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<document id="50FE6634C8E3A5EC2DA729D83B429187" ID-DOI="10.11646/zootaxa.5594.2.11" ID-ISSN="1175-5326" ID-Zenodo-Dep="14954282" ID-ZooBank="65B73AB6-C13D-4BD7-BCCF-2190FE65D479" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1740853219037" checkinUser="plazi" docAuthor="Feng, Xueli, Xiao, Jie &amp; Zhao, Zhengxue" docDate="2025" docId="A8116825CD36E92CFF1D49B8E2DFA973" docLanguage="en" docName="zootaxa.5594.2.11.pdf" docOrigin="Zootaxa 5594 (2)" docSource="https://doi.org/10.11646/zootaxa.5594.2.11" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tachycines (Gymnaeta) Lianhuaensis 1902, sp. nov." docType="treatment" docVersion="3" lastPageNumber="398" masterDocId="5428105DCD37E92FFF8A4A1CE653A86E" masterDocTitle="Two new species of the subgenus Tachycines (Gymnaeta) (Orthoptera: Rhaphidophoridae: Aemodogryllinae) from a karst cave in China" masterLastPageNumber="400" masterPageNumber="395" pageNumber="396" updateTime="1741272033012" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="8FCB2DD20B55B4E27F54537F69A1CA45">Two new species of the subgenus Tachycines (Gymnaeta) (Orthoptera: Rhaphidophoridae: Aemodogryllinae) from a karst cave in China</mods:title>
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<mods:namePart id="E1282A3EBC7AB7D37F515D87D5487AD1">Feng, Xueli</mods:namePart>
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<mods:affiliation id="5176F6B7748C577EC13912BF9A2C37F9">College of Agriculture, Anshun University, Anshun, China.</mods:affiliation>
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<mods:namePart id="DF4BEDEBAF9F3E106267EF181B6B4C10">Xiao, Jie</mods:namePart>
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<mods:affiliation id="73A25DDFF682C19F3CB96A6681370B4B">College of Agriculture, Anshun University, Anshun, China.</mods:affiliation>
<mods:nameIdentifier id="0E6D4CE8C5210FBE9BA618CE0DD0EF1A" type="email">jiexiao0852@163.com</mods:nameIdentifier>
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<mods:namePart id="AA4C2FD78AAE402BC7F43013673445EE">Zhao, Zhengxue</mods:namePart>
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<mods:affiliation id="8A0A887F780E96DF6B9E68BD798F5384">College of Agriculture, Anshun University, Anshun, China.</mods:affiliation>
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<paragraph id="2007D933CD36E92EFF1D49B8E496ABD1" blockId="1.[151,709,932,995]" box="[151,709,932,959]" pageId="1" pageNumber="396">
<heading id="7B4F6E5FCD36E92EFF1D49B8E496ABD1" bold="true" box="[151,709,932,959]" fontSize="11" level="1" pageId="1" pageNumber="396" reason="1">
<emphasis id="12CC0521CD36E92EFF1D49B8E496ABD1" bold="true" box="[151,709,932,959]" pageId="1" pageNumber="396">
<taxonomicName id="E7B8A2B0CD36E92EFF1D49B8E433ABD1" ID-CoL="8PWB9" authority="(Gymnaeta) Lianhuaensis" authorityName="Lianhuaensis" authorityYear="1902" baseAuthorityName="Gymnaeta" box="[151,608,932,959]" class="Insecta" family="Rhaphidophoridae" genus="Tachycines" higherTaxonomySource="CoL" kingdom="Animalia" order="Orthoptera" pageId="1" pageNumber="396" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="12CC0521CD36E92EFF1D49B8E74EABD1" bold="true" box="[151,285,932,959]" italics="true" pageId="1" pageNumber="396">Tachycines</emphasis>
(
<emphasis id="12CC0521CD36E92EFEA449B8E7F9ABD0" bold="true" box="[302,426,932,958]" italics="true" pageId="1" pageNumber="396">Gymnaeta</emphasis>
)
<emphasis id="12CC0521CD36E92EFE3149B8E433ABD1" bold="true" box="[443,608,932,959]" italics="true" pageId="1" pageNumber="396">Lianhuaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="09FFB85ACD36E92EFDED49B9E496ABD1" box="[615,709,933,959]" pageId="1" pageNumber="396" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="68A28AB8CD36E92EFF1D49D4E79FAF2C" pageId="1" pageNumber="396" type="description">
<paragraph id="2007D933CD36E92EFF1D49D4E740AB8D" blockId="1.[151,709,932,995]" box="[151,275,968,995]" pageId="1" pageNumber="396">
(
<figureCitation id="B883C5B6CD36E92EFF1549D4E6BEAB8D" box="[159,237,968,995]" captionStart="FIGURE 1" captionStartId="2.[151,250,1143,1167]" captionTargetBox="[214,1373,179,1109]" captionTargetId="figure-33@2.[214,1373,179,1117]" captionTargetPageId="2" captionText="FIGURE 1. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (male). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. vertex of head, showing conical tubercles; E. pronotum, dorsal view; F. hind tarsus, lateral view; G. genitalia, ventral view; H. genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14954284" httpUri="https://zenodo.org/record/14954284/files/figure.png" pageId="1" pageNumber="396">Figs. 1</figureCitation>
,
<figureCitation id="B883C5B6CD36E92EFF7149D5E759AB8D" box="[251,266,969,995]" captionStart="FIGURE 2" captionStartId="2.[151,250,1959,1983]" captionTargetBox="[214,1374,1263,1935]" captionTargetId="figure-89@2.[214,1374,1263,1935]" captionTargetPageId="2" captionText="FIGURE 2. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (female). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. ovipositor, lateral view; E. subgenital plate, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14954286" httpUri="https://zenodo.org/record/14954286/files/figure.png" pageId="1" pageNumber="396">2</figureCitation>
)
</paragraph>
<paragraph id="2007D933CD36E92EFF1D4E0CE436ACF8" blockId="1.[151,1437,1040,2038]" pageId="1" pageNumber="396">
<emphasis id="12CC0521CD36E92EFF1D4E0CE77AAC44" bold="true" box="[151,297,1040,1066]" pageId="1" pageNumber="396">Description.</emphasis>
Male. Body small size (
<figureCitation id="B883C5B6CD36E92EFDC94E0CE498AC45" box="[579,715,1040,1067]" captionStart="FIGURE 1" captionStartId="2.[151,250,1143,1167]" captionTargetBox="[214,1373,179,1109]" captionTargetId="figure-33@2.[214,1373,179,1117]" captionTargetPageId="2" captionText="FIGURE 1. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (male). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. vertex of head, showing conical tubercles; E. pronotum, dorsal view; F. hind tarsus, lateral view; G. genitalia, ventral view; H. genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14954284" httpUri="https://zenodo.org/record/14954284/files/figure.png" pageId="1" pageNumber="396">Figs. 1A–C</figureCitation>
). Vertex of head not divided into two conical tubercles, ocellar disappearance (
<figureCitation id="B883C5B6CD36E92EFECC4E28E7F2AC21" box="[326,417,1076,1103]" captionStart="FIGURE 1" captionStartId="2.[151,250,1143,1167]" captionTargetBox="[214,1373,179,1109]" captionTargetId="figure-33@2.[214,1373,179,1117]" captionTargetPageId="2" captionText="FIGURE 1. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (male). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. vertex of head, showing conical tubercles; E. pronotum, dorsal view; F. hind tarsus, lateral view; G. genitalia, ventral view; H. genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14954284" httpUri="https://zenodo.org/record/14954284/files/figure.png" pageId="1" pageNumber="396">Fig. 1D</figureCitation>
). Eyes slightly reduced (
<figureCitation id="B883C5B6CD36E92EFD364E28E545AC21" box="[700,790,1076,1103]" captionStart="FIGURE 1" captionStartId="2.[151,250,1143,1167]" captionTargetBox="[214,1373,179,1109]" captionTargetId="figure-33@2.[214,1373,179,1117]" captionTargetPageId="2" captionText="FIGURE 1. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (male). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. vertex of head, showing conical tubercles; E. pronotum, dorsal view; F. hind tarsus, lateral view; G. genitalia, ventral view; H. genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14954284" httpUri="https://zenodo.org/record/14954284/files/figure.png" pageId="1" pageNumber="396">Fig. 1D</figureCitation>
). Anterior margin of pronotum straight, posterior margin protruding caudad, ventral margin arcshaped (
<figureCitation id="B883C5B6CD36E92EFD1F4E44E4BBAC1D" box="[661,744,1112,1139]" captionStart="FIGURE 1" captionStartId="2.[151,250,1143,1167]" captionTargetBox="[214,1373,179,1109]" captionTargetId="figure-33@2.[214,1373,179,1117]" captionTargetPageId="2" captionText="FIGURE 1. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (male). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. vertex of head, showing conical tubercles; E. pronotum, dorsal view; F. hind tarsus, lateral view; G. genitalia, ventral view; H. genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14954284" httpUri="https://zenodo.org/record/14954284/files/figure.png" pageId="1" pageNumber="396">Fig. 1E</figureCitation>
); posterior margin of mesonotum protruding caudad; posterior margin of metanotum protruding caudad.
</paragraph>
<paragraph id="2007D933CD36E92EFF4D4EBDE24CAD90" blockId="1.[151,1437,1040,2038]" pageId="1" pageNumber="396">
Legs long and slender. Coxae with small medial projections. Fore femur about 2.6 times longer than the pronotum, ventrally unarmed, internal genicular lobe with 1 small spine, external genicular lobe with 1 long spine; fore tibia ventrally with 1 inner spine and 2 outer spines, apex with 1 small outer spine on dorsal surface and 1 pair of long spines on ventral surface, between the paired ventral spines with 1 small spine. Middle femur ventrally unarmed, internal and external genicular lobes with 1 long spine respectively; middle tibiae ventrally with 1 inner spine and 1 outer spine, apex with 1 pair of dorsal spines and 1 pair of ventral spines, between the paired ventral spines with 1 small spine. Hind femur ventrally unarmed; hind tibia dorsally with 23–25 inner spines and 22–23 outer spines, arrange in groups, ventral and dorsal surface with a pair of small subapical spines respectively, apex with 1 pair of long dorsal spines and 1 pair of long ventral spines; the longest dorso-apical spine not exceeding the dorso-apical spine of hind metatarsus (
<figureCitation id="B883C5B6CD36E92EFDC34FF8E4CEAD91" box="[585,669,1508,1535]" captionStart="FIGURE 1" captionStartId="2.[151,250,1143,1167]" captionTargetBox="[214,1373,179,1109]" captionTargetId="figure-33@2.[214,1373,179,1117]" captionTargetPageId="2" captionText="FIGURE 1. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (male). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. vertex of head, showing conical tubercles; E. pronotum, dorsal view; F. hind tarsus, lateral view; G. genitalia, ventral view; H. genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14954284" httpUri="https://zenodo.org/record/14954284/files/figure.png" pageId="1" pageNumber="396">Fig. 1F</figureCitation>
). Hind metatarsus keeled beneath.
</paragraph>
<paragraph id="2007D933CD36E92EFF4D4C14E521AE05" blockId="1.[151,1437,1040,2038]" pageId="1" pageNumber="396">
Abdominal sternites with short ventral projections. Epiphallus of male genitalia like the shape of “H”; median lobe of genitalia with 1 pair of apical lobules, divided by a shallow notch; paired lateral sclerites small; dorsal and ventral pairs of lateral lobes almost equal in length (
<figureCitation id="B883C5B6CD36E92EFD544C4CE535AE05" box="[734,870,1616,1643]" captionStart="FIGURE 1" captionStartId="2.[151,250,1143,1167]" captionTargetBox="[214,1373,179,1109]" captionTargetId="figure-33@2.[214,1373,179,1117]" captionTargetPageId="2" captionText="FIGURE 1. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (male). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. vertex of head, showing conical tubercles; E. pronotum, dorsal view; F. hind tarsus, lateral view; G. genitalia, ventral view; H. genitalia, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.14954284" httpUri="https://zenodo.org/record/14954284/files/figure.png" pageId="1" pageNumber="396">Figs. 1G, H</figureCitation>
).
</paragraph>
<paragraph id="2007D933CD36E92EFF4D4C68E539AE95" blockId="1.[151,1437,1040,2038]" pageId="1" pageNumber="396">
Female. General appearance similar to that of male (
<figureCitation id="B883C5B6CD36E92EFC864C68E5C1AEE1" box="[780,914,1652,1679]" captionStart="FIGURE 2" captionStartId="2.[151,250,1959,1983]" captionTargetBox="[214,1374,1263,1935]" captionTargetId="figure-89@2.[214,1374,1263,1935]" captionTargetPageId="2" captionText="FIGURE 2. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (female). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. ovipositor, lateral view; E. subgenital plate, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14954286" httpUri="https://zenodo.org/record/14954286/files/figure.png" pageId="1" pageNumber="396">Figs. 2A–C</figureCitation>
). Ovipositor longer than half the length of hind femur; base of ovipositor broad, narrowing to apex; apex pointed and slightly curved upwards; almost indistinct denticles on ventral edge of distal part of inferior valves (
<figureCitation id="B883C5B6CD36E92EFC944CA0E52AAEB9" box="[798,889,1724,1751]" captionStart="FIGURE 2" captionStartId="2.[151,250,1959,1983]" captionTargetBox="[214,1374,1263,1935]" captionTargetId="figure-89@2.[214,1374,1263,1935]" captionTargetPageId="2" captionText="FIGURE 2. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (female). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. ovipositor, lateral view; E. subgenital plate, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14954286" httpUri="https://zenodo.org/record/14954286/files/figure.png" pageId="1" pageNumber="396">Fig. 2D</figureCitation>
). Subgenital plate with 3 nearly triangular lobes, median lobe rather large and paired lateral lobes small (
<figureCitation id="B883C5B6CD36E92EFC8C4CFCE50EAE95" box="[774,861,1760,1787]" captionStart="FIGURE 2" captionStartId="2.[151,250,1959,1983]" captionTargetBox="[214,1374,1263,1935]" captionTargetId="figure-89@2.[214,1374,1263,1935]" captionTargetPageId="2" captionText="FIGURE 2. Tachycines (Gymnaeta) Lianhuaensis sp. nov. (female). A. habitus, lateral view; B. habitus, dorsal view; C. habitus, ventral view; D. ovipositor, lateral view; E. subgenital plate, ventral view." figureDoi="http://doi.org/10.5281/zenodo.14954286" httpUri="https://zenodo.org/record/14954286/files/figure.png" pageId="1" pageNumber="396">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="2007D933CD36E92EFF4D4D18E79FAF2C" blockId="1.[151,1437,1040,2038]" pageId="1" pageNumber="396">
<emphasis id="12CC0521CD36E92EFF4D4D18E71CAF70" bold="true" box="[199,335,1796,1822]" pageId="1" pageNumber="396">Coloration.</emphasis>
Body yellowish brown, mottled with dark brown blotches. Face without distinct stripes. Legs with light brown annular stripes.
</paragraph>
</subSubSection>
<subSubSection id="68A28AB8CD36E92EFF4D4D50E2CDAFE4" pageId="1" pageNumber="396" type="materials_examined">
<paragraph id="2007D933CD36E92EFF4D4D50E2CDAFE4" blockId="1.[151,1437,1040,2038]" pageId="1" pageNumber="396">
<emphasis id="12CC0521CD36E92EFF4D4D50E7E3AF08" bold="true" box="[199,432,1868,1894]" pageId="1" pageNumber="396">Material examined.</emphasis>
<materialsCitation id="90D0D36ECD36E92EFE3F4D50E4C0AFE5" collectingDate="2024-09-07" collectorName="Jie Xiao &amp; Zhengxue Zhao &amp; Xueli Feng." county="Puding County" location="Anshun City" municipality="Lianhua Cave" pageId="1" pageNumber="396" specimenCount="1" specimenCount-male="1" stateProvince="Guizhou" typeStatus="holotype">
<typeStatus id="FF036791CD36E92EFE3F4D50E471AF08" box="[437,546,1868,1894]" pageId="1" pageNumber="396" type="holotype">Holotype</typeStatus>
:
<specimenCount id="36BE12BACD36E92EFDA14D51E403AF09" box="[555,592,1869,1895]" count="1" pageId="1" pageNumber="396" type="male">1♂</specimenCount>
,
<collectingMunicipality id="C0634349CD36E92EFDD14D50E4AAAF09" box="[603,761,1868,1895]" pageId="1" pageNumber="396">Lianhua Cave</collectingMunicipality>
,
<collectingCounty id="C966A1BFCD36E92EFC894D50E5FEAF09" box="[771,941,1868,1895]" pageId="1" pageNumber="396">Puding County</collectingCounty>
,
<location id="25678FE8CD36E92EFC3F4D50E210AF08" LSID="urn:lsid:plazi:treatment:A8116825CD36E92CFF1D49B8E2DFA973:25678FE8CD36E92EFC3F4D50E210AF08" box="[949,1091,1868,1894]" county="Puding County" municipality="Lianhua Cave" name="Anshun City" pageId="1" pageNumber="396" stateProvince="Guizhou">Anshun City</location>
,
<collectingRegion id="E27C17D1CD36E92EFBC74D50E34AAF08" box="[1101,1305,1868,1894]" country="China" name="Guizhou" pageId="1" pageNumber="396">Guizhou Province</collectingRegion>
,
<date id="5406FFF3CD36E92EFAAE4D51E3C4AF09" box="[1316,1431,1869,1895]" pageId="1" pageNumber="396" value="2024-09-07">
<collectingDate id="4442061BCD36E92EFAAE4D51E3C4AF09" box="[1316,1431,1869,1895]" pageId="1" pageNumber="396" value="2024-09-07">7.IX.2024</collectingDate>
</date>
, coll.
<collectorName id="8D4DBCE5CD36E92EFF454D6CE77DAFE4" box="[207,302,1904,1930]" pageId="1" pageNumber="396">Jie Xiao</collectorName>
,
<collectorName id="8D4DBCE5CD36E92EFEB04D6CE7B8AFE4" box="[314,491,1904,1930]" pageId="1" pageNumber="396">Zhengxue Zhao</collectorName>
&amp;
<collectorName id="8D4DBCE5CD36E92EFD844D6CE4C0AFE5" box="[526,659,1904,1931]" pageId="1" pageNumber="396">Xueli Feng.</collectorName>
</materialsCitation>
<materialsCitation id="90D0D36ECD36E92EFD134D6DE2C8AFE4" box="[665,1179,1904,1931]" collectingDate="2024-09-07" collectorName="Jie Xiao &amp; Zhengxue Zhao &amp; Xueli Feng." county="Puding County" location="Anshun City" municipality="Lianhua Cave" pageId="1" pageNumber="396" specimenCount="10" specimenCount-female="5" specimenCount-male="5" stateProvince="Guizhou" typeStatus="paratype">
<typeStatus id="FF036791CD36E92EFD134D6DE559AFE5" box="[665,778,1905,1931]" pageId="1" pageNumber="396" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="36BE12BACD36E92EFC9F4D6CE56AAFE5" box="[789,825,1904,1931]" count="5" pageId="1" pageNumber="396" type="male">5♂</specimenCount>
,
<specimenCount id="36BE12BACD36E92EFCCC4D6CE539AFE5" box="[838,874,1904,1931]" count="5" pageId="1" pageNumber="396" type="female">5♀</specimenCount>
, same data as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="68A28AB8CD36E92EFF4D4D89E283AFBD" pageId="1" pageNumber="396" type="description">
<paragraph id="2007D933CD36E92EFF4D4D89E283AFBD" blockId="1.[151,1437,1040,2038]" pageId="1" pageNumber="396">
<emphasis id="12CC0521CD36E92EFF4D4D89E794AFC0" bold="true" box="[199,455,1940,1967]" pageId="1" pageNumber="396">Measurements (mm).</emphasis>
Body:
<specimenCount id="36BE12BACD36E92EFD944D88E462AFC1" box="[542,561,1940,1967]" count="1" pageId="1" pageNumber="396" type="male">♂</specimenCount>
10.40–15.32,
<specimenCount id="36BE12BACD36E92EFD454D88E48EAFC1" box="[719,733,1940,1967]" count="1" pageId="1" pageNumber="396" type="female">♀</specimenCount>
9.58–12.36; pronotum:
<specimenCount id="36BE12BACD36E92EFC7B4D89E257AFC1" box="[1009,1028,1941,1967]" count="1" pageId="1" pageNumber="396" type="male">♂</specimenCount>
3.84–4.02,
<specimenCount id="36BE12BACD36E92EFB0C4D89E2C7AFC1" box="[1158,1172,1941,1967]" count="1" pageId="1" pageNumber="396" type="female">♀</specimenCount>
3.74–4.34; fore femur:
<specimenCount id="36BE12BACD36E92EFF1D4DA4E6F9AFBC" box="[151,170,1976,2002]" count="1" pageId="1" pageNumber="396" type="male">♂</specimenCount>
10.10–10.64,
<specimenCount id="36BE12BACD36E92EFECC4DA4E707AFBC" box="[326,340,1976,2002]" count="1" pageId="1" pageNumber="396" type="female">♀</specimenCount>
9.32–9.68; hind femur:
<specimenCount id="36BE12BACD36E92EFDEE4DA4E424AFBC" box="[612,631,1976,2002]" count="1" pageId="1" pageNumber="396" type="male">♂</specimenCount>
16.84–17.06,
<specimenCount id="36BE12BACD36E92EFC994DA4E572AFBD" box="[787,801,1976,2003]" count="1" pageId="1" pageNumber="396" type="female">♀</specimenCount>
16.58–17.32; ovipositor:
<specimenCount id="36BE12BACD36E92EFBCE4DA5E201AFBD" box="[1092,1106,1977,2003]" count="1" pageId="1" pageNumber="396" type="female">♀</specimenCount>
9.02–9.10.
</paragraph>
</subSubSection>
<subSubSection id="68A28AB8CD36E92EFF4D4DC0E47FAF98" box="[199,556,2012,2038]" pageId="1" pageNumber="396" type="distribution">
<paragraph id="2007D933CD36E92EFF4D4DC0E47FAF98" blockId="1.[151,1437,1040,2038]" box="[199,556,2012,2038]" pageId="1" pageNumber="396">
<emphasis id="12CC0521CD36E92EFF4D4DC0E732AF98" bold="true" box="[199,353,2012,2038]" pageId="1" pageNumber="396">Distribution.</emphasis>
<collectingCountry id="58AF99A3CD36E92EFEED4DC0E7F9AF98" box="[359,426,2012,2038]" name="China" pageId="1" pageNumber="396">China</collectingCountry>
(
<collectingRegion id="E27C17D1CD36E92EFE334DC0E473AF98" box="[441,544,2012,2038]" country="China" name="Guizhou" pageId="1" pageNumber="396">Guizhou</collectingRegion>
).
</paragraph>
</subSubSection>
<caption id="74C789BBCD35E92DFF1D4E6BE471ACB9" ID-DOI="http://doi.org/10.5281/zenodo.14954284" ID-Zenodo-Dep="14954284" httpUri="https://zenodo.org/record/14954284/files/figure.png" pageId="2" pageNumber="397" startId="2.[151,250,1143,1167]" targetBox="[214,1373,179,1109]" targetPageId="2" targetType="figure">
<paragraph id="2007D933CD35E92DFF1D4E6BE471ACB9" blockId="2.[151,1437,1143,1239]" pageId="2" pageNumber="397">
<emphasis id="12CC0521CD35E92DFF1D4E6BE740ACE1" bold="true" box="[151,275,1143,1167]" pageId="2" pageNumber="397">FIGURE 1.</emphasis>
<taxonomicName id="E7B8A2B0CD35E92DFE934E6BE454ACE1" authorityName="Adelung" authorityYear="1902" box="[281,519,1143,1167]" class="Insecta" family="Rhaphidophoridae" genus="Tachycines" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="subGenus" subGenus="Gymnaeta">
<emphasis id="12CC0521CD35E92DFE934E6BE7DAACE1" box="[281,393,1143,1167]" italics="true" pageId="2" pageNumber="397">Tachycines</emphasis>
(
<emphasis id="12CC0521CD35E92DFE1D4E6BE7ACACE0" box="[407,511,1143,1166]" italics="true" pageId="2" pageNumber="397">Gymnaeta</emphasis>
)
</taxonomicName>
<taxonomicName id="E7B8A2B0CD35E92DFD844E6BE4C6ACE1" authorityName="Feng &amp; Xiao &amp; Zhao" authorityYear="2025" box="[526,661,1143,1167]" class="Insecta" family="Carabidae" genus="Lianhuaensis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="12CC0521CD35E92DFD844E6BE4C6ACE1" box="[526,661,1143,1167]" italics="true" pageId="2" pageNumber="397">Lianhuaensis</emphasis>
</taxonomicName>
<emphasis id="12CC0521CD35E92DFD114E6BE4B8ACE1" bold="true" box="[667,747,1143,1167]" pageId="2" pageNumber="397">
<taxonomicNameLabel id="09FFB85ACD35E92DFD114E6BE4B8ACE1" box="[667,747,1143,1167]" pageId="2" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(male).
<emphasis id="12CC0521CD35E92DFCCA4E6BE501ACE1" bold="true" box="[832,850,1143,1167]" pageId="2" pageNumber="397">A</emphasis>
. habitus, lateral view;
<emphasis id="12CC0521CD35E92DFBB34E6BE219ACE1" bold="true" box="[1081,1098,1143,1167]" pageId="2" pageNumber="397">B</emphasis>
. habitus, dorsal view;
<emphasis id="12CC0521CD35E92DFAA44E6BE313ACE1" bold="true" box="[1326,1344,1143,1167]" pageId="2" pageNumber="397">C</emphasis>
. habitus, ventral view;
<emphasis id="12CC0521CD35E92DFEA94E87E766ACDD" bold="true" box="[291,309,1179,1203]" pageId="2" pageNumber="397">D</emphasis>
. vertex of head, showing conical tubercles;
<emphasis id="12CC0521CD35E92DFD7E4E87E556ACDD" bold="true" box="[756,773,1179,1203]" pageId="2" pageNumber="397">E</emphasis>
. pronotum, dorsal view;
<emphasis id="12CC0521CD35E92DFB8E4E87E244ACDD" bold="true" box="[1028,1047,1179,1203]" pageId="2" pageNumber="397">F.</emphasis>
hind tarsus, lateral view;
<emphasis id="12CC0521CD35E92DFA944E87E361ACDD" bold="true" box="[1310,1330,1179,1203]" pageId="2" pageNumber="397">G</emphasis>
. genitalia, ventral view;
<emphasis id="12CC0521CD35E92DFEA84EA3E765ACB9" bold="true" box="[290,310,1215,1239]" pageId="2" pageNumber="397">H</emphasis>
. genitalia, dorsal view.
</paragraph>
</caption>
<caption id="74C789BBCD35E92DFF1D4DBBE5D5AF8D" ID-DOI="http://doi.org/10.5281/zenodo.14954286" ID-Zenodo-Dep="14954286" httpUri="https://zenodo.org/record/14954286/files/figure.png" pageId="2" pageNumber="397" startId="2.[151,250,1959,1983]" targetBox="[214,1374,1263,1935]" targetPageId="2" targetType="figure">
<paragraph id="2007D933CD35E92DFF1D4DBBE5D5AF8D" blockId="2.[151,1436,1959,2019]" pageId="2" pageNumber="397">
<emphasis id="12CC0521CD35E92DFF1D4DBBE741AFD1" bold="true" box="[151,274,1959,1983]" pageId="2" pageNumber="397">FIGURE 2.</emphasis>
<taxonomicName id="E7B8A2B0CD35E92DFE9D4DBBE457AFD1" authorityName="Adelung" authorityYear="1902" box="[279,516,1959,1983]" class="Insecta" family="Rhaphidophoridae" genus="Tachycines" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="subGenus" subGenus="Gymnaeta">
<emphasis id="12CC0521CD35E92DFE9D4DBBE7D4AFD1" box="[279,391,1959,1983]" italics="true" pageId="2" pageNumber="397">Tachycines</emphasis>
(
<emphasis id="12CC0521CD35E92DFE1E4DBBE7AFAFD0" box="[404,508,1959,1982]" italics="true" pageId="2" pageNumber="397">Gymnaeta</emphasis>
)
</taxonomicName>
<taxonomicName id="E7B8A2B0CD35E92DFD834DBBE4C3AFD1" authorityName="Feng &amp; Xiao &amp; Zhao" authorityYear="2025" box="[521,656,1959,1983]" class="Insecta" family="Carabidae" genus="Lianhuaensis" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="397" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="12CC0521CD35E92DFD834DBBE4C3AFD1" box="[521,656,1959,1983]" italics="true" pageId="2" pageNumber="397">Lianhuaensis</emphasis>
</taxonomicName>
<emphasis id="12CC0521CD35E92DFD1F4DBBE4B7AFD1" bold="true" box="[661,740,1959,1983]" pageId="2" pageNumber="397">
<taxonomicNameLabel id="09FFB85ACD35E92DFD1F4DBBE4B7AFD1" box="[661,740,1959,1983]" pageId="2" pageNumber="397" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(female).
<emphasis id="12CC0521CD35E92DFCC14DBBE537AFD1" bold="true" box="[843,868,1959,1983]" pageId="2" pageNumber="397">A.</emphasis>
habitus, lateral view;
<emphasis id="12CC0521CD35E92DFBB44DBBE205AFD1" bold="true" box="[1086,1110,1959,1983]" pageId="2" pageNumber="397">B.</emphasis>
habitus, dorsal view;
<emphasis id="12CC0521CD35E92DFAA54DBBE31BAFD1" bold="true" box="[1327,1352,1959,1983]" pageId="2" pageNumber="397">C.</emphasis>
habitus, ventral view;
<emphasis id="12CC0521CD35E92DFEA84DD7E768AF8D" bold="true" box="[290,315,1995,2019]" pageId="2" pageNumber="397">D.</emphasis>
ovipositor, lateral view;
<emphasis id="12CC0521CD35E92DFDB24DD7E41CAF8D" bold="true" box="[568,591,1995,2019]" pageId="2" pageNumber="397">E.</emphasis>
subgenital plate, ventral view.
</paragraph>
</caption>
<subSubSection id="68A28AB8CD34E92CFF4D4A8AE4DAA897" pageId="3" pageNumber="398" type="discussion">
<paragraph id="2007D933CD34E92CFF4D4A8AE4DAA897" blockId="3.[151,1436,150,285]" pageId="3" pageNumber="398">
<emphasis id="12CC0521CD34E92CFF4D4A8AE71FA8DE" bold="true" box="[199,332,150,176]" pageId="3" pageNumber="398">Discussion.</emphasis>
The new species resembles most species in the subgenus
<taxonomicName id="E7B8A2B0CD34E92CFC694A8BE2BEA8DF" authorityName="Adelung" authorityYear="1902" box="[995,1261,151,177]" class="Insecta" family="Rhaphidophoridae" genus="Tachycines" kingdom="Animalia" order="Orthoptera" pageId="3" pageNumber="398" phylum="Arthropoda" rank="subGenus" subGenus="Gymnaeta">
<emphasis id="12CC0521CD34E92CFC694A8BE20DA8DF" box="[995,1118,151,177]" italics="true" pageId="3" pageNumber="398">Tachycines</emphasis>
(
<emphasis id="12CC0521CD34E92CFBFA4A8BE2B0A8DF" box="[1136,1251,151,177]" italics="true" pageId="3" pageNumber="398">Gymnaeta</emphasis>
)
</taxonomicName>
in terms of the shape of the male genitalia (approximately“H”), yet it is clearly distinguishable by the vestigial conical tubercles on the vertex of the head and the vestigial eyes.
</paragraph>
</subSubSection>
<subSubSection id="68A28AB8CD34E92CFF4D4B1FE2DFA973" box="[199,1164,259,285]" pageId="3" pageNumber="398" type="etymology">
<paragraph id="2007D933CD34E92CFF4D4B1FE2DFA973" blockId="3.[151,1436,150,285]" box="[199,1164,259,285]" pageId="3" pageNumber="398">
<emphasis id="12CC0521CD34E92CFF4D4B1FE71DA973" bold="true" box="[199,334,259,285]" pageId="3" pageNumber="398">Etymology.</emphasis>
The specific epithet refers to the Lianhua Cave where this species occurs.
</paragraph>
</subSubSection>
</treatment>
</document>