionCode id="ED31AE3BFFFE6E12FBDB1536FBA6FAFD" box="[1034,1125,1331,1357]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="9" pageNumber="96">MNHN</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
,
|
||||
<materialsCitation id="3B483CA3FFFE6E12FBA91536FA55FAFD" box="[1144,1430,1331,1357]" collectionCode="QM" country="Papua New Guinea" pageId="9" pageNumber="96" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="S" typeStatus="paratype">
|
||||
<materialsCitation id="3B483CA3FFFE6E12FBA91536FA55FAFD" box="[1144,1430,1331,1357]" collectingDate="1974-07-05" collectingDateMax="1974-07-08" collectingDateMin="1974-07-05" collectionCode="QM" collectorName="S. Peck" country="Papua New Guinea" elevation="1829" location="Mount Hagen" longitude="144.23334" pageId="9" pageNumber="96" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="S" typeStatus="paratype">
|
||||
<specimenCount id="9D26FD77FFFE6E12FBA91536FB67FAFD" box="[1144,1188,1331,1357]" count="1" pageId="9" pageNumber="96" type="male">1 ♂</specimenCount>
|
||||
,
|
||||
<specimenCount id="9D26FD77FFFE6E12FB611536FB19FAFD" box="[1200,1242,1331,1357]" count="1" pageId="9" pageNumber="96" type="female">1 ♀</specimenCount>
|
||||
<typeStatus id="549B885CFFFE6E12FB301536FA8FFAFD" box="[1249,1356,1331,1357]" pageId="9" pageNumber="96" type="paratype">paratypes</typeStatus>
|
||||
(
|
||||
<collectionCode id="ED31AE3BFFFE6E12FA8D1536FA4CFAFD" box="[1372,1423,1331,1357]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="9" pageNumber="96" type="Museum">QM</collectionCode>
|
||||
<collectionCode id="ED31AE3BFFFE6E12FA8D1536FA4CFAFD" box="[1372,1423,1331,1357]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="9" pageNumber="96" type="Museum">QM</collectionCode>
|
||||
)
|
||||
</materialsCitation>
|
||||
.
|
||||
|
|
|
|||
Loading…
Reference in a new issue