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<document id="62399FB09A105758890B9B8C5096F5BB" ID-DOI="10.5194/fr-22-51-2019" ID-ISSN="2193-0074" ID-Zenodo-Dep="11359653" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1716846063762" checkinUser="felipe" docAuthor="Wichard, Wilfried &amp; Neumann, Christian" docDate="2019" docId="4E126219FFB2FFE1E9E6FDDCFAB1FEFF" docLanguage="en" docName="FossRec.22.2.51-56.pdf" docOrigin="Fossil Record 22 (2)" docSource="http://dx.doi.org/10.5194/fr-22-51-2019" docStyle="DocumentStyle:1B3764FFAC190C6D25C29F616DE4358D.2:FossRec.2014-2021.journal_article" docStyleId="1B3764FFAC190C6D25C29F616DE4358D" docStyleName="FossRec.2014-2021.journal_article" docStyleVersion="2" docTitle="Dysoneuridae Sukatsheva 1968" docType="treatment" docVersion="3" lastPageNumber="52" masterDocId="B22B1A61FFB3FFE0E957FFA3FFC5FFB8" masterDocTitle="A new bizarre dysoneurid species (Insecta, Trichoptera) in Burmese amber" masterLastPageNumber="56" masterPageNumber="51" pageNumber="52" updateTime="1726170637509" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8CE5B930188CB32D591708F6EFA48FA4">A new bizarre dysoneurid species (Insecta, Trichoptera) in Burmese amber</mods:title>
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<mods:namePart id="AC4077621FF965872CA5CE3A26C09F61">Wichard, Wilfried</mods:namePart>
<mods:affiliation id="14F1C6B797D614B1BF17EA0AFEECC309">Christian Neumann &amp; Institute of Biology and its Didactic, University of Cologne, Herbert Lewinstrasse 2, 50931 Cologne, Germany</mods:affiliation>
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<mods:namePart id="023A155F6DC2DB7AB5BAB49CD85510A1">Neumann, Christian</mods:namePart>
<mods:affiliation id="091AB8C953B5D77D90342313DBBD30E3">Museum für Naturkunde - Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany</mods:affiliation>
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<treatment id="4E126219FFB2FFE1E9E6FDDCFAB1FEFF" LSID="urn:lsid:plazi:treatment:4E126219FFB2FFE1E9E6FDDCFAB1FEFF" httpUri="http://treatment.plazi.org/id/4E126219FFB2FFE1E9E6FDDCFAB1FEFF" lastPageNumber="52" pageId="1" pageNumber="52">
<subSubSection id="8EA18084FFB2FFE1E9E6FDDCFDA5FD2E" box="[177,608,638,662]" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="C604D30FFFB2FFE1E9E6FDDCFDA5FD2E" blockId="1.[177,608,638,662]" box="[177,608,638,662]" pageId="1" pageNumber="52">
<heading id="9D4C6463FFB2FFE1E9E6FDDCFDA5FD2E" box="[177,608,638,662]" fontSize="10" level="2" pageId="1" pageNumber="52" reason="2">
Family
<taxonomicName id="01BBA88CFFB2FFE1E854FDDDFDA5FD2E" authority="Sukatsheva, 1968" authorityName="Sukatsheva" authorityYear="1968" box="[259,608,638,662]" class="Insecta" family="Dysoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">
<emphasis id="F4CF0F1DFFB2FFE1E854FDDDFE5AFD2E" bold="true" box="[259,415,638,662]" pageId="1" pageNumber="52">Dysoneuridae</emphasis>
<bibRefCitation id="A22AAEFEFFB2FFE1E8F1FDDCFDA5FD2E" author="Sukatsheva, I. D." box="[422,608,639,662]" pageId="1" pageNumber="52" pagination="175 - 179" refId="ref3042" refString="Sukatsheva, I. D.: New Jurassic Caddisflies from the Kara Tau (Trichoptera), Yurskie nasekomye Karatau, 175 - 179, 1968 (in Russian)." type="book chapter" year="1968">Sukatsheva, 1968</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="8EA18084FFB2FFE1E9E6FD19FEF3FD49" pageId="1" pageNumber="52" type="materials_examined">
<paragraph id="C604D30FFFB2FFE1E9E6FD19FEF3FD49" blockId="1.[177,766,698,753]" pageId="1" pageNumber="52">
<typeStatus id="19006DADFFB2FFE1E9E6FD19FF20FD69" box="[177,229,698,721]" pageId="1" pageNumber="52">Type</typeStatus>
genus and species:
<taxonomicName id="01BBA88CFFB2FFE1E89DFD19FEF7FD49" authority="Sukatsheva, 1968" authorityName="Sukatsheva" authorityYear="1968" class="Insecta" family="Dysoneuridae" genus="Dysoneura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="trifurcata">
<emphasis id="F4CF0F1DFFB2FFE1E89DFD19FD6BFD6A" box="[458,686,698,722]" italics="true" pageId="1" pageNumber="52">Dysoneura trifurcata</emphasis>
<bibRefCitation id="A22AAEFEFFB2FFE1EBEEFD19FEF7FD49" author="Sukatsheva, I. D." pageId="1" pageNumber="52" pagination="175 - 179" refId="ref3042" refString="Sukatsheva, I. D.: New Jurassic Caddisflies from the Kara Tau (Trichoptera), Yurskie nasekomye Karatau, 175 - 179, 1968 (in Russian)." type="book chapter" year="1968">Sukatsheva, 1968</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="8EA18084FFB2FFE1E92BFCBBFF06FA12" pageId="1" pageNumber="52" type="diagnosis">
<paragraph id="C604D30FFFB2FFE1E92BFCBBFEFEFC88" blockId="1.[124,315,792,816]" box="[124,315,792,816]" pageId="1" pageNumber="52">
<emphasis id="F4CF0F1DFFB2FFE1E92BFCBBFEFEFC88" bold="true" box="[124,315,792,816]" pageId="1" pageNumber="52">Family diagnosis</emphasis>
</paragraph>
<paragraph id="C604D30FFFB2FFE1E92BFCF6FE50FB73" blockId="1.[124,767,853,1450]" pageId="1" pageNumber="52">
<taxonomicName id="01BBA88CFFB2FFE1E92BFCF6FED5FCD4" authorityName="Sukatsheva" authorityYear="1968" box="[124,272,853,876]" class="Insecta" family="Dysoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">Dysoneuridae</taxonomicName>
clearly distinguished from all extant and extinct families of
<taxonomicName id="01BBA88CFFB2FFE1E87AFCD6FE6DFC34" authorityName="Kirby" authorityYear="1815" box="[301,424,885,908]" class="Insecta" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
by the tibial spur formula 2/2/
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combination with the following fore- and hind wing venational characters. In forewing, forks I and
<collectionCode id="A0AA4BCAFFB2FFE1EB31FC16FD47FC74" box="[614,642,949,972]" pageId="1" pageNumber="52">IV</collectionCode>
absent and forks
<collectionCode id="A0AA4BCAFFB2FFE1E9E8FC76FF16FC54" box="[191,211,981,1004]" pageId="1" pageNumber="52">II</collectionCode>
,
<collectionCode id="A0AA4BCAFFB2FFE1E9B3FC76FEC4FC54" box="[228,257,981,1004]" pageId="1" pageNumber="52">III</collectionCode>
, and
<collectionCode id="A0AA4BCAFFB2FFE1E812FC76FE9DFC54" box="[325,344,981,1004]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="52" type="Museum">V</collectionCode>
present; the nygma absent; discoidal and median cells closed or open; the thyridial cell long and closed; Cu2 reaches the desclerotized line (crossvein) running from the wing margin to Cu1b. In the hind wing, forks I,
<collectionCode id="A0AA4BCAFFB2FFE1E92BFBF7FF5CFBD3" box="[124,153,1108,1131]" pageId="1" pageNumber="52">III</collectionCode>
, and
<collectionCode id="A0AA4BCAFFB2FFE1E984FBF7FF23FBD3" box="[211,230,1108,1131]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="52" type="Museum">V</collectionCode>
are present; and forks
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and
<collectionCode id="A0AA4BCAFFB2FFE1EB72FBF7FD84FBD3" box="[549,577,1108,1131]" pageId="1" pageNumber="52">IV</collectionCode>
and the discoidal cell are absent. Antennae about as long as the forewings or longer; the scapus as long as the head or longer, often bearing a brush of long dark setae.
</paragraph>
<paragraph id="C604D30FFFB2FFE1E9C0FB77FF06FA12" blockId="1.[124,767,853,1450]" pageId="1" pageNumber="52">
The family
<taxonomicName id="01BBA88CFFB2FFE1E849FB77FE77FB53" authorityName="Sukatsheva" authorityYear="1968" box="[286,434,1236,1259]" class="Insecta" family="Dysoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">Dysoneuridae</taxonomicName>
belongs to superfamily
<taxonomicName id="01BBA88CFFB2FFE1EB9CFB77FED0FAB3" pageId="1" pageNumber="52" rank="superFamily" superFamily="Sericostomatoidea">Sericostomatoidea</taxonomicName>
indicated by the absence of ocelli, with the terminal maxillary and labial segments being nonannulated and inflexible and the tibial spur formula being 2/2/4. The
<taxonomicName id="01BBA88CFFB2FFE1E92BFAF0FED5FAD2" authorityName="Sukatsheva" authorityYear="1968" box="[124,272,1363,1386]" class="Insecta" family="Dysoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">Dysoneuridae</taxonomicName>
are distinct from all other families of
<taxonomicName id="01BBA88CFFB2FFE1EBF0FAF0FF37FA32" pageId="1" pageNumber="55" rank="superFamily" superFamily="Sericostomatoidea">Sericostomatoidea</taxonomicName>
by their typical wing venation of fore- and hind wings.
</paragraph>
</subSubSection>
<subSubSection id="8EA18084FFB2FFE1E92BFA6DFAB1FEFF" pageId="1" pageNumber="52" type="discussion">
<paragraph id="C604D30FFFB2FFE1E92BFA6DFF26FA5E" blockId="1.[124,227,1486,1510]" box="[124,227,1486,1510]" pageId="1" pageNumber="52">
<emphasis id="F4CF0F1DFFB2FFE1E92BFA6DFF26FA5E" bold="true" box="[124,227,1486,1510]" pageId="1" pageNumber="52">Remarks</emphasis>
</paragraph>
<paragraph id="C604D30FFFB2FFE1E92BF9A8FAB1FEFF" blockId="1.[124,767,1547,1921]" lastBlockId="1.[819,1462,176,327]" pageId="1" pageNumber="52">
The family
<taxonomicName id="01BBA88CFFB2FFE1E84BF9A8FE75F99A" authorityName="Sukatsheva" authorityYear="1968" box="[284,432,1547,1570]" class="Insecta" family="Dysoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">Dysoneuridae</taxonomicName>
as established by
<bibRefCitation id="A22AAEFEFFB2FFE1EBEEF9A8FED5F9FA" author="Sukatsheva, I. D." pageId="1" pageNumber="52" pagination="175 - 179" refId="ref3042" refString="Sukatsheva, I. D.: New Jurassic Caddisflies from the Kara Tau (Trichoptera), Yurskie nasekomye Karatau, 175 - 179, 1968 (in Russian)." type="book chapter" year="1968">Sukatsheva (1968)</bibRefCitation>
is based merely on the venation of an isolated forewing preserved in sedimentary rocks from the Upper Jurassic in southern
<collectingCountry id="BEAC939FFFB2FFE1E88BF9C8FD9EF93A" box="[476,603,1643,1666]" name="Kazakhstan" pageId="1" pageNumber="52">Kazakhstan</collectingCountry>
. The forewing venation was interpreted as a reduced venation because of the absence of apical forks I and
<collectionCode id="A0AA4BCAFFB2FFE1E88BF908FE35F97A" box="[476,496,1707,1730]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="52" type="Museum">V</collectionCode>
; in addition, Cu2 reaches a desclerotized line running from the wing margin (arculus) to Cu1b; both characteristics correspond approximately to those of the wings of the family
<taxonomicName id="01BBA88CFFB2FFE1E88FF8A9FE85F8F9" authority="(Sukatsheva and Vassilenko, 2013)" baseAuthorityName="Sukatsheva and Vassilenko" baseAuthorityYear="2013" class="Insecta" family="Kokiriidae" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">
Kokiriidae (
<bibRefCitation id="A22AAEFEFFB2FFE1EB00F8A9FEFDF8F9" author="Sukatsheva, I. D. &amp; Vassilenko, D. V." pageId="1" pageNumber="52" pagination="57 - 63" refId="ref3074" refString="Sukatsheva, I. D. and Vassilenko, D. V.: New Taxa of Caddisflies (Insecta, Trichoptera) with reduced forewing venation from the Mesozoic of Asia, Paleontologicheskii Zh., 2013, 57 - 63, 2013." type="journal article" year="2013">Sukatsheva and Vassilenko, 2013</bibRefCitation>
)
</taxonomicName>
. However, for a complete and significant description and classification of
<taxonomicName id="01BBA88CFFB2FFE1E88DF8E9FD9DF8D9" authorityName="Kirby" authorityYear="1815" box="[474,600,1866,1889]" class="Insecta" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
, a complete set of morphological characters is imperatively needed. In the case of the family
<taxonomicName id="01BBA88CFFB2FFE1EAAFFF13FB4BFF7F" authorityName="Sukatsheva" authorityYear="1968" box="[1016,1166,176,199]" class="Insecta" family="Dysoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">Dysoneuridae</taxonomicName>
, this became recently available by excellently preserved specimens from Cretaceous Burmese amber, subsequently leading to a redescription of the family
<taxonomicName id="01BBA88CFFB2FFE1EA8EFEB3FBA8FE9F" authorityName="Sukatsheva" authorityYear="1968" box="[985,1133,272,295]" class="Insecta" family="Dysoneuridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="family">Dysoneuridae</taxonomicName>
and their assignment to the superfamily
<taxonomicName id="01BBA88CFFB2FFE1EAEEFE93FAB5FEFF" authority="(Wichard et al., 2018)" baseAuthorityName="Wichard" baseAuthorityYear="2018" box="[953,1392,304,327]" class="Insecta" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="superFamily" superFamily="Sericostomatoidea">
Sericostomatoidea (
<bibRefCitation id="A22AAEFEFFB2FFE1EDDCFE93FAADFEFF" author="Wichard, W. &amp; Neumann, C. &amp; Muller, P. &amp; Wang, B." box="[1163,1384,304,327]" pageId="1" pageNumber="52" pagination="138 - 146" refId="ref3172" refString="Wichard, W., Neumann, C., Muller, P., and Wang, B.: Family Dysoneuridae (Insecta, Trichoptera) in Cretaceous Burmese amber, Cretaceous Res., 82, 138 - 146, 2018." type="journal article" year="2018">Wichard et al., 2018</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>

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<mods:title id="9F6DA34F8F909C6845E37E8E0657785D">A new bizarre dysoneurid species (Insecta, Trichoptera) in Burmese amber</mods:title>
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<mods:affiliation id="45B340C72E12C8C3D7A484A4EF9FD4CB">Christian Neumann &amp; Institute of Biology and its Didactic, University of Cologne, Herbert Lewinstrasse 2, 50931 Cologne, Germany</mods:affiliation>
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<taxonomicName id="01BBA88CFFB2FFE1EA3EFECEFB0FFE3C" authority="Wichard et al., 2018" authorityName="Wichard" authorityYear="2018" box="[873,1226,365,389]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="F4CF0F1DFFB2FFE1EA3EFECEFC29FE3D" box="[873,1004,365,389]" italics="true" pageId="1" pageNumber="52">Cretapsyche</emphasis>
<bibRefCitation id="A22AAEFEFFB2FFE1EAA4FECEFB0FFE3C" author="Wichard, W. &amp; Neumann, C. &amp; Muller, P. &amp; Wang, B." box="[1011,1226,365,388]" pageId="1" pageNumber="52" pagination="138 - 146" refId="ref3172" refString="Wichard, W., Neumann, C., Muller, P., and Wang, B.: Family Dysoneuridae (Insecta, Trichoptera) in Cretaceous Burmese amber, Cretaceous Res., 82, 138 - 146, 2018." type="journal article" year="2018">Wichard et al., 2018</bibRefCitation>
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</paragraph>
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<typeStatus id="19006DADFFB2FFE1EA3EFE05FC58FE05" box="[873,925,422,445]" pageId="1" pageNumber="52">Type</typeStatus>
species:
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<emphasis id="F4CF0F1DFFB2FFE1EAA9FE06FB17FE05" box="[1022,1234,421,445]" italics="true" pageId="1" pageNumber="52">Cretapsyche circula</emphasis>
<bibRefCitation id="A22AAEFEFFB2FFE1ED8EFE05FA75FE05" author="Wichard, W. &amp; Neumann, C. &amp; Muller, P. &amp; Wang, B." box="[1241,1456,422,445]" pageId="1" pageNumber="52" pagination="138 - 146" refId="ref3172" refString="Wichard, W., Neumann, C., Muller, P., and Wang, B.: Family Dysoneuridae (Insecta, Trichoptera) in Cretaceous Burmese amber, Cretaceous Res., 82, 138 - 146, 2018." type="journal article" year="2018">Wichard et al., 2018</bibRefCitation>
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</paragraph>
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<paragraph id="C604D30FFFB2FFE1EA3EFE7CFBE4FDCD" blockId="1.[873,1462,479,629]" pageId="1" pageNumber="52">Genus diagnosis: the five-segmented maxillary palps give a clear and striking characteristic – second segment longest, longer than the following three segments together, which are successively smaller and shorter; fifth segment shortest.</paragraph>
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<paragraph id="C604D30FFFB2FFE1EA3EFD34FB70FD16" blockId="1.[873,1205,663,687]" box="[873,1205,663,687]" pageId="1" pageNumber="52">
<heading id="9D4C6463FFB2FFE1EA3EFD34FB70FD16" box="[873,1205,663,687]" fontSize="10" level="2" pageId="1" pageNumber="52" reason="7">
<taxonomicName id="01BBA88CFFB2FFE1EA3EFD34FB74FD16" authority="Wichard &amp; Neumann, 2019" authorityName="Wichard &amp; Neumann" authorityYear="2019" box="[873,1201,663,687]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="palpinova" status="sp. nov.">
<emphasis id="F4CF0F1DFFB2FFE1EA3EFD34FB98FD17" box="[873,1117,663,687]" italics="true" pageId="1" pageNumber="52">Cretapsyche palpinova</emphasis>
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<paragraph id="C604D30FFFB2FFE1EA3EFD73FC17FD5F" blockId="1.[873,978,720,743]" box="[873,978,720,743]" pageId="1" pageNumber="52">
<figureCitation id="5E80CF8AFFB2FFE1EA3EFD73FC0BFD5F" box="[873,974,720,743]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStartId-0="2.[124,191,1452,1474]" captionStartId-1="3.[124,191,1764,1786]" captionTargetBox-0="[150,1435,177,1420]" captionTargetBox-1="[150,1435,189,1733]" captionTargetId-0="figure-11@2.[150,1435,177,1420]" captionTargetId-1="figure-11@3.[150,1435,189,1733]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figure 1. Cretapsyche palpinova sp. nov. (a) Male in ventral view. (b) Forewing venation: II, III, and V denote apical forks II, III, and V; TC denotes thyridial cell." captionText-1="Figure 2. Cretapsyche palpinova sp. nov. (a) Drawing of the scapi, pedicelli, and first antennal segments; the five-segmented maxillary palps (max palp), the three-segmented labial palps (lab palp); and a pair of pilifers arising from the maxillary palps. (b) Head with palps and antennae in ventral view. (c) Male genitalia in ventral view. (d) Drawing of the male genitalia in ventral view with the inferior appendages (inf ap) consisting of the basal coxopodite and the apical harpago, with the preanal appendages (pre ap) and with tergum X divided in two apical lobes (X ap)." figureDoi-0="http://doi.org/10.5281/zenodo.11359655" figureDoi-1="http://doi.org/10.5281/zenodo.11359657" httpUri-0="https://zenodo.org/record/11359655/files/figure.png" httpUri-1="https://zenodo.org/record/11359657/files/figure.png" pageId="1" pageNumber="52">Figs. 1–2</figureCitation>
.
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<paragraph id="C604D30FFFB2FFE1EA64FCAFFC5FFC9C" blockId="1.[819,922,780,804]" box="[819,922,780,804]" pageId="1" pageNumber="52">
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Kept in the amber collection of the Museum für Naturkunde,
<collectingRegion id="047F1DEDFFB2FFE1EA64FCCBFCB0FCC7" box="[819,885,872,895]" country="Germany" name="Berlin" pageId="1" pageNumber="52">Berlin</collectingRegion>
(MfN:
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inventory number MB.I. 7340 coll., former Patrick Müller collection, BUB 2587). Fossil from
<collectingMunicipality id="26604975FFB2FFE1EA64FC0BFC33FC07" box="[819,1014,936,959]" pageId="1" pageNumber="52">Upper Cretaceous</collectingMunicipality>
,
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amber, body well preserved; forewing in dorsal view, hind wing covered dorsally by forewing. Antennae and legs present. Male genitalia visible in ventral view.
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<emphasis id="F4CF0F1DFFB2FFE1EA64FBE1FC6BFBE2" bold="true" box="[819,942,1090,1114]" pageId="1" pageNumber="52">Etymology</emphasis>
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<heading id="9D4C6463FFB2FFE1EA64FBDDFBCAFB0D" centered="true" fontSize="10" level="2" pageId="1" pageNumber="52" reason="5">
<taxonomicName id="01BBA88CFFB2FFE1EA64FBDDFB45FB2D" authority="Wichard &amp; Neumann, 2019" authorityName="Wichard &amp; Neumann" authorityYear="2019" box="[819,1152,1150,1174]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="palpinova" status="sp. nov.">
<emphasis id="F4CF0F1DFFB2FFE1EA64FBDDFBE2FB2E" box="[819,1063,1150,1174]" italics="true" pageId="1" pageNumber="52">Cretapsyche palpinova</emphasis>
<taxonomicNameLabel id="EFFCB266FFB2FFE1ED79FBDDFB45FB2D" box="[1070,1152,1150,1173]" pageId="1" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
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is named for its novel maxillary and labial palpi.
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<emphasis id="F4CF0F1DFFB2FFE1EA64FB7AFC65FB49" bold="true" box="[819,928,1241,1265]" pageId="1" pageNumber="52">Diagnosis</emphasis>
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<paragraph id="C604D30FFFB2FFE1EA64FAB6FA72F9F3" blockId="1.[819,1463,1301,1611]" pageId="1" pageNumber="52">Scapus about as long as the head and bearing a brush of long setae; the pedicellus short, thickened at end; first flagellomere almost twice as long as the following flagellomeres and basally thickened. The new species is distinguished from other dysoneurid species by the maxillary and labial palps: maxillary palps five-segmented; second segment longest, narrow, and slightly bent; third segment hairy, pin-shaped pendant (pilifer); third to fifth segments successively decreasing in size. Labial palps three-segmented; the terminal segment long, oval, and flattened, with an apicolateral bulge.</paragraph>
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<emphasis id="F4CF0F1DFFB2FFE1EA64F9CDFC73F93E" bold="true" box="[819,950,1646,1670]" pageId="1" pageNumber="52">Description</emphasis>
</paragraph>
<paragraph id="C604D30FFFB2FFE2EA64F909FDF2F8A0" blockId="1.[819,1462,1706,1921]" lastBlockId="2.[124,767,1570,1912]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<emphasis id="F4CF0F1DFFB2FFE1EA64F909FB90F97A" box="[819,1109,1706,1730]" italics="true" pageId="1" pageNumber="52">Male head without ocelli.</emphasis>
Each flagellum about
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in length, consisting of 26 rod-shaped flagellomeres; scapus strong, narrow, distally slightly broader than basally, about 0.5 mm in length and slightly longer than head, hairy, apically with tuft of hair; pedicellus short, thickened at ends; first flagellomere twice as long as following flagellomeres, basally thickened. Maxillary palpus five-segmented: first segment long but shorter than second segment; second segment longest, slim and slightly curved, longer than third, fourth, and fifth segments together. Rod-shaped appendage (pilifer) originating from third segment. Labial palpus three-segmented: first segment longest, almost as long as second and third segments together; third segment about as long as second segment but wider, flattened, oval, rounded margin, apicolaterally with small rounded tongue.
</paragraph>
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<emphasis id="F4CF0F1DFFB1FFE2E92BFA0FFF12FA7A" bold="true" box="[124,215,1452,1474]" pageId="2" pageNumber="53">Figure 1.</emphasis>
<taxonomicName id="01BBA88CFFB1FFE2E98BFA0FFE72FA79" authority="Wichard &amp; Neumann, 2019" authorityName="Wichard &amp; Neumann" authorityYear="2019" box="[220,439,1452,1473]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="palpinova" status="sp. nov.">
<emphasis id="F4CF0F1DFFB1FFE2E98BFA0FFE72FA79" box="[220,439,1452,1473]" italics="true" pageId="2" pageNumber="53">Cretapsyche palpinova</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFFCB266FFB1FFE2E8EAFA0FFDC2FA79" box="[445,519,1452,1473]" pageId="2" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="F4CF0F1DFFB1FFE2EB5CFA0FFDE2FA7A" bold="true" box="[523,551,1452,1474]" pageId="2" pageNumber="53">(a)</emphasis>
Male in ventral view.
<emphasis id="F4CF0F1DFFB1FFE2EBACFA0FFCDDFA7A" bold="true" box="[763,792,1452,1474]" pageId="2" pageNumber="53">(b)</emphasis>
Forewing venation: II, III, and V denote apical forks II, III, and V; TC denotes thyridial cell.
</paragraph>
</caption>
<paragraph id="C604D30FFFB1FFE2E9C0F882FAE5F880" blockId="2.[124,767,1570,1912]" lastBlockId="2.[819,1463,1570,1912]" pageId="2" pageNumber="53">
<emphasis id="F4CF0F1DFFB1FFE2E9C0F882FD99F881" box="[151,604,1825,1849]" italics="true" pageId="2" pageNumber="53">
Forewings each
<quantity id="01437EEAFFB1FFE2E81EF882FE44F881" box="[329,385,1825,1849]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="53" unit="mm" value="3.0">3 mm</quantity>
in length (
<figureCitation id="5E80CF8AFFB1FFE2E8AAF882FD8BF881" box="[509,590,1825,1849]" captionStart="Figure 1" captionStartId="2.[124,191,1452,1474]" captionTargetBox="[150,1435,177,1420]" captionTargetId="figure-11@2.[150,1435,177,1420]" captionTargetPageId="2" captionText="Figure 1. Cretapsyche palpinova sp. nov. (a) Male in ventral view. (b) Forewing venation: II, III, and V denote apical forks II, III, and V; TC denotes thyridial cell." figureDoi="http://doi.org/10.5281/zenodo.11359655" httpUri="https://zenodo.org/record/11359655/files/figure.png" pageId="2" pageNumber="53">Fig. 1b</figureCitation>
).
</emphasis>
Apical forks I and
<collectionCode id="A0AA4BCAFFB1FFE2E9FDF8E2FF03F8E0" box="[170,198,1857,1880]" pageId="2" pageNumber="53">IV</collectionCode>
absent;
<collectionCode id="A0AA4BCAFFB1FFE2E875F8E2FEF3F8E0" box="[290,310,1857,1880]" pageId="2" pageNumber="53">II</collectionCode>
,
<collectionCode id="A0AA4BCAFFB1FFE2E815F8E2FE9AF8E0" box="[322,351,1857,1880]" pageId="2" pageNumber="53">III</collectionCode>
, and
<collectionCode id="A0AA4BCAFFB1FFE2E8CEF8E2FE69F8E0" box="[409,428,1857,1880]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="53" type="Museum">V</collectionCode>
present; fork
<collectionCode id="A0AA4BCAFFB1FFE2EB12F8E2FDA5F8E0" box="[581,608,1857,1880]" pageId="2" pageNumber="53">III</collectionCode>
petiolate; fork
<collectionCode id="A0AA4BCAFFB1FFE2E92BF8C2FF4AF8C0" box="[124,143,1889,1912]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="53" type="Museum">V</collectionCode>
probably slightly petiolate. Crossveins
<emphasis id="F4CF0F1DFFB1FFE2EB6FF8C3FD87F8C0" box="[568,578,1888,1912]" italics="true" pageId="2" pageNumber="53">r</emphasis>
and
<emphasis id="F4CF0F1DFFB1FFE2EB2DF8C3FD41F8C0" box="[634,644,1888,1912]" italics="true" pageId="2" pageNumber="53">s</emphasis>
not clearly visible,
<emphasis id="F4CF0F1DFFB1FFE2EADAF981FC72F982" box="[909,951,1570,1594]" italics="true" pageId="2" pageNumber="53">r -m</emphasis>
short and aligned diagonally,
<emphasis id="F4CF0F1DFFB1FFE2EC46F981FAE3F982" box="[1297,1318,1570,1594]" italics="true" pageId="2" pageNumber="53">m</emphasis>
absent. Discoidal and median cells open, thyridial cell long and closed. Cu2 running to Cu1b and reaching desclerotized crossvein connecting Cu1b to margin; this crossvein not clearly visible. Hind wings smaller than forewings, each with wing venation and setation as follows (in ventral view, so far as visible): fork
<collectionCode id="A0AA4BCAFFB1FFE2EA3FF942FC46F940" box="[872,899,1761,1784]" pageId="2" pageNumber="53">III</collectionCode>
clearly visible, petiolate; crossvein
<emphasis id="F4CF0F1DFFB1FFE2EC5FF942FAF6F941" box="[1288,1331,1761,1785]" italics="true" pageId="2" pageNumber="53">r -m</emphasis>
also clearly visible; fork
<collectionCode id="A0AA4BCAFFB1FFE2EAEDF8A2FC09F8A0" box="[954,972,1793,1816]" pageId="2" pageNumber="53">II</collectionCode>
absent; forks I and
<collectionCode id="A0AA4BCAFFB1FFE2EDCBF8A2FB6AF8A0" box="[1180,1199,1793,1816]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="53" type="Museum">V</collectionCode>
probably present; dorsal surface covered with small scale-shaped hairs.
</paragraph>
<paragraph id="C604D30FFFB1FFE4EA19F8E2FE2EFE5E" blockId="2.[819,1463,1570,1912]" lastBlockId="4.[124,767,176,486]" lastPageId="4" lastPageNumber="55" pageId="2" pageNumber="53">
<emphasis id="F4CF0F1DFFB1FFE2EA19F8E2FA4EF8E1" box="[846,1419,1857,1881]" italics="true" pageId="2" pageNumber="53">
Male genitalia visible in ventral view (
<figureCitation id="5E80CF8AFFB1FFE2EC5CF8E2FABBF8E1" box="[1291,1406,1857,1881]" captionStart="Figure 2" captionStartId="3.[124,191,1764,1786]" captionTargetBox="[150,1435,189,1733]" captionTargetId="figure-11@3.[150,1435,189,1733]" captionTargetPageId="3" captionText="Figure 2. Cretapsyche palpinova sp. nov. (a) Drawing of the scapi, pedicelli, and first antennal segments; the five-segmented maxillary palps (max palp), the three-segmented labial palps (lab palp); and a pair of pilifers arising from the maxillary palps. (b) Head with palps and antennae in ventral view. (c) Male genitalia in ventral view. (d) Drawing of the male genitalia in ventral view with the inferior appendages (inf ap) consisting of the basal coxopodite and the apical harpago, with the preanal appendages (pre ap) and with tergum X divided in two apical lobes (X ap)." figureDoi="http://doi.org/10.5281/zenodo.11359657" httpUri="https://zenodo.org/record/11359657/files/figure.png" pageId="2" pageNumber="53">Fig. 2c, d</figureCitation>
).
</emphasis>
Inferior appendages two-segmented, each consisting of coxopodite and harpago; small subtriangular accessory appendage basoventrally; coxopodite strong, about 0.2 mm in length, about as long as harpago; harpago originating from coxopodite, basally narrow, thicker mesally near middle, and tapered apically; curved mesad, enclosing central phallic apparatus. Tergum X with mesodorsal lobe divided apparently into two small parallel processes covering phallic apparatus dorsally. In ventral view dorsolateral preanal appendages also deeply behind central phallic apparatus, each forming small, elongate, triangular spatula.
</paragraph>
</subSubSection>
</treatment>
</document>

128
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<mods:title id="8D68C60D2FA646D62071CC3A8929871B">A new bizarre dysoneurid species (Insecta, Trichoptera) in Burmese amber</mods:title>
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<mods:namePart id="33FA75D54A67BAC77C1737CCFCDEB490">Wichard, Wilfried</mods:namePart>
<mods:affiliation id="420CB3F136B9A0E4813966A272F67EDF">Christian Neumann &amp; Institute of Biology and its Didactic, University of Cologne, Herbert Lewinstrasse 2, 50931 Cologne, Germany</mods:affiliation>
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<mods:namePart id="FE145AD8928310200EEBFF0A501BB881">Neumann, Christian</mods:namePart>
<mods:affiliation id="FCFFA90285CB3A1D10C9FBCAB3896136">Museum für Naturkunde - Leibniz Institute for Evolution and Biodiversity Science, Invalidenstrasse 43, 10115 Berlin, Germany</mods:affiliation>
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<treatment id="4E126219FFB7FFE4EA64FF0CFBD5FBF6" LSID="urn:lsid:plazi:treatment:4E126219FFB7FFE4EA64FF0CFBD5FBF6" httpUri="http://treatment.plazi.org/id/4E126219FFB7FFE4EA64FF0CFBD5FBF6" lastPageNumber="55" pageId="4" pageNumber="55">
<subSubSection id="8EA18084FFB7FFE4EA64FF0CFBD5FF5F" pageId="4" pageNumber="55" type="nomenclature">
<paragraph id="C604D30FFFB7FFE4EA64FF0CFBD5FF5F" blockId="4.[819,1454,175,231]" pageId="4" pageNumber="55">
<heading id="9D4C6463FFB7FFE4EA64FF0CFBD5FF5F" bold="true" centered="true" fontSize="10" level="1" pageId="4" pageNumber="55" reason="1">
<emphasis id="F4CF0F1DFFB7FFE4EA64FF0CFBD5FF5F" bold="true" pageId="4" pageNumber="55">
4 Key to species of genus
<taxonomicName id="01BBA88CFFB7FFE4ED32FF0CFB2EFF7F" box="[1125,1259,175,199]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="F4CF0F1DFFB7FFE4ED32FF0CFB2EFF7F" bold="true" box="[1125,1259,175,199]" italics="true" pageId="4" pageNumber="55">Cretapsyche</emphasis>
</taxonomicName>
from Cretaceous Burmese amber
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8EA18084FFB7FFE4EA64FEA8FC33FE3A" pageId="4" pageNumber="55" type="description">
<paragraph id="C604D30FFFB7FFE4EA64FEA8FC33FE3A" blockId="4.[819,1462,267,386]" pageId="4" pageNumber="55">
(Family and genus characters: five-segmented maxillary palps each with second segment longest; in forewing forks
<collectionCode id="A0AA4BCAFFB7FFE4EA64FEE8FC82FEDA" box="[819,839,331,354]" pageId="4" pageNumber="55">II</collectionCode>
,
<collectionCode id="A0AA4BCAFFB7FFE4EA02FEE8FCB7FEDA" box="[853,882,331,354]" pageId="4" pageNumber="55">III</collectionCode>
, and
<collectionCode id="A0AA4BCAFFB7FFE4EAE7FEE8FC06FEDA" box="[944,963,331,354]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="55" type="Museum">V</collectionCode>
and in hind wing forks I,
<collectionCode id="A0AA4BCAFFB7FFE4EDBCFEE8FACDFEDA" box="[1259,1288,331,354]" pageId="4" pageNumber="55">III</collectionCode>
, and
<collectionCode id="A0AA4BCAFFB7FFE4EC12FEE8FA9DFEDA" box="[1349,1368,331,354]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="55" type="Museum">V</collectionCode>
present; tibial spurs 2/2/4).
</paragraph>
</subSubSection>
<subSubSection id="8EA18084FFB7FFE4EA10FE05FBD5FBF6" pageId="4" pageNumber="55" type="key">
<paragraph id="C604D30FFFB7FFE4EA10FE05FAF1FE45" blockId="4.[839,1462,422,509]" pageId="4" pageNumber="55">
1. In forewing, Rs and M branching on about the same level, fork
<collectionCode id="A0AA4BCAFFB7FFE4EA8DFE65FC28FE65" box="[986,1005,454,477]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="55" type="Museum">V</collectionCode>
petiolate; in male genitalia coxopodite and harpago at a ratio of about 1:
<quantity id="01437EEAFFB7FFE4EDF0FE45FB15FE45" box="[1191,1232,486,509]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="55" unit="in" value="1.0">1 in</quantity>
length:
<emphasis id="F4CF0F1DFFB7FFE4EC70FE46FAF1FE45" bold="true" box="[1319,1332,485,509]" pageId="4" pageNumber="55">2</emphasis>
</paragraph>
<paragraph id="C604D30FFFB7FFE4EA2BFDBEFB29FDCC" blockId="4.[892,1462,541,628]" pageId="4" pageNumber="55">
– In forewing, M branching before Rs, fork
<collectionCode id="A0AA4BCAFFB7FFE4EC0EFDBEFAA9FD8C" box="[1369,1388,541,564]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="55" type="Museum">V</collectionCode>
nearly sessile; in male genitalia coxopodite and harpago at a ratio of about 2:
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length:
<emphasis id="F4CF0F1DFFB7FFE4ED88FDFFFB29FDCC" bold="true" box="[1247,1260,604,628]" pageId="4" pageNumber="55">3</emphasis>
</paragraph>
<paragraph id="C604D30FFFB7FFE4EA10FD30FB71FD52" blockId="4.[839,1462,659,746]" pageId="4" pageNumber="55">
2. Labial palps three-segmented, terminal segment oval flattened; maxillary palps with pilifer arising from third segment:
<emphasis id="F4CF0F1DFFB7FFE4EA98FD71FB71FD52" bold="true" box="[975,1204,722,746]" pageId="4" pageNumber="55">
<taxonomicName id="01BBA88CFFB7FFE4EA98FD71FB9CFD52" authority="Wichard &amp; Neumann, 2019" authorityName="Wichard &amp; Neumann" authorityYear="2019" box="[975,1113,722,746]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="palpinova" status="sp. nov.">
<emphasis id="F4CF0F1DFFB7FFE4EA98FD71FB9CFD52" bold="true" box="[975,1113,722,746]" italics="true" pageId="4" pageNumber="55">C. palpinova</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EFFCB266FFB7FFE4ED37FD71FB71FD52" box="[1120,1204,722,746]" pageId="4" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="C604D30FFFB7FFE4EA2BFCA9FBCFFCD9" blockId="4.[892,1462,778,865]" pageId="4" pageNumber="55">
– Labial palps three-segmented, simple; maxillary palps without pilifer on third segment:
<taxonomicName id="01BBA88CFFB7FFE4EAC0FCEAFBC6FCD9" box="[919,1027,841,865]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="circula">
<emphasis id="F4CF0F1DFFB7FFE4EAC0FCEAFBC6FCD9" bold="true" box="[919,1027,841,865]" italics="true" pageId="4" pageNumber="55">C. circula</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C604D30FFFB7FFE4EA10FC23FBFCFC6F" blockId="4.[839,1462,896,983]" pageId="4" pageNumber="55">
3. Forewing with median cell open; in male genitalia dorsomedian process of tergum X distinct, ventrally protruding:
<taxonomicName id="01BBA88CFFB7FFE4EA94FC1CFBF7FC6F" box="[963,1074,959,983]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="insueta">
<emphasis id="F4CF0F1DFFB7FFE4EA94FC1CFBF7FC6F" bold="true" box="[963,1074,959,983]" italics="true" pageId="4" pageNumber="55">C. insueta</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C604D30FFFB7FFE4EA2BFC54FBD5FBF6" blockId="4.[892,1461,1015,1102]" pageId="4" pageNumber="55">
– Forewing with median cell closed; in male genitalia dorsomedian process of tergum X simple:
<taxonomicName id="01BBA88CFFB7FFE4EAC0FB95FBCCFBF6" box="[919,1033,1078,1102]" class="Insecta" family="Dysoneuridae" genus="Cretapsyche" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="F4CF0F1DFFB7FFE4EAC0FB95FBCCFBF6" bold="true" box="[919,1033,1078,1102]" italics="true" pageId="4" pageNumber="55">C. elegans</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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