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<document id="83FF502A4BC675AB3AEFE9AF4D8B787D" ID-DOI="10.5852/ejt.2024.958.2665" ID-ISSN="2118-9773" ID-Zenodo-Dep="13837028" ID-ZooBank="425BCF68-144B-4861-BCED-CB570D8246D1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727252768769" checkinUser="plazi" docAuthor="Constant, Jérôme &amp; Pham, Hong Thai" docDate="2024" docId="03B887FAFFA2FFC22377FBF75CC01653" docLanguage="en" docName="EJT.2024.958.114-150.pdf" docOrigin="European Journal of Taxonomy 958" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2665/12349" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zanna limbourgi Constant 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="34" masterDocId="FF81FF82FFBAFFE3213DFFAE5E6A1776" masterDocTitle="Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)" masterLastPageNumber="150" masterPageNumber="114" pageNumber="25" updateTime="1727253238722" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6B640478990980B8BA01AB1C8F0B438E">Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<mods:namePart id="F6AE05473E9047A360ECD0AE3EB1129E">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="7CDB052243FC290F75E89B94E6B4A3A0">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier Street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
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<mods:namePart id="5F28FC0CE607E5B9B4D32DE7D2FE0581">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="5A385E169BB498429790AC5CCB10F8AA">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.</mods:affiliation>
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<mods:date id="A505BA82DA91F20B940AEAC8B34AF199">2024</mods:date>
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<paragraph id="8BAE36ECFFA2FFFB2377FBF75D801305" blockId="24.[358,1229,1112,1210]" box="[586,1002,1112,1139]" pageId="24" pageNumber="25">
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<taxonomicName id="4C114D6FFFA2FFFB2377FBF75DE21305" authority="Constant" authorityName="Constant" authorityYear="2024" box="[586,904,1112,1139]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA2FFFB2377FBF75D721305" bold="true" box="[586,792,1112,1139]" italics="true" pageId="24" pageNumber="25">Zanna limbourgi</emphasis>
Constant
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<taxonomicNameLabel id="A2565785FFA2FFFB22B2FBF75D801305" box="[911,1002,1113,1139]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C30B6567FFA2FFFB2369FB315D8A13CF" box="[596,992,1183,1210]" pageId="24" pageNumber="25" type="description">
<paragraph id="8BAE36ECFFA2FFFB2369FB315D8A13CF" blockId="24.[358,1229,1112,1210]" box="[596,992,1183,1210]" pageId="24" pageNumber="25">
<figureCitation id="132A2A69FFA2FFFB2369FB315CF413CC" box="[596,670,1183,1210]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="24" pageNumber="25">Figs 5</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB2396FB315C9D13CC" box="[683,759,1183,1210]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="25.[189,230,1925,1951]" captionStartId-1="26.[197,236,1958,1984]" captionStartId-2="28.[185,226,1960,1986]" captionTargetBox-0="[240,1354,270,1868]" captionTargetBox-1="[230,1373,285,1917]" captionTargetBox-2="[213,1370,261,1921]" captionTargetId-0="figure-46@25.[230,1354,265,1868]" captionTargetId-1="figure-62@26.[226,1383,268,1920]" captionTargetId-2="figure-36@28.[213,1370,258,1921]" captionTargetPageId-0="25" captionTargetPageId-1="26" captionTargetPageId-2="28" captionText-0="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." captionText-1="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" captionText-2="Fig. 17. Zanna limbourgi Constant sp. nov., holotype, ♂, in nature in Phnom Aural Wildlife Sanctuary, Cambodia, 12 May 2018. A. Habitus, lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi-0="http://doi.org/10.5281/zenodo.13837073" figureDoi-1="http://doi.org/10.5281/zenodo.13837076" figureDoi-2="http://doi.org/10.5281/zenodo.13837078" httpUri-0="https://zenodo.org/record/13837073/files/figure.png" httpUri-1="https://zenodo.org/record/13837076/files/figure.png" httpUri-2="https://zenodo.org/record/13837078/files/figure.png" pageId="24" pageNumber="25">1517</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB223EFB315D1613CC" box="[771,892,1183,1210]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="24" pageNumber="25">18D, H, L</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB22B4FB315D8A13CF" box="[905,992,1183,1209]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="24" pageNumber="25">19GH</figureCitation>
</paragraph>
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<subSubSection id="C30B6567FFA2FFFB2180FB4E5D2F11B9" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="8BAE36ECFFA2FFFB2180FB4E5F5C138C" blockId="24.[189,310,1248,1274]" box="[189,310,1248,1274]" pageId="24" pageNumber="25">
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<emphasis id="B965EAFEFFA2FFFB2180FB4E5F5C138C" bold="true" box="[189,310,1248,1274]" pageId="24" pageNumber="25">Diagnosis</emphasis>
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<paragraph id="8BAE36ECFFA2FFFB2180FAA15D2F11B9" blockId="24.[189,1399,1295,1744]" pageId="24" pageNumber="25">
The species is closest to
<taxonomicName id="4C114D6FFFA2FFFB20CCFABE5CAE125F" authorityName="Constant &amp; Pham" authorityYear="2024" box="[497,708,1295,1321]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFA2FFFB20CCFABE5CAE125F" box="[497,708,1295,1321]" italics="true" pageId="24" pageNumber="25">Zanna bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA2FFFB23F2FAA15D5A125F" box="[719,816,1295,1321]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C114D6FFFA2FFFB227DFABE5A56125F" authority="Constant" authorityName="Constant" authorityYear="2024" box="[832,1084,1295,1321]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFA2FFFB227DFABE5DA3125F" box="[832,969,1295,1321]" italics="true" pageId="24" pageNumber="25">Z. chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA2FFFB257AFAA15AC2125F" box="[1095,1192,1295,1321]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C114D6FFFA2FFFB25D0FABE5B1D125F" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1261,1399,1295,1321]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFA2FFFB25D0FABE5B1D125F" box="[1261,1399,1295,1321]" italics="true" pageId="24" pageNumber="25">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA2FFFB2180FA9D5F76123B" box="[189,284,1331,1357]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can be separated by the following characters: basal portion of head (
<figureCitation id="132A2A69FFA2FFFB243BFA9C5E9A1219" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="24" pageNumber="25">Fig. 16B, EF</figureCitation>
) covered in moderately large, rather well-spaced, black pitting with background of dense, very small black pitting (dense, irregular, rather large, coarse, sometimes coalescent black pitting in
<taxonomicName id="4C114D6FFFA2FFFB2180FA335F3312C0" authorityName="Constant &amp; Pham" authorityYear="2024" box="[189,345,1436,1462]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bidoupana">
<emphasis id="B965EAFEFFA2FFFB2180FA335F3312C0" box="[189,345,1436,1462]" italics="true" pageId="24" pageNumber="25">Z. bidoupana</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA2FFFB2045FA325C7E12C0" box="[376,532,1436,1462]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="24" pageNumber="25">Fig. 3B, EF</figureCitation>
; moderately large, rather well-spaced, black pitting without background of dense, very small black pitting in
<taxonomicName id="4C114D6FFFA2FFFB2342FA6E5D6212AF" authorityName="Constant &amp; Pham" authorityYear="2024" box="[639,776,1471,1497]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFA2FFFB2342FA6E5D6212AF" box="[639,776,1471,1497]" italics="true" pageId="24" pageNumber="25">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA2FFFB2216FA115DB712AF" box="[811,989,1471,1497]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="24" pageNumber="25">Fig. 12B, EF</figureCitation>
); anal tube of male rather high in lateral view (
<figureCitation id="132A2A69FFA2FFFB2069FA4C5FAA128A" box="[340,448,1506,1532]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="24" pageNumber="25">Fig. 16A</figureCitation>
), about 1.65 times as long as high, with ventral angle in basal half (rather flattened dorsoventrally, 1.952.14 times as long as high in lateral view, with ventral angle at midlength in
<taxonomicName id="4C114D6FFFA2FFFB2180F9875F291134" authorityName="Constant" authorityYear="2024" box="[189,323,1576,1602]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="chartieri">
<emphasis id="B965EAFEFFA2FFFB2180F9875F291134" box="[189,323,1576,1602]" italics="true" pageId="24" pageNumber="25">Z. chartieri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFA2FFFB2040F9875C6E1134" authorityName="Constant &amp; Pham" authorityYear="2024" box="[381,516,1576,1602]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFA2FFFB2040F9875C6E1134" box="[381,516,1576,1602]" italics="true" pageId="24" pageNumber="25">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA2FFFB231CF9865CE81135" box="[545,642,1576,1603]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="24" pageNumber="25">Figs 7A</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB23B3F9875CAF1135" box="[654,709,1577,1603]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">13A</figureCitation>
); pygofer of male in lateral view (
<figureCitation id="132A2A69FFA2FFFB256AF9865AA81134" box="[1111,1218,1576,1602]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="24" pageNumber="25">Fig. 16A</figureCitation>
) with posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle (posterior margin projecting into a large posterior lobe angularly rounded in midheight in
<taxonomicName id="4C114D6FFFA2FFFB25E6F9DE5B0B11FF" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1243,1377,1647,1673]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFA2FFFB25E6F9DE5B0B11FF" box="[1243,1377,1647,1673]" italics="true" pageId="24" pageNumber="25">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA2FFFB2180F93C5F4211DA" box="[189,296,1682,1708]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">Fig. 13A</figureCitation>
; posterior margin moderately sinuate, forming a large, broadly rounded posterior lobe in dorsal half in
<taxonomicName id="4C114D6FFFA2FFFB202DF9185FC611B9" authorityName="Constant &amp; Pham" authorityYear="2024" box="[272,428,1717,1743]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bidoupana">
<emphasis id="B965EAFEFFA2FFFB202DF9185FC611B9" box="[272,428,1717,1743]" italics="true" pageId="24" pageNumber="25">Z. bidoupana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFA2FFFB20DBF9185C1211B9" authorityName="Constant" authorityYear="2024" box="[486,632,1717,1743]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFA2FFFB20DBF9185C1211B9" box="[486,632,1717,1743]" italics="true" pageId="24" pageNumber="25">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA2FFFB23A9F91B5C9E11B9" box="[660,756,1717,1743]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="24" pageNumber="25">Figs 3A</figureCitation>
,
<figureCitation id="132A2A69FFA2FFFB223CF91B5D5D11B9" box="[769,823,1717,1743]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="24" pageNumber="25">16A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA2FFF92180F9595B151F70" lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="etymology">
<paragraph id="8BAE36ECFFA2FFFB2180F9595F2E1067" blockId="24.[189,324,1783,1809]" box="[189,324,1783,1809]" pageId="24" pageNumber="25">
<heading id="D0E68180FFA2FFFB2180F9595F2E1067" bold="true" box="[189,324,1783,1809]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B965EAFEFFA2FFFB2180F9595F2E1067" bold="true" box="[189,324,1783,1809]" pageId="24" pageNumber="25">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA2FFFB2180F88B5FCA10F0" blockId="24.[189,1399,1829,1926]" pageId="24" pageNumber="25">
The species epithet is a patronym dedicated to my great (although still a bit fat, see
<bibRefCitation id="EF804B1DFFA2FFFB2582F88B5B041049" author="Constant J." box="[1215,1390,1829,1855]" pageId="24" pageNumber="25" pagination="41 - 54" refId="ref13027" refString="Constant J. 2008. Revision of the Eurybrachidae (XIII). The new Australian genus Chewobrachys (Hemiptera: Fulgoromorpha). Zootaxa 1898: 41 - 54." type="journal article" year="2008">Constant 2008</bibRefCitation>
) colleague Pol Limbourg (RBINS) who spotted the
<typeStatus id="54AA884EFFA2FFFB2212F8E75D081015" box="[815,866,1865,1891]" pageId="24" pageNumber="25">type</typeStatus>
specimens on a tree trunk in Phnom Aural Wildlife Sanctuary.
</paragraph>
<caption id="DF6E6664FFA3FFFA2180F82B5FB31090" ID-DOI="http://doi.org/10.5281/zenodo.13837073" ID-Zenodo-Dep="13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="25" pageNumber="26" startId="25.[189,230,1925,1951]" targetBox="[240,1354,270,1868]" targetPageId="25">
<paragraph id="8BAE36ECFFA3FFFA2180F82B5FB31090" blockId="25.[189,1399,1925,2022]" pageId="25" pageNumber="26">
<emphasis id="B965EAFEFFA3FFFA2180F82B5F7210E9" bold="true" box="[189,280,1925,1951]" pageId="25" pageNumber="26">Fig. 15.</emphasis>
<taxonomicName id="4C114D6FFFA3FFFA201DF8285C3910E9" authority="Constant" authorityName="Constant" authorityYear="2024" box="[288,595,1925,1951]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA3FFFA201DF8285F8E10E9" box="[288,484,1925,1951]" italics="true" pageId="25" pageNumber="26">Zanna limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA3FFFA2367F8285CDD10D6" box="[602,695,1926,1952]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS).
<emphasis id="B965EAFEFFA3FFFA22F1F82B5D8B10E9" bold="true" box="[972,993,1925,1951]" pageId="25" pageNumber="26">A</emphasis>
. Habitus dorsal view.
<emphasis id="B965EAFEFFA3FFFA25D8F82B5A9310E9" bold="true" box="[1253,1273,1925,1951]" pageId="25" pageNumber="26">B</emphasis>
. Head and thorax, dorsal view.
<emphasis id="B965EAFEFFA3FFFA2093F8065FA910B4" bold="true" box="[430,451,1960,1986]" pageId="25" pageNumber="26">C</emphasis>
. Habitus, ventral view.
<emphasis id="B965EAFEFFA3FFFA23E3F8075C9910B5" bold="true" box="[734,755,1961,1987]" pageId="25" pageNumber="26">D</emphasis>
. Habitus, lateral view.
<emphasis id="B965EAFEFFA3FFFA253BF8075A7010B5" bold="true" box="[1030,1050,1961,1987]" pageId="25" pageNumber="26">E</emphasis>
. Head, ventral view.
<emphasis id="B965EAFEFFA3FFFA242AF8075B4310B5" bold="true" box="[1303,1321,1961,1987]" pageId="25" pageNumber="26">F</emphasis>
. Head and thorax, lateral view.
</paragraph>
</caption>
<caption id="DF6E6664FFA0FFF921F8F8085B141095" ID-DOI="http://doi.org/10.5281/zenodo.13837076" ID-Zenodo-Dep="13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="26" pageNumber="27" startId="26.[197,236,1958,1984]" targetBox="[230,1373,285,1917]" targetPageId="26">
<paragraph id="8BAE36ECFFA0FFF921F8F8085B141095" blockId="26.[197,1407,1958,2020]" pageId="26" pageNumber="27">
<emphasis id="B965EAFEFFA0FFF921F8F8085F7410B6" bold="true" box="[197,286,1958,1984]" pageId="26" pageNumber="27">Fig. 16.</emphasis>
<taxonomicName id="4C114D6FFFA0FFF92018F8095C2410B6" authority="Constant" authorityName="Constant" authorityYear="2024" box="[293,590,1958,1984]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA0FFF92018F8095F8910B6" box="[293,483,1958,1984]" italics="true" pageId="26" pageNumber="27">Zanna limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA0FFF92368F8095CC510B7" box="[597,687,1959,1985]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS), terminalia.
<emphasis id="B965EAFEFFA0FFF92506F8085A1810B6" bold="true" box="[1083,1138,1958,1984]" pageId="26" pageNumber="27">AD</emphasis>
. Pygofer, anal tube and gonostyli.
<emphasis id="B965EAFEFFA0FFF92001F8675F3B1095" bold="true" box="[316,337,1993,2019]" pageId="26" pageNumber="27">A</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFA0FFF92317F8675C541095" bold="true" box="[554,574,1993,2019]" pageId="26" pageNumber="27">B</emphasis>
. Caudal view.
<emphasis id="B965EAFEFFA0FFF923D4F8675C941095" bold="true" box="[745,766,1993,2019]" pageId="26" pageNumber="27">C</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFA0FFF92298F8675DD01095" bold="true" box="[933,954,1993,2019]" pageId="26" pageNumber="27">D</emphasis>
. Ventral view.
<emphasis id="B965EAFEFFA0FFF9255AF8675AF41095" bold="true" box="[1127,1182,1993,2019]" pageId="26" pageNumber="27">EH</emphasis>
. Aedeagus.
<emphasis id="B965EAFEFFA0FFF92416F8675B541095" bold="true" box="[1323,1342,1993,2019]" pageId="26" pageNumber="27">E</emphasis>
. Left
</paragraph>
</caption>
<paragraph id="8BAE36ECFFA0FFF921FBF8425B151F70" blockId="26.[198,1407,2028,2055]" box="[198,1407,2028,2055]" pageId="26" pageNumber="27">
lateral view.
<emphasis id="B965EAFEFFA0FFF92072F8435F0B1F71" bold="true" box="[335,353,2029,2055]" pageId="26" pageNumber="27">F</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFA0FFF920C1F8425C791F70" bold="true" box="[508,531,2028,2054]" pageId="26" pageNumber="27">G</emphasis>
. Laterodorsal view.
<emphasis id="B965EAFEFFA0FFF923CCF8435D621F71" bold="true" box="[753,776,2029,2055]" pageId="26" pageNumber="27">H</emphasis>
. Ventral view.Abbreviations: see Materials and methods.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA1FFF82180FEA75F7115B9" pageId="27" pageNumber="28" type="materials_examined">
<paragraph id="8BAE36ECFFA1FFF82180FEA75F061654" blockId="27.[189,364,264,291]" box="[189,364,264,291]" pageId="27" pageNumber="28">
<heading id="D0E68180FFA1FFF82180FEA75F061654" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B965EAFEFFA1FFF82180FEA75F061654" bold="true" box="[189,364,264,291]" pageId="27" pageNumber="28">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA1FFF821D7FE995F361627" blockId="27.[189,1399,311,513]" box="[234,348,311,337]" pageId="27" pageNumber="28">
<emphasis id="B965EAFEFFA1FFF821D7FE995F361627" bold="true" box="[234,348,311,337]" pageId="27" pageNumber="28">
<typeStatus id="54AA884EFFA1FFF821D7FE995F361627" box="[234,348,311,337]" pageId="27" pageNumber="28" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FEF45FE21577" blockId="27.[189,1399,311,513]" pageId="27" pageNumber="28">
<materialsCitation id="3B793CB1FFA1FFF82180FEF45FE21577" collectingDate="2018-05-10" collectingDateMax="2018-05-14" collectingDateMin="2018-05-10" collectionCode="RBINS" collectorName="J. Constant &amp; P. Limbourg &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc. &amp; Leg. J. Constant" country="Cambodia" elevation="300" latitude="11.983334" location="Srea Ken Village" longLatPrecision="1294" longitude="104.13333" municipality="Phnom Aural Wildlife Sanctuary" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Speu" typeStatus="holotype">
<collectingCountry id="F306767CFFA1FFF82180FEF45F311602" box="[189,347,346,372]" name="Cambodia" pageId="27" pageNumber="28">CAMBODIA</collectingCountry>
<specimenCount id="9D17FD65FFA1FFF82049FEF45FC81603" box="[372,418,346,373]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D5F80EFFA1FFF82092FEF55CBC1602" box="[431,726,346,373]" country="Cambodia" name="Kampong Speu" pageId="27" pageNumber="28">Kampong Speu Province</collectingRegion>
,
<collectingMunicipality id="6BCAAC96FFA1FFF823DFFEF45A081602" box="[738,1122,346,372]" pageId="27" pageNumber="28">Phnom Aural Wildlife Sanctuary</collectingMunicipality>
, near
<location id="8ECE6037FFA1FFF8259AFEF45B181602" LSID="urn:lsid:plazi:treatment:03B887FAFFA2FFC22377FBF75CC01653:8ECE6037FFA1FFF8259AFEF45B181602" box="[1191,1394,346,373]" country="Cambodia" latitude="11.983334" longLatPrecision="1294" longitude="104.13333" municipality="Phnom Aural Wildlife Sanctuary" name="Srea Ken Village" pageId="27" pageNumber="28" stateProvince="Kampong Speu">Srea Ken Village</location>
;
<geoCoordinate id="EE25502BFFA1FFF82180FED25F4216EE" box="[189,296,380,408]" degrees="11" direction="north" minutes="59" orientation="latitude" pageId="27" pageNumber="28" precision="925" value="11.983334">11°59ʹ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFA1FFF82008FED25FC716EE" box="[309,429,380,408]" degrees="104" direction="east" minutes="08" orientation="longitude" pageId="27" pageNumber="28" precision="925" value="104.13333">104°08ʹ E</geoCoordinate>
;
<date id="FFAF102CFFA1FFF82087FED05CEC16EE" box="[442,646,382,408]" pageId="27" pageNumber="28" value="2018-05-10" valueMax="2018-05-14" valueMin="2018-05-10">
<collectingDate id="EFEBE9C4FFA1FFF82087FED05CEC16EE" box="[442,646,382,408]" pageId="27" pageNumber="28" value="2018-05-10" valueMax="2018-05-14" valueMin="2018-05-10">1014 May 2018</collectingDate>
</date>
;
<elevation id="003CD1DFFFA1FFF823AEFED05D3216E1" box="[659,856,381,408]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="27" pageNumber="28" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">
<quantity id="4CE99B09FFA1FFF823AEFED05D7216EE" box="[659,792,382,408]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="27" pageNumber="28" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200400 m</quantity>
a.s.l.
</elevation>
; GTI Project;
<collectorName id="26E4533AFFA1FFF8253FFED05AEE16EE" box="[1026,1156,382,408]" pageId="27" pageNumber="28">J. Constant</collectorName>
&amp;
<collectorName id="26E4533AFFA1FFF82596FED05B5616E1" box="[1195,1340,381,408]" pageId="27" pageNumber="28">P. Limbourg</collectorName>
leg.; I.G.: 33.732; “
<collectorName id="26E4533AFFA1FFF82059FE0F5C5A16CD" box="[356,560,417,443]" pageId="27" pageNumber="28">Coll. I.R.Sc.N.B.</collectorName>
, Cambodia,
<collectingRegion id="49D5F80EFFA1FFF823FFFE0F5DD116CD" box="[706,955,417,443]" country="Cambodia" name="Kampong Speu" pageId="27" pageNumber="28">Kampong Speu prov.</collectingRegion>
, Phnom Aural W.S., nr Srea Ken Vill.,
<date id="FFAF102CFFA1FFF82180FE6A5F3016A8" box="[189,346,452,478]" pageId="27" pageNumber="28" value="2018-05-10" valueMax="2018-05-14" valueMin="2018-05-10">
<collectingDate id="EFEBE9C4FFA1FFF82180FE6A5F3016A8" box="[189,346,452,478]" pageId="27" pageNumber="28" value="2018-05-10" valueMax="2018-05-14" valueMin="2018-05-10">10-14.v.2018</collectingDate>
</date>
,
<quantity id="4CE99B09FFA1FFF82055FE6A5F9B16A8" box="[360,497,452,478]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="27" pageNumber="28" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">
<elevation id="003CD1DFFFA1FFF82055FE6A5F9B16A8" box="[360,497,452,478]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="27" pageNumber="28" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200400 m</elevation>
</quantity>
,
<geoCoordinate id="EE25502BFFA1FFF820C2FE6C5C0616A8" box="[511,620,450,478]" degrees="11" direction="north" minutes="59" orientation="latitude" pageId="27" pageNumber="28" precision="925" value="11.983334">11°59ʹ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFA1FFF82347FE6C5C9916A8" box="[634,755,450,478]" degrees="104" direction="east" minutes="08" orientation="longitude" pageId="27" pageNumber="28" precision="925" value="104.13333">104°08ʹ E</geoCoordinate>
, GTI Project,
<collectorName id="26E4533AFFA1FFF822A2FE6A5A0B16A8" box="[927,1121,452,478]" pageId="27" pageNumber="28">Leg. J. Constant</collectorName>
and
<collectorName id="26E4533AFFA1FFF825A3FE6A5B5916A8" box="[1182,1331,452,478]" pageId="27" pageNumber="28">P. Limbourg</collectorName>
, I.G.: 33.732”;
<collectionCode id="ED00AE29FFA1FFF82017FE495FEE1577" box="[298,388,487,513]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="27" pageNumber="28" type="Museum">RBINS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAE36ECFFA1FFF821D7FD875F0D1535" blockId="27.[189,1399,553,719]" box="[234,359,553,579]" pageId="27" pageNumber="28">
<emphasis id="B965EAFEFFA1FFF821D7FD875F0D1535" bold="true" box="[234,359,553,579]" pageId="27" pageNumber="28">
<typeStatus id="54AA884EFFA1FFF821D7FD875F0D1535" box="[234,359,553,579]" pageId="27" pageNumber="28" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FDE25F7115B9" blockId="27.[189,1399,553,719]" pageId="27" pageNumber="28">
<materialsCitation id="3B793CB1FFA1FFF82180FDE25DD31510" box="[189,953,588,614]" collectingDate="2018-05-10" collectingDateMax="2018-05-14" collectingDateMin="2018-05-10" collectionCode="RBINS" collectorName="J. Constant &amp; P. Limbourg &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc. &amp; Leg. J. Constant" country="Cambodia" elevation="300" latitude="11.983334" location="Srea Ken Village" longLatPrecision="1294" longitude="104.13333" municipality="Phnom Aural Wildlife Sanctuary" pageId="27" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="Kampong Speu" typeStatus="paratype">
<collectingCountry id="F306767CFFA1FFF82180FDE25F311510" box="[189,347,588,614]" name="Cambodia" pageId="27" pageNumber="28">CAMBODIA</collectingCountry>
<specimenCount id="9D17FD65FFA1FFF8204EFDE25FC81510" box="[371,418,588,614]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED00AE29FFA1FFF8225EFDE25DD31510" box="[867,953,588,614]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="27" pageNumber="28" type="Museum">RBINS</collectionCode>
</materialsCitation>
<materialsCitation id="3B793CB1FFA1FFF822EFFDE25F7D15B9" collectingDate="2005-05-28" collectionCode="RBINS" collectorName="I. Var &amp; P. Grootaert &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc." country="Cambodia" latitude="13.708333" location="Kbal Spean" longLatPrecision="21" longitude="104.02556" pageId="27" pageNumber="28" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Siem Reap" typeStatus="paratype">
<specimenCount id="9D17FD65FFA1FFF822EFFDE25A681510" box="[978,1026,588,614]" count="1" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D17FD65FFA1FFF82535FDE25A581510" box="[1032,1074,588,614]" count="1" pageId="27" pageNumber="28" type="female">1 ♀</specimenCount>
;
<collectingRegion id="49D5F80EFFA1FFF8257DFDE25B581510" box="[1088,1330,588,614]" country="Cambodia" name="Siem Reap" pageId="27" pageNumber="28">Siem Reap Province</collectingRegion>
,
<location id="8ECE6037FFA1FFF82400FDE25F6215FF" LSID="urn:lsid:plazi:treatment:03B887FAFFA2FFC22377FBF75CC01653:8ECE6037FFA1FFF82400FDE25F6215FF" country="Cambodia" latitude="13.708333" longLatPrecision="21" longitude="104.02556" name="Kbal Spean" pageId="27" pageNumber="28" stateProvince="Siem Reap">Kbal Spean</location>
; [
<geoCoordinate id="EE25502BFFA1FFF8201DFDC35FD615FF" box="[288,444,621,649]" degrees="13" direction="north" minutes="42" orientation="latitude" pageId="27" pageNumber="28" precision="15" seconds="30" value="13.708333">13°42ʹ30ʺ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFA1FFF820F0FDC35C1815FF" box="[461,626,621,649]" degrees="104" direction="east" minutes="01" orientation="longitude" pageId="27" pageNumber="28" precision="15" seconds="32" value="104.02556">104°01ʹ32ʺ E</geoCoordinate>
];
<collectingMethod id="52504EFBFFA1FFF823B1FDC15C9515FF" box="[652,767,623,649]" pageId="27" pageNumber="28">light trap</collectingMethod>
;
<date id="FFAF102CFFA1FFF8222DFDC15DD015FC" box="[784,954,623,650]" pageId="27" pageNumber="28" value="2005-05-28">
<collectingDate id="EFEBE9C4FFA1FFF8222DFDC15DD015FC" box="[784,954,623,650]" pageId="27" pageNumber="28" value="2005-05-28">28 May 2005</collectingDate>
</date>
;
<collectorName id="26E4533AFFA1FFF822F6FDC15A7A15FF" box="[971,1040,623,649]" pageId="27" pageNumber="28">I. Var</collectorName>
and
<collectorName id="26E4533AFFA1FFF8256EFDC15A8915FF" box="[1107,1251,623,649]" pageId="27" pageNumber="28">P. Grootaert</collectorName>
leg.; “
<collectorName id="26E4533AFFA1FFF82406FDC15F2D15DA" pageId="27" pageNumber="28">Coll. I.R.Sc.N.B.</collectorName>
, Cambodia (
<collectingRegion id="49D5F80EFFA1FFF820E0FD3C5C0A15DB" box="[477,608,658,685]" country="Cambodia" name="Siem Reap" pageId="27" pageNumber="28">Siem Reap</collectingRegion>
prov), Kbal Spean,
<collectingMethod id="52504EFBFFA1FFF8226DFD3C5DBA15DB" box="[848,976,658,685]" pageId="27" pageNumber="28">Light Trap</collectingMethod>
,
<date id="FFAF102CFFA1FFF822E0FD3C5A3215DB" box="[989,1112,658,685]" pageId="27" pageNumber="28" value="2005-05-28">
<collectingDate id="EFEBE9C4FFA1FFF822E0FD3C5A3215DB" box="[989,1112,658,685]" pageId="27" pageNumber="28" value="2005-05-28">28.V.2005</collectingDate>
</date>
, Leg. Var &amp; Grootaert”;
<collectionCode id="ED00AE29FFA1FFF82180FD1B5F7D15B9" box="[189,279,693,719]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="27" pageNumber="28" type="Museum">RBINS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA1FFFC2180FD585FE31310" lastPageId="31" lastPageNumber="32" pageId="27" pageNumber="28" type="description">
<paragraph id="8BAE36ECFFA1FFF82180FD585F241466" blockId="27.[189,334,758,784]" box="[189,334,758,784]" pageId="27" pageNumber="28">
<heading id="D0E68180FFA1FFF82180FD585F241466" bold="true" box="[189,334,758,784]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
<emphasis id="B965EAFEFFA1FFF82180FD585F241466" bold="true" box="[189,334,758,784]" pageId="27" pageNumber="28">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FC8B5B641415" blockId="27.[189,1399,805,867]" pageId="27" pageNumber="28">
MEASUREMENTS AND RATIOS. LT:
<specimenCount id="9D17FD65FFA1FFF82374FC885C351436" box="[585,607,806,832]" count="1" pageId="27" pageNumber="28" type="male"></specimenCount>
(n=2):
<quantity id="4CE99B09FFA1FFF823F9FC8B5D1E1436" box="[708,884,805,832]" metricMagnitude="-2" metricUnit="m" metricValue="4.515" metricValueMax="4.68" metricValueMin="4.35" pageId="27" pageNumber="28" unit="mm" value="45.15" valueMax="46.8" valueMin="43.5">43.546.8 mm</quantity>
. LTg/BTg =3.3; LH/BH =5.1; LF/BF= 5.3; BH/BPrH = 2.56; LH/LT =0.43; wingspan (extrapolated):
<specimenCount id="9D17FD65FFA1FFF8225FFCE75D161415" box="[866,892,841,867]" count="1" pageId="27" pageNumber="28" type="male"></specimenCount>
:
<quantity id="4CE99B09FFA1FFF822BAFCE75D9A1415" box="[903,1008,841,867]" metricMagnitude="-2" metricUnit="m" metricValue="4.4799999999999995" pageId="27" pageNumber="28" unit="mm" value="44.8">44.8 mm</quantity>
(41.647),
<specimenCount id="9D17FD65FFA1FFF82545FCE75AFA1415" box="[1144,1168,841,867]" count="1" pageId="27" pageNumber="28" type="female"></specimenCount>
:
<quantity id="4CE99B09FFA1FFF825A3FCE65B601415" box="[1182,1290,840,867]" metricMagnitude="-2" metricUnit="m" metricValue="4.59" pageId="27" pageNumber="28" unit="mm" value="45.9">45.9 mm</quantity>
.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FC245DFB1397" blockId="27.[189,1400,906,1249]" pageId="27" pageNumber="28">
HEAD (
<figureCitation id="132A2A69FFA1FFF82033FC245FD314D2" box="[270,441,906,933]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="27" pageNumber="28">Fig. 15B, EF</figureCitation>
). Strongly elongate, representing 43% of total length, gently and evenly narrowing towards apex with apical 1/2 more or less parallel-sided in dorsal view; apex weakly widening, then obliquely cut in lateral view and with incomplete, rim-shaped orange carina (missing in ventral portion); pinkish brown densely covered in irregular very small black pitting, with some moderately larger pits mostly in basal half on sides and vertex and rarely coalescent; some slightly larger black pustules on side in distal portion, with always 23 along carina of vertex. Lateral carinae of frons indistinct in basal half, then nearly straight and slightly prominent towards apex, upcurved near apex and merging into a short dorsal carina reaching apex; lateral carinae of vertex indistinct in basal half, then rather weak and zigzagged, merging with upcurved part of lateral carinae of frons; median carina of vertex weakly marked in distal ⅓. Labium pale brown, reaching mesocoxae.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FAA65FB9121E" blockId="27.[189,1398,1288,1384]" pageId="27" pageNumber="28">
THORAX (
<figureCitation id="132A2A69FFA1FFF8200CFAA65F891254" box="[305,483,1288,1315]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="27" pageNumber="28">Fig. 15B, EF</figureCitation>
). Pro- and mesonotum pinkish brown entirely covered in moderately dense, irregular, moderate black pitting with pits mostly smaller than space between them and rarely coalescent; median carina obsolete.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180FA3E5AA11141" blockId="27.[188,1399,1424,1591]" pageId="27" pageNumber="28">
TEGMINA (
<figureCitation id="132A2A69FFA1FFF82007FA3E5F9912DC" box="[314,499,1424,1451]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="27" pageNumber="28">Fig. 15A, CD</figureCitation>
). Strongly elongate and reticulate, pale pinkish brown, progressively paler, subhyaline towards apex, basal portion densely covered with minute dark red spots; all surface irregularly covered in small to moderately large black spots, mostly on veins, larger ones slightly protruding; costal and postclaval margins subparallel, weakly diverging from base towards apex and curved at level of apex of clavus; apical margin rounded. Veins orange in large basal portion. Clavus open.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180F9F05B2211ED" blockId="27.[189,1398,1629,1692]" pageId="27" pageNumber="28">VENATION. ScP+R forked in basal ⅓, MP forked more basally in basal 1/4; CuA forked near apex of clavus; PCu and A1 fused near apex of clavus; PCu+A1 fused with postclaval margin at nodal line.</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180F96D5A341076" blockId="27.[189,1398,1731,1793]" pageId="27" pageNumber="28">
HIND WINGS (
<figureCitation id="132A2A69FFA1FFF8205BF96D5F9611AB" box="[358,508,1731,1758]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="27" pageNumber="28">Fig. 15A, C</figureCitation>
). Subhyaline, milky white with veins slightly darker, tinged with reddish brown; vein PCu brown in basal half; postclaval margin emarginate at A1 vein.
</paragraph>
<paragraph id="8BAE36ECFFA1FFF82180F8865FBF10DD" blockId="27.[189,1398,1831,1963]" pageId="27" pageNumber="28">
VENATION. ScP+R and MP forked beyond midlength; CuA forked slightly more basally then MP fork; PCu forked at basal ⅓; A1 forked rather close to base, A1 2 fused with A
<quantity id="4CE99B09FFA1FFF82530F8E55A501013" box="[1037,1082,1867,1893]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="27" pageNumber="28" unit="in" value="2.0">2 in</quantity>
distal portion and A1 2 +A2 forked slightly before reaching postclaval margin. Numerous crossveins in large portion along apical and postclaval margins.
</paragraph>
<caption id="DF6E6664FFA6FFFF2184F8065AD91093" ID-DOI="http://doi.org/10.5281/zenodo.13837078" ID-Zenodo-Dep="13837078" httpUri="https://zenodo.org/record/13837078/files/figure.png" pageId="28" pageNumber="29" startId="28.[185,226,1960,1986]" targetBox="[213,1370,261,1921]" targetPageId="28">
<paragraph id="8BAE36ECFFA6FFFF2184F8065AD91093" blockId="28.[185,1394,1960,2022]" pageId="28" pageNumber="29">
<emphasis id="B965EAFEFFA6FFFF2184F8065F7C10B4" bold="true" box="[185,278,1960,1986]" pageId="28" pageNumber="29">Fig. 17.</emphasis>
<taxonomicName id="4C114D6FFFA6FFFF2023F8075C3910B4" authority="Constant" authorityName="Constant" authorityYear="2024" box="[286,595,1960,1986]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA6FFFF2023F8075F8E10B4" box="[286,484,1960,1986]" italics="true" pageId="28" pageNumber="29">Zanna limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA6FFFF2366F8065CD210B4" box="[603,696,1960,1986]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂, in nature in Phnom Aural Wildlife Sanctuary, Cambodia, 12 May 2018.
<emphasis id="B965EAFEFFA6FFFF20D1F8655C6B1093" bold="true" box="[492,513,1995,2021]" pageId="28" pageNumber="29">A</emphasis>
. Habitus, lateral view.
<emphasis id="B965EAFEFFA6FFFF222DF8655D4E1093" bold="true" box="[784,804,1995,2021]" pageId="28" pageNumber="29">B</emphasis>
. Habitus, dorsal view.
<emphasis id="B965EAFEFFA6FFFF250CF8655A2C1093" bold="true" box="[1073,1094,1995,2021]" pageId="28" pageNumber="29">C</emphasis>
. Habitat.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFA7FFFC2180FEA75AB51630" blockId="29.[189,1399,265,467]" lastBlockId="31.[189,1399,265,327]" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
LEGS (
<figureCitation id="132A2A69FFA7FFFE2037FEA75FAB1655" box="[266,449,265,292]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="29" pageNumber="30">Fig. 15A, CD</figureCitation>
). Pinkish stramineous, pilose, rather short and robust. Pro- and mesocoxae with some black markings; all femora with small to minute black spots and conspicuous black spots aligned in a ring in distal ⅓, rarely coalescent; pro- and mesotibiae with some small black spots (some arranged into irregular ring in basal ¼), and with apex brown; pro- and mesotarsi with segments darker apically; metatibiae weakly brown basally with all spines apically black, 45 lateral spines and 8 apical spines; first metatarsomere with 1114 apical spines ventrally; second metatarsomere with 10 apical spines ventrally; first and second metatarsomere with a dense pad of microsetae ventrally; third metatarsomere elongate with a subapical blackish brown ring. Metatibiotarsal formula: (45) 8/1114/10.
</paragraph>
<caption id="DF6E6664FFA7FFFE2182F8F65FA310AA" ID-DOI="http://doi.org/10.5281/zenodo.13837080" ID-Zenodo-Dep="13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="29" pageNumber="30" startId="29.[191,234,1880,1906]" targetBox="[185,1406,518,1841]" targetPageId="29">
<paragraph id="8BAE36ECFFA7FFFE2182F8F65FA310AA" blockId="29.[190,1401,1880,2013]" pageId="29" pageNumber="30">
<emphasis id="B965EAFEFFA7FFFE2182F8F65F711005" bold="true" box="[191,283,1880,1907]" pageId="29" pageNumber="30">Fig. 18.</emphasis>
<taxonomicName id="4C114D6FFFA7FFFE201CF8F45F061005" authorityName="Kirkaldy" authorityYear="1902" box="[289,364,1882,1907]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFA7FFFE201CF8F45F061005" box="[289,364,1882,1907]" italics="true" pageId="29" pageNumber="30">Zanna</emphasis>
</taxonomicName>
, new spp. from Cambodia and Vietnam, key characters.
<emphasis id="B965EAFEFFA7FFFE22C6F8F65A5E1004" bold="true" box="[1019,1076,1880,1906]" pageId="29" pageNumber="30">AD</emphasis>
. Prothotax and basal half of head, lateral view.
<emphasis id="B965EAFEFFA7FFFE20A5F8D25FBB10E0" bold="true" box="[408,465,1916,1942]" pageId="29" pageNumber="30">EH</emphasis>
. Head and thorax, dorsal view.
<emphasis id="B965EAFEFFA7FFFE2200F8D25D0010E0" bold="true" box="[829,874,1916,1942]" pageId="29" pageNumber="30">IL</emphasis>
. Right tegmen.
<emphasis id="B965EAFEFFA7FFFE251CF8D25A1A10E0" bold="true" box="[1057,1136,1916,1942]" pageId="29" pageNumber="30">A, E, I</emphasis>
.
<taxonomicName id="4C114D6FFFA7FFFE2542F8D35B7010E0" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1151,1306,1916,1942]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFA7FFFE2542F8D35B7010E0" box="[1151,1306,1916,1942]" italics="true" pageId="29" pageNumber="30">Z. bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA7FFFE241DF8D25B1310E0" box="[1312,1401,1916,1942]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA7FFFE2183F8315F7810CF" bold="true" box="[190,274,1951,1977]" pageId="29" pageNumber="30">B, F, J</emphasis>
.
<taxonomicName id="4C114D6FFFA7FFFE2019F80E5C4A10CF" authority="Constant" authorityName="Constant" authorityYear="2024" box="[292,544,1951,1978]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="34" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
Z.
<emphasis id="B965EAFEFFA7FFFE2075F8315FC510CF" box="[328,431,1951,1977]" italics="true" pageId="29" pageNumber="30">chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA7FFFE2317F80E5CED10CC" box="[554,647,1952,1978]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA7FFFE23ACF8315C9410CF" bold="true" box="[657,766,1951,1977]" pageId="29" pageNumber="30">C, G, K.</emphasis>
<taxonomicName id="4C114D6FFFA7FFFE223AF80E5DFA10CF" authorityName="Constant &amp; Pham" authorityYear="2024" box="[775,912,1951,1977]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFA7FFFE223AF80E5DFA10CF" box="[775,912,1951,1977]" italics="true" pageId="29" pageNumber="30">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA7FFFE22A4F80E5D9C10CC" box="[921,1014,1952,1978]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA7FFFE253DF8315A0810CF" bold="true" box="[1024,1122,1951,1977]" pageId="29" pageNumber="30">D, H, L</emphasis>
.
<taxonomicName id="4C114D6FFFA7FFFE254EF80E5B1310CF" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1139,1401,1951,1977]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA7FFFE254EF80E5B6D10CF" box="[1139,1287,1951,1977]" italics="true" pageId="29" pageNumber="30">Z. limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA7FFFE2182F86D5F7710AB" box="[191,285,1987,2013]" pageId="29" pageNumber="30" rank="species">sp. nov.</taxonomicNameLabel>
, (not to scale).
</paragraph>
</caption>
<caption id="DF6E6664FFA4FFFD21FCF8CA5CDC109E" ID-DOI="http://doi.org/10.5281/zenodo.13837082" ID-Zenodo-Dep="13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="30" pageNumber="31" startId="30.[193,236,1892,1918]" targetBox="[193,1402,242,1853]" targetPageId="30">
<paragraph id="8BAE36ECFFA4FFFD21FCF8CA5CDC109E" blockId="30.[193,1403,1892,2024]" pageId="30" pageNumber="31">
<emphasis id="B965EAFEFFA4FFFD21FCF8CA5F761008" bold="true" box="[193,284,1892,1918]" pageId="30" pageNumber="31">Fig. 19.</emphasis>
<taxonomicName id="4C114D6FFFA4FFFD201DF8CB5F011008" authorityName="Kirkaldy" authorityYear="1902" box="[288,363,1893,1918]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFA4FFFD201DF8CB5F011008" box="[288,363,1893,1918]" italics="true" pageId="30" pageNumber="31">Zanna</emphasis>
</taxonomicName>
, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli.
<emphasis id="B965EAFEFFA4FFFD21FCF8295F2410D7" bold="true" box="[193,334,1927,1953]" pageId="30" pageNumber="31">A, C, E, G</emphasis>
. Lateral view.
<emphasis id="B965EAFEFFA4FFFD2334F8295CFA10D7" bold="true" box="[521,656,1927,1953]" pageId="30" pageNumber="31">B, D, F, H</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFA4FFFD2278F8295D1710D7" bold="true" box="[837,893,1927,1953]" pageId="30" pageNumber="31">AB</emphasis>
.
<taxonomicName id="4C114D6FFFA4FFFD22ADF8265A5A10D7" authorityName="Constant &amp; Pham" authorityYear="2024" box="[912,1072,1927,1953]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFA4FFFD22ADF8265A5A10D7" box="[912,1072,1927,1953]" italics="true" pageId="30" pageNumber="31">Z. bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA4FFFD2506F8265AF010D4" box="[1083,1178,1928,1954]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA4FFFD2598F8295AB410D7" bold="true" box="[1189,1246,1927,1953]" pageId="30" pageNumber="31">CD</emphasis>
.
<taxonomicName id="4C114D6FFFA4FFFD25CCF8265F4310B3" authority="Constant" authorityName="Constant" authorityYear="2024" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFA4FFFD25CCF8265B1110D7" box="[1265,1403,1927,1953]" italics="true" pageId="30" pageNumber="31">Z. chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA4FFFD200FF8055FE410B3" box="[306,398,1963,1989]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA4FFFD20A5F8055FA110B3" bold="true" box="[408,459,1963,1989]" pageId="30" pageNumber="31">EF</emphasis>
.
<taxonomicName id="4C114D6FFFA4FFFD20E6F8055C0910B2" authorityName="Constant &amp; Pham" authorityYear="2024" box="[475,611,1962,1988]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFA4FFFD20E6F8055C0910B2" box="[475,611,1962,1988]" italics="true" pageId="30" pageNumber="31">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA4FFFD2356F8055CAD10B3" box="[619,711,1963,1989]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B965EAFEFFA4FFFD23ECF8045D6610B2" bold="true" box="[721,780,1962,1988]" pageId="30" pageNumber="31">GH</emphasis>
.
<taxonomicName id="4C114D6FFFA4FFFD2221F8055A4A10B3" authority="Constant" authorityName="Constant" authorityYear="2024" box="[796,1056,1962,1989]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA4FFFD2221F8055DDA10B2" box="[796,944,1962,1988]" italics="true" pageId="30" pageNumber="31">Z. limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA4FFFD2514F8055AEC10B3" box="[1065,1158,1963,1989]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
(scale bars =1 mm). Abbreviations: see Materials and methods.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFA5FFFC2180FEC05DCD1684" blockId="31.[189,1399,365,498]" pageId="31" pageNumber="32">
ABDOMEN (
<figureCitation id="132A2A69FFA5FFFC207DFEC05FA116FE" box="[320,459,365,392]" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="31" pageNumber="32">Fig. 15A, C</figureCitation>
). Tergites smooth, segments blackish brown with more or less developed marking at posterior angles, and posterior margin, yellowish; sternites (except pregenital one) yellowish brown with irregular small black dots arranged in two groups in middle portion, and two groups on side of each segment, often with two black dots anteriorly to middle groups.
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180FDB75FE31310" blockId="31.[189,1399,536,1126]" pageId="31" pageNumber="32">
MALE TERMINALIA (
<figureCitation id="132A2A69FFA5FFFC2090FDB75C621544" box="[429,520,536,563]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16</figureCitation>
). Pygofer (
<emphasis id="B965EAFEFFA5FFFC23AFFDB75CD81544" box="[658,690,537,562]" italics="true" pageId="31" pageNumber="32">Py</emphasis>
<figureCitation id="132A2A69FFA5FFFC23ECFDB75D0E1544" box="[721,868,536,563]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16AD</figureCitation>
) about 1.69 times as high as long in lateral view, robust with posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle; sides weakly excavate in dorsal portion (to observe in dorsal aspect,
<figureCitation id="132A2A69FFA5FFFC2183FD2C5F4615EA" box="[190,300,642,668]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16C</figureCitation>
). Anal tube (
<emphasis id="B965EAFEFFA5FFFC20FAFD2D5F8215EB" box="[455,488,643,669]" italics="true" pageId="31" pageNumber="32">An</emphasis>
<figureCitation id="132A2A69FFA5FFFC2338FD2C5CFF15EA" box="[517,661,642,668]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16AC</figureCitation>
) robust, moderately flattened, 1.65 times as long as high and with ventral margin oblique in distal half and moderately angular before midlength in lateral view; in dorsal view, 1.08 times as long as wide, with sides sinuate, subparallel in basal portion, then roundly diverging towards apex, forming rounded posterior angles and with apical margin incurved; suboval in caudal view, flattened dorsally; epiproct and paraproct rather strongly surpassing anal tube, subtriangular and rounded posteriorly in dorsal aspect. Gonostyli (
<emphasis id="B965EAFEFFA5FFFC250DFC9D5A2F143B" box="[1072,1093,819,845]" italics="true" pageId="31" pageNumber="32">G</emphasis>
<figureCitation id="132A2A69FFA5FFFC2559FC9C5A9C143A" box="[1124,1270,818,844]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16AD</figureCitation>
) elongate, 2.58 times as long as high; in lateral view, ventral margin broadly rounded and dorsal margin rather broadly rounded after lateral hook (
<emphasis id="B965EAFEFFA5FFFC2340FCD75CCE14E5" box="[637,676,889,915]" italics="true" pageId="31" pageNumber="32">lhg</emphasis>
), apical margin rather broadly rounded; lateral hook with basal portion projecting posterolaterad and apex strongly curved lateroventrad; in ventral view, lateral protrusion at about midlength, rather strongly, angularly marked and with posterior angle curved. Aedeagus (
<figureCitation id="132A2A69FFA5FFFC2094FC4C5C51148A" box="[425,571,994,1020]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="31" pageNumber="32">Fig. 16EH</figureCitation>
) moderately elongate with lateral processes (
<emphasis id="B965EAFEFFA5FFFC2551FC4C5AF9148A" box="[1132,1171,994,1020]" italics="true" pageId="31" pageNumber="32">lpa</emphasis>
) forming multiple lobes; dorsal process (
<emphasis id="B965EAFEFFA5FFFC20E8FBAB5C681369" box="[469,514,1029,1055]" italics="true" pageId="31" pageNumber="32">dpa</emphasis>
) weakly protruding dorsally in lateral view; dorsal endosomal process (
<emphasis id="B965EAFEFFA5FFFC21FAFB875E991335" box="[199,243,1065,1091]" italics="true" pageId="31" pageNumber="32">dep</emphasis>
) in dorsal view subtriangular with lateral margins nearly straight and rather wide median slit in distal portion.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA5FFFC2180FB235A7C126A" pageId="31" pageNumber="32" type="biology_ecology">
<paragraph id="8BAE36ECFFA5FFFC2180FB235F7613D1" blockId="31.[189,284,1165,1191]" box="[189,284,1165,1191]" pageId="31" pageNumber="32">
<heading id="D0E68180FFA5FFFC2180FB235F7613D1" bold="true" box="[189,284,1165,1191]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B965EAFEFFA5FFFC2180FB235F7613D1" bold="true" box="[189,284,1165,1191]" pageId="31" pageNumber="32">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180FB125A7C126A" blockId="31.[189,1399,1212,1308]" pageId="31" pageNumber="32">
The specimens from
<collectingRegion id="49D5F80EFFA5FFFC2089FB125C5913A0" box="[436,563,1212,1238]" country="Cambodia" name="Siem Reap" pageId="31" pageNumber="32">Siem Reap</collectingRegion>
were attracted to a light trap in a strongly anthropized area while those from Phnom Aural were found sitting on the trunk of an unidentified tree (
<figureCitation id="132A2A69FFA5FFFC2578FB715ABE138F" box="[1093,1236,1247,1273]" captionStart="Fig" captionStartId="28.[185,226,1960,1986]" captionTargetBox="[213,1370,261,1921]" captionTargetId="figure-36@28.[213,1370,258,1921]" captionTargetPageId="28" captionText="Fig. 17. Zanna limbourgi Constant sp. nov., holotype, ♂, in nature in Phnom Aural Wildlife Sanctuary, Cambodia, 12 May 2018. A. Habitus, lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi="http://doi.org/10.5281/zenodo.13837078" httpUri="https://zenodo.org/record/13837078/files/figure.png" pageId="31" pageNumber="32">Fig. 17AB</figureCitation>
) about
<quantity id="4CE99B09FFA5FFFC240CFB715B1D138C" box="[1329,1399,1247,1274]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="m" value="1.5">1.5 m</quantity>
above the ground in rather heavily disturbed secondary forest (
<figureCitation id="132A2A69FFA5FFFC22A0FAAC5A63126A" box="[925,1033,1282,1308]" captionStart="Fig" captionStartId="28.[185,226,1960,1986]" captionTargetBox="[213,1370,261,1921]" captionTargetId="figure-36@28.[213,1370,258,1921]" captionTargetPageId="28" captionText="Fig. 17. Zanna limbourgi Constant sp. nov., holotype, ♂, in nature in Phnom Aural Wildlife Sanctuary, Cambodia, 12 May 2018. A. Habitus, lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi="http://doi.org/10.5281/zenodo.13837078" httpUri="https://zenodo.org/record/13837078/files/figure.png" pageId="31" pageNumber="32">Fig. 17C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA5FFFC2180FAED5A0812A7" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8BAE36ECFFA5FFFC2180FAED5F3C122B" blockId="31.[189,342,1347,1373]" box="[189,342,1347,1373]" pageId="31" pageNumber="32">
<heading id="D0E68180FFA5FFFC2180FAED5F3C122B" bold="true" box="[189,342,1347,1373]" fontSize="11" level="3" pageId="31" pageNumber="32" reason="3">
<emphasis id="B965EAFEFFA5FFFC2180FAED5F3C122B" bold="true" box="[189,342,1347,1373]" pageId="31" pageNumber="32">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180FADC5DFE12FB" blockId="31.[189,916,1394,1421]" box="[189,916,1394,1421]" pageId="31" pageNumber="32">
<collectingCountry id="F306767CFFA5FFFC2180FADC5F5012FA" box="[189,314,1394,1420]" name="Cambodia" pageId="31" pageNumber="32">Cambodia</collectingCountry>
:
<collectingRegion id="49D5F80EFFA5FFFC2078FADD5F9112FA" box="[325,507,1394,1421]" country="Cambodia" name="Kampong Speu" pageId="31" pageNumber="32">Kampong Speu</collectingRegion>
and
<collectingRegion id="49D5F80EFFA5FFFC2309FADC5CD912FB" box="[564,691,1394,1421]" country="Cambodia" name="Siem Reap" pageId="31" pageNumber="32">Siem Reap</collectingRegion>
provinces (
<figureCitation id="132A2A69FFA5FFFC2200FADC5DEF12FB" box="[829,901,1394,1421]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="31" pageNumber="32">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180FA1A5A0812A7" blockId="31.[189,1122,1460,1489]" box="[189,1122,1460,1489]" pageId="31" pageNumber="32">
<emphasis id="B965EAFEFFA5FFFC2180FA1A5A0812A7" bold="true" box="[189,1122,1460,1489]" italics="true" pageId="31" pageNumber="32">
Identification key to the species of
<taxonomicName id="4C114D6FFFA5FFFC23B9FA1B5CB112A7" authorityName="Kirkaldy" authorityYear="1902" box="[644,731,1461,1489]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">Zanna</taxonomicName>
from
<collectingCountry id="F306767CFFA5FFFC2216FA1A5DD812A7" box="[811,946,1460,1489]" name="Cambodia" pageId="31" pageNumber="32">Cambodia</collectingCountry>
and
<collectingCountry id="F306767CFFA5FFFC22C9FA1B5A0812A7" box="[1012,1122,1461,1489]" name="Vietnam" pageId="31" pageNumber="32">Vietnam</collectingCountry>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA5FFC22180FA475CC01653" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="key">
<key id="FF8061DDFFA5FFC32180FA475B1D1479" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<keyStep id="30E52E49FFA5FFFC2180FA475B1C111B" pageId="31" pageNumber="32">
<paragraph id="8BAE36ECFFA5FFFC2180FA475B1D1150" blockId="31.[189,1399,1513,1645]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180FA475B1D1150" pageId="31" pageNumber="32">
1. Smaller, wingspan up to
<quantity id="4CE99B09FFA5FFFC232DFA475C0D1172" box="[528,615,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="32" unit="mm" value="55.0">55 mm</quantity>
, usually less than
<quantity id="4CE99B09FFA5FFFC2203FA475DFE1172" box="[830,916,1513,1540]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="mm" value="50.0">50 mm</quantity>
; cephalic process exceeding 1.3 times as long as abdomen ................................................................................................................................ 2
</keyLead>
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180F99E5B1C111B" blockId="31.[189,1399,1513,1645]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180F99E5B1C111B" pageId="31" pageNumber="32">
Larger, wingspan
<quantity id="4CE99B09FFA5FFFC20FDF9815C29113C" box="[448,579,1583,1610]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" metricValueMax="6.5" metricValueMin="6.0" pageId="31" pageNumber="32" unit="mm" value="62.5" valueMax="65.0" valueMin="60.0">6065 mm</quantity>
; cephalic process less than 1.3 times as long as abdomen (
<figureCitation id="132A2A69FFA5FFFC25C0F9815E94111B" captionStart="Fig" captionStartId="32.[188,229,1947,1973]" captionTargetBox="[189,1398,864,1901]" captionTargetId="figure-207@32.[186,1401,864,1901]" captionTargetPageId="32" captionText="Fig. 20. Zanna chinensis (Distant, 1893), syntype, ♀ (BMNH). A. Dorsal view. B. Lateral view. C. Labels (photograph by D. Rendon-Mera, © The Trustees of the Natural History Museum, London)." figureDoi="http://doi.org/10.5281/zenodo.13837084" httpUri="https://zenodo.org/record/13837084/files/figure.png" pageId="31" pageNumber="32">Fig. 20A B</figureCitation>
) ............................................................................................................
<taxonomicName id="4C114D6FFFA5FFFC2516F9FD5B1C111B" ID-CoL="5CW46" authority="(Distant, 1893)" baseAuthorityName="Distant" baseAuthorityYear="1893" box="[1067,1398,1618,1645]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B965EAFEFFA5FFFC2516F9FD5AD7111B" bold="true" box="[1067,1213,1618,1645]" italics="true" pageId="31" pageNumber="32">Z. chinensis</emphasis>
(
<bibRefCitation id="EF804B1DFFA5FFFC25F0F9FD5B04111B" author="Distant W. L." box="[1229,1390,1619,1645]" pageId="31" pageNumber="32" pagination="443 - 449" refId="ref13113" refString="Distant W. L. 1893. On the Homopterous genus Pyrops, with descriptions of two new species. Transactions of the Entomological Society of London 1893: 443 - 449." type="journal article" year="1893">Distant, 1893</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E49FFA5FFFC2180F93A5B1D106E" pageId="31" pageNumber="32">
<paragraph id="8BAE36ECFFA5FFFC2180F93A5B1D11A7" blockId="31.[189,1399,1684,1816]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180F93A5B1D11A7" pageId="31" pageNumber="32">
2. Anal tube of male rather flattened dorsoventrally, 1.952.14 times as long as high in lateral view; ventral angle of anal tube at midlength (
<figureCitation id="132A2A69FFA5FFFC2382F9195D2311A7" box="[703,841,1719,1745]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="31" pageNumber="32">Fig. 19C, E</figureCitation>
) ....................................................................... 3
</keyLead>
</paragraph>
<paragraph id="8BAE36ECFFA5FFFC2180F9755B1D106E" blockId="31.[189,1399,1684,1816]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180F9755B1D106E" pageId="31" pageNumber="32">
Anal tube of male rather high, 1.601.65 times as long as high in lateral view; ventral angle of anal tube before midlength (
<figureCitation id="132A2A69FFA5FFFC20C0F9505CE1106E" box="[509,651,1790,1816]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="31" pageNumber="32">Fig. 19A, G</figureCitation>
) ................................................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E49FFA5FFC32180F8915B1D16FB" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<paragraph id="8BAE36ECFFA5FFFC2180F8915B1D1091" blockId="31.[189,1399,1855,2023]" pageId="31" pageNumber="32">
<keyLead id="30E095D9FFA5FFFC2180F8915B1D1091" pageId="31" pageNumber="32">
3. Pygofer with posterior margin projecting into a large rounded posterior lobe in dorsal half (
<figureCitation id="132A2A69FFA5FFFC243BF8915B04102F" box="[1286,1390,1855,1881]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="31" pageNumber="32">Fig. 19C</figureCitation>
) and with sides rather strongly excavate in dorsal portion (
<figureCitation id="132A2A69FFA5FFFC22A9F8CC5A68100B" box="[916,1026,1890,1917]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="31" pageNumber="32">Fig. 19D</figureCitation>
); basal portion of head covered in dense, irregular black pitting with smaller pits and some dispersed, moderately large, not or very rarely coalescent pits (
<figureCitation id="132A2A69FFA5FFFC20CFF8075C1610B5" box="[498,636,1961,1987]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="31" pageNumber="32">Fig. 18B, F</figureCitation>
); tegmina more elongate, 3.80 times as long as wide (
<figureCitation id="132A2A69FFA5FFFC25CEF8075B3E10B5" box="[1267,1364,1961,1987]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="31" pageNumber="32">Fig. 18J</figureCitation>
) .. ..............................................................................................................
<taxonomicName id="4C114D6FFFA5FFFC2521F8625B7F1090" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1052,1301,1995,2022]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFA5FFFC2521F8625ACF1090" bold="true" box="[1052,1189,1995,2022]" italics="true" pageId="31" pageNumber="32">Z. chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA5FFFC2421F8635B1D1091" box="[1308,1399,1997,2023]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAE36ECFF9AFFC32180FEA75B1D16FB" blockId="32.[189,1399,265,397]" pageId="32" pageNumber="33">
<keyLead id="30E095D9FF9AFFC32180FEA75B1D16FB" pageId="32" pageNumber="33">
Pygofer with posterior margin projecting into a large posterior lobe angularly rounded in midheight (
<figureCitation id="132A2A69FF9AFFC321CEFE825F311630" box="[243,347,300,326]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19E</figureCitation>
) and with sides weakly excavate in dorsal portion (
<figureCitation id="132A2A69FF9AFFC322FDFE825A431630" box="[960,1065,300,326]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19F</figureCitation>
); basal portion of head with irregular, moderately large, rather well-spaced, black pitting but without background of smaller pits (
<figureCitation id="132A2A69FF9AFFC321CEFEDC5FEA16FB" box="[243,384,370,397]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="32" pageNumber="33">Fig. 18C, G</figureCitation>
); tegmina less elongate, 3.25 times as long as wide (
<figureCitation id="132A2A69FF9AFFC322DCFEDC5A2016FB" box="[993,1098,370,397]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="32" pageNumber="33">Fig. 18K</figureCitation>
).......
<taxonomicName id="4C114D6FFF9AFFC325B6FEDD5B7F16FB" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1163,1301,370,397]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFF9AFFC325B6FEDD5B7F16FB" bold="true" box="[1163,1301,370,397]" italics="true" pageId="32" pageNumber="33">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FF9AFFC32420FEDD5B1D16FB" box="[1309,1399,371,397]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E52E49FF9AFFC32180FE1A5B1D1479" pageId="32" pageNumber="33">
<paragraph id="8BAE36ECFF9AFFC32180FE1A5B1D152D" blockId="32.[189,1399,436,783]" pageId="32" pageNumber="33">
<keyLead id="30E095D9FF9AFFC32180FE1A5B1D152D" pageId="32" pageNumber="33">
4. Pygofer in lateral view with posterior margin sinuate, forming a large, broadly rounded posterior lobe in dorsal half (
<figureCitation id="132A2A69FF9AFFC320F5FE795C591687" box="[456,563,471,497]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19A</figureCitation>
); anal tube in dorsal view 1.2 times as long as wide and with sides nearly straight, weakly diverging towards apex (
<figureCitation id="132A2A69FF9AFFC323FAFE545D5B1563" box="[711,817,506,533]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19B</figureCitation>
); basal portion of head covered in dense, irregular, rather large, coarse, sometimes coalescent black pitting with few smaller pits (
<figureCitation id="132A2A69FF9AFFC325BDFDB05B66154E" box="[1152,1292,542,568]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="32" pageNumber="33">Fig. 18A, E</figureCitation>
) ............ ..........................................................................................................................
<taxonomicName id="4C114D6FFF9AFFC32549FDEF5B7F152D" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1140,1301,576,603]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFF9AFFC32549FDEF5B7F152D" bold="true" box="[1140,1301,576,603]" italics="true" pageId="32" pageNumber="33">Z. bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FF9AFFC32421FDEF5B1D152D" box="[1308,1399,577,603]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BAE36ECFF9AFFC32180FDCB5B1D1479" blockId="32.[189,1399,436,783]" pageId="32" pageNumber="33">
<keyLead id="30E095D9FF9AFFC32180FDCB5B1D1479" pageId="32" pageNumber="33">
Pygofer in lateral view with posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle (
<figureCitation id="132A2A69FF9AFFC323DFFD295D3815D7" box="[738,850,647,673]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19G</figureCitation>
); anal tube in dorsal view 1.08 times as long as wide, with sides roundly diverging towards apex (
<figureCitation id="132A2A69FF9AFFC32240FD045D9A15B3" box="[893,1008,682,709]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="32" pageNumber="33">Fig. 19H</figureCitation>
); basal portion of head densely covered in irregular, very small black pitting, with some moderately larger pits, rarely coalescent (
<figureCitation id="132A2A69FF9AFFC321CEFD5A5FE81478" box="[243,386,756,782]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="32" pageNumber="33">Fig. 18D, H</figureCitation>
).......................................................................................
<taxonomicName id="4C114D6FFF9AFFC32533FD5A5B7F1478" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1038,1301,755,782]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFF9AFFC32533FD5A5ACF1478" bold="true" box="[1038,1189,755,782]" italics="true" pageId="32" pageNumber="33">Z. limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FF9AFFC32421FD5B5B1D1479" box="[1308,1399,757,783]" pageId="32" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
<caption id="DF6E6664FF9AFFC32181F8355B0710AF" ID-DOI="http://doi.org/10.5281/zenodo.13837084" ID-Zenodo-Dep="13837084" httpUri="https://zenodo.org/record/13837084/files/figure.png" pageId="32" pageNumber="33" startId="32.[188,229,1947,1973]" targetBox="[189,1398,864,1901]" targetPageId="32">
<paragraph id="8BAE36ECFF9AFFC32181F8355B0710AF" blockId="32.[188,1398,1947,2009]" pageId="32" pageNumber="33">
<emphasis id="B965EAFEFF9AFFC32181F8355F7510C0" bold="true" box="[188,287,1947,1974]" pageId="32" pageNumber="33">Fig. 20.</emphasis>
<taxonomicName id="4C114D6FFF9AFFC32010F8325CD310C0" authority="(Distant, 1893)" baseAuthorityName="Distant" baseAuthorityYear="1893" box="[301,697,1947,1974]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B965EAFEFF9AFFC32010F8325F9810C3" box="[301,498,1947,1973]" italics="true" pageId="32" pageNumber="33">Zanna chinensis</emphasis>
(
<bibRefCitation id="EF804B1DFF9AFFC3233AF8355CDA10C0" author="Distant W. L." box="[519,688,1947,1974]" pageId="32" pageNumber="33" pagination="443 - 449" refId="ref13113" refString="Distant W. L. 1893. On the Homopterous genus Pyrops, with descriptions of two new species. Transactions of the Entomological Society of London 1893: 443 - 449." type="journal article" year="1893">Distant, 1893</bibRefCitation>
)
</taxonomicName>
, syntype, ♀ (BMNH).
<emphasis id="B965EAFEFF9AFFC322E3F8355D9910C3" bold="true" box="[990,1011,1947,1973]" pageId="32" pageNumber="33">A</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFF9AFFC32592F8325AA910C0" bold="true" box="[1199,1219,1948,1974]" pageId="32" pageNumber="33">B</emphasis>
. Lateral view.
<emphasis id="B965EAFEFF9AFFC32181F8105EBB10AE" bold="true" box="[188,209,1982,2008]" pageId="32" pageNumber="33">C</emphasis>
. Labels (photograph by D. Rendon-Mera, © The Trustees of the Natural History Museum, London).
</paragraph>
</caption>
<paragraph id="8BAE36ECFF9BFFC22180FEA65CC01653" blockId="33.[189,682,264,293]" box="[189,682,264,293]" pageId="33" pageNumber="34">
<emphasis id="B965EAFEFF9BFFC22180FEA65CC01653" bold="true" box="[189,682,264,293]" italics="true" pageId="33" pageNumber="34">Other species examined for this study</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>

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<document id="BB47138F8E3CBD6B9954858DC49C7119" ID-DOI="10.5852/ejt.2024.958.2665" ID-ISSN="2118-9773" ID-Zenodo-Dep="13837028" ID-ZooBank="425BCF68-144B-4861-BCED-CB570D8246D1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727252768769" checkinUser="plazi" docAuthor="Constant, Jérôme &amp; Pham, Hong Thai" docDate="2024" docId="03B887FAFFA9FFFB23BAFAAA5C131362" docLanguage="en" docName="EJT.2024.958.114-150.pdf" docOrigin="European Journal of Taxonomy 958" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2665/12349" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Zanna kusamae Constant &amp; Pham, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="25" masterDocId="FF81FF82FFBAFFE3213DFFAE5E6A1776" masterDocTitle="Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)" masterLastPageNumber="150" masterPageNumber="114" pageNumber="20" updateTime="1727253238722" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="695F9810E744B45C089437B1C01D1281">Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<mods:namePart id="C012BEF4A597DCCF4BEDC9DAC4065299">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="86008B95F06790DBE13C5F72570C5287">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier Street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
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<mods:namePart id="FA029A91845B7B262B04E8FE2967A97A">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="CB5D465A52F7F89C8A7C25BE04B23D9C">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.</mods:affiliation>
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<paragraph id="8BAE36ECFFA9FFF023BAFAAA5DCF1268" blockId="19.[348,1240,1283,1385]" box="[647,933,1283,1310]" pageId="19" pageNumber="20">
<heading id="D0E68180FFA9FFF023BAFAAA5DCF1268" box="[647,933,1283,1310]" centered="true" fontSize="11" level="2" pageId="19" pageNumber="20" reason="2">
<taxonomicName id="4C114D6FFFA9FFF023BAFAAA5D201268" authorityName="Constant &amp; Pham" authorityYear="2024" box="[647,842,1283,1310]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFA9FFF023BAFAAA5D201268" bold="true" box="[647,842,1283,1310]" italics="true" pageId="19" pageNumber="20">Zanna kusamae</emphasis>
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sp.
<emphasis id="B965EAFEFFA9FFF0224EFAAA5DCF1268" bold="true" box="[883,933,1284,1310]" italics="true" pageId="19" pageNumber="20">nov.</emphasis>
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<subSubSection id="C30B6567FFA9FFF0236EFAE35D8B121F" box="[595,993,1356,1385]" pageId="19" pageNumber="20" type="description">
<paragraph id="8BAE36ECFFA9FFF0236EFAE35D8B121F" blockId="19.[348,1240,1283,1385]" box="[595,993,1356,1385]" pageId="19" pageNumber="20">
<figureCitation id="132A2A69FFA9FFF0236EFAE35CF7121E" box="[595,669,1357,1384]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="19" pageNumber="20">Figs 5</figureCitation>
,
<figureCitation id="132A2A69FFA9FFF02397FAE35C991211" box="[682,755,1357,1383]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="21.[188,229,1927,1953]" captionStartId-1="22.[189,230,1926,1952]" captionStartId-2="23.[189,230,1950,1976]" captionTargetBox-0="[215,1367,257,1885]" captionTargetBox-1="[218,1365,253,1887]" captionTargetBox-2="[187,1398,522,1886]" captionTargetId-0="figure-46@21.[210,1367,254,1885]" captionTargetId-1="figure-62@22.[214,1371,238,1891]" captionTargetId-2="figure-106@23.[187,1401,522,1886]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionTargetPageId-2="23" captionText-0="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." captionText-1="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." captionText-2="Fig. 14. Zanna kusamae sp. nov., holotype, ♂, in nature, Dong Nai Biosphere Reserve, Vietnam, 3 Jul. 2012. A. Habitus lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi-0="http://doi.org/10.5281/zenodo.13837062" figureDoi-1="http://doi.org/10.5281/zenodo.13837066" figureDoi-2="http://doi.org/10.5281/zenodo.13837070" httpUri-0="https://zenodo.org/record/13837062/files/figure.png" httpUri-1="https://zenodo.org/record/13837066/files/figure.png" httpUri-2="https://zenodo.org/record/13837070/files/figure.png" pageId="19" pageNumber="20">1214</figureCitation>
,
<figureCitation id="132A2A69FFA9FFF0223FFAE25DE8121E" box="[770,898,1356,1385]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="19" pageNumber="20">18C, G, K</figureCitation>
,
<figureCitation id="132A2A69FFA9FFF022B3FAE25D8B121F" box="[910,993,1356,1385]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="19" pageNumber="20">19EF</figureCitation>
</paragraph>
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<paragraph id="8BAE36ECFFA9FFF02180FA205F5C12DE" blockId="19.[189,310,1422,1448]" box="[189,310,1422,1448]" pageId="19" pageNumber="20">
<heading id="D0E68180FFA9FFF02180FA205F5C12DE" bold="true" box="[189,310,1422,1448]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B965EAFEFFA9FFF02180FA205F5C12DE" bold="true" box="[189,310,1422,1448]" pageId="19" pageNumber="20">Diagnosis</emphasis>
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<paragraph id="8BAE36ECFFA9FFF02180FA135D8B100B" blockId="19.[188,1399,1469,1918]" pageId="19" pageNumber="20">
The species is closest to
<taxonomicName id="4C114D6FFFA9FFF020D0FA105CD512A1" authorityName="Constant &amp; Pham" authorityYear="2024" box="[493,703,1469,1495]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFA9FFF020D0FA105CD512A1" box="[493,703,1469,1495]" italics="true" pageId="19" pageNumber="20">Zanna bidoupana</emphasis>
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<taxonomicNameLabel id="A2565785FFA9FFF023F7FA105D4012AE" box="[714,810,1470,1496]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C114D6FFFA9FFF02204FA105A5E12A1" authority="Constant" authorityName="Constant" authorityYear="2024" box="[825,1076,1469,1495]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFA9FFF02204FA105DA812A1" box="[825,962,1469,1495]" italics="true" pageId="19" pageNumber="20">Z. chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA9FFF02503FA105AF412AE" box="[1086,1182,1470,1496]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, and
<taxonomicName id="4C114D6FFFA9FFF025DFFA105F4F128D" authority="Constant" authorityName="Constant" authorityYear="2024" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFA9FFF025DFFA105B1D12A1" box="[1250,1399,1469,1495]" italics="true" pageId="19" pageNumber="20">Z.limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFA9FFF02012FA4F5FE5128D" box="[303,399,1505,1531]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can be separated by the following characters: basal portion of head (
<figureCitation id="132A2A69FFA9FFF021FBF9AD5F1A1168" box="[198,368,1539,1566]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="19" pageNumber="20">Fig. 12B, EF</figureCitation>
) covered in moderately large, rather well-spaced, black pitting without background of dense, very small black pitting (dense, irregular, rather large, coarse, sometimes coalescent black pitting in
<taxonomicName id="4C114D6FFFA9FFF021E1F9E55F131112" authorityName="Constant &amp; Pham" authorityYear="2024" box="[220,377,1610,1636]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bidoupana">
<emphasis id="B965EAFEFFA9FFF021E1F9E55F131112" box="[220,377,1610,1636]" italics="true" pageId="19" pageNumber="20">Z. bidoupana</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA9FFF020A4F9E45C5C1113" box="[409,566,1610,1637]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="19" pageNumber="20">Fig. 3B, EF</figureCitation>
; moderately large, rather well-spaced, black pitting with background of dense, very small black pitting in
<taxonomicName id="4C114D6FFFA9FFF02356F9C05C9B11F1" authorityName="Constant" authorityYear="2024" box="[619,753,1645,1671]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="chartieri">
<emphasis id="B965EAFEFFA9FFF02356F9C05C9B11F1" box="[619,753,1645,1671]" italics="true" pageId="19" pageNumber="20">Z. chartieri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFA9FFF02217F9C05DD611F1" authorityName="Constant" authorityYear="2024" box="[810,956,1645,1671]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFA9FFF02217F9C05DD611F1" box="[810,956,1645,1671]" italics="true" pageId="19" pageNumber="20">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA9FFF022E4F9C35A1F11FE" box="[985,1141,1645,1672]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="19" pageNumber="20">Figs 7B, EF</figureCitation>
,
<figureCitation id="132A2A69FFA9FFF025BFF9C35A9D11F1" box="[1154,1271,1645,1671]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="19" pageNumber="20">16B, EF</figureCitation>
); anal tube of male (
<figureCitation id="132A2A69FFA9FFF02014F93E5FFF11DD" box="[297,405,1680,1707]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="19" pageNumber="20">Fig. 13A</figureCitation>
) rather flattened dorsoventrally, 1.95 times as long as high in lateral view and with ventral angle at midlength (rather high in lateral view, 1.601.65 times as long as high, with ventral angle in basal half in
<taxonomicName id="4C114D6FFFA9FFF020FDF9765C371187" authorityName="Constant &amp; Pham" authorityYear="2024" box="[448,605,1751,1777]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="bidoupana">
<emphasis id="B965EAFEFFA9FFF020FDF9765C371187" box="[448,605,1751,1777]" italics="true" pageId="19" pageNumber="20">Z. bidoupana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFA9FFF023A4F9765D461187" authorityName="Constant" authorityYear="2024" box="[665,812,1751,1777]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFA9FFF023A4F9765D461187" box="[665,812,1751,1777]" italics="true" pageId="19" pageNumber="20">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA9FFF02277F9795DC11187" box="[842,939,1751,1777]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="19" pageNumber="20">Figs 3A</figureCitation>
,
<figureCitation id="132A2A69FFA9FFF02284F9795D851187" box="[953,1007,1751,1777]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="19" pageNumber="20">16A</figureCitation>
); pygofer of male in lateral view (
<figureCitation id="132A2A69FFA9FFF021FBF9545F5B1062" box="[198,305,1786,1812]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="19" pageNumber="20">Fig. 13A</figureCitation>
) with posterior margin projecting into a large posterior lobe angularly rounded in midheight (posterior margin sinuate, forming a large, broadly rounded posterior lobe in dorsal half in
<taxonomicName id="4C114D6FFFA9FFF025CCF8B05B1D1041" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1265,1399,1821,1847]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFA9FFF025CCF8B05B1D1041" box="[1265,1399,1821,1847]" italics="true" pageId="19" pageNumber="20">Z. kusamae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFA9FFF021CFF8EF5F11102C" authorityName="Constant" authorityYear="2024" box="[242,379,1856,1882]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="chartieri">
<emphasis id="B965EAFEFFA9FFF021CFF8EF5F11102C" box="[242,379,1856,1882]" italics="true" pageId="19" pageNumber="20">Z. chartieri</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA9FFF020A2F8EE5C68102D" box="[415,514,1856,1883]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="19" pageNumber="20">Figs 3A</figureCitation>
,
<figureCitation id="132A2A69FFA9FFF0232FF8EF5C53102D" box="[530,569,1857,1883]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="19" pageNumber="20">7A</figureCitation>
; posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle in
<taxonomicName id="4C114D6FFFA9FFF02385F8CA5D20100B" authorityName="Constant" authorityYear="2024" box="[696,842,1891,1917]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFA9FFF02385F8CA5D20100B" box="[696,842,1891,1917]" italics="true" pageId="19" pageNumber="20">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFA9FFF0225AF8CD5DB9100B" box="[871,979,1891,1917]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="19" pageNumber="20">Fig. 16A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFAEFFF72180FEA75F0616E1" pageId="20" pageNumber="21" type="etymology">
<paragraph id="8BAE36ECFFAEFFF72180FEA75F2E1655" blockId="20.[189,324,265,291]" box="[189,324,265,291]" pageId="20" pageNumber="21">
<heading id="D0E68180FFAEFFF72180FEA75F2E1655" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B965EAFEFFAEFFF72180FEA75F2E1655" bold="true" box="[189,324,265,291]" pageId="20" pageNumber="21">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180FE995F0616E1" blockId="20.[189,1399,311,407]" pageId="20" pageNumber="21">
The species epithet is a patronym dedicated to the famous Japanese artist Mrs Yayoi Kusama (1929), whose artwork, extensively covered in dots, is reminiscent of the pattern on the body and tegmina of
<taxonomicName id="4C114D6FFFAEFFF72180FED05F0616E1" box="[189,364,381,407]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B965EAFEFFAEFFF72180FED05F6216E1" box="[189,264,382,407]" italics="true" pageId="20" pageNumber="21">Zanna</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFAEFFF72180FE115FAF15E2" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BAE36ECFFAEFFF72180FE115F0616AF" blockId="20.[189,364,447,473]" box="[189,364,447,473]" pageId="20" pageNumber="21">
<heading id="D0E68180FFAEFFF72180FE115F0616AF" bold="true" box="[189,364,447,473]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B965EAFEFFAEFFF72180FE115F0616AF" bold="true" box="[189,364,447,473]" pageId="20" pageNumber="21">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFAEFFF721D7FE405F36157E" blockId="20.[189,1398,494,660]" box="[234,348,494,520]" pageId="20" pageNumber="21">
<emphasis id="B965EAFEFFAEFFF721D7FE405F36157E" bold="true" box="[234,348,494,520]" pageId="20" pageNumber="21">
<typeStatus id="54AA884EFFAEFFF721D7FE405F36157E" box="[234,348,494,520]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180FDBF5FAF15E2" blockId="20.[189,1398,494,660]" pageId="20" pageNumber="21">
<materialsCitation id="3B793CB1FFAEFFF72180FDBF5FAF15E2" collectingDate="2012-06-25" collectingDateMax="2012-07-06" collectingDateMin="2012-06-25" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc. &amp; Leg. J. Constant" country="Vietnam" latitude="11.3" location="Biosphere Reserve" longLatPrecision="1295" longitude="107.1" municipality="Vietnam" pageId="20" pageNumber="21" specimenCount="1" specimenCount-male="1" stateProvince="Dong Nai" typeStatus="holotype">
<collectingCountry id="F306767CFFAEFFF72180FDBF5F2E155D" box="[189,324,529,555]" name="Vietnam" pageId="20" pageNumber="21">VIETNAM</collectingCountry>
<specimenCount id="9D17FD65FFAEFFF72066FDBF5FE3155D" box="[347,393,529,555]" count="1" pageId="20" pageNumber="21" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BCAAC96FFAEFFF720AEFDBF5F93155D" box="[403,505,529,555]" pageId="20" pageNumber="21">Vietnam</collectingMunicipality>
,
<collectingRegion id="49D5F80EFFAEFFF72339FDBF5C1F155D" box="[516,629,529,555]" country="Vietnam" name="Dong Nai" pageId="20" pageNumber="21">Dong Nai</collectingRegion>
<location id="8ECE6037FFAEFFF72341FDBF5D31155D" LSID="urn:lsid:plazi:treatment:03B887FAFFA9FFFB23BAFAAA5C131362:8ECE6037FFAEFFF72341FDBF5D31155D" box="[636,859,529,555]" country="Vietnam" latitude="11.3" longLatPrecision="1295" longitude="107.1" municipality="Vietnam" name="Biosphere Reserve" pageId="20" pageNumber="21" stateProvince="Dong Nai">Biosphere Reserve</location>
;
<geoCoordinate id="EE25502BFFAEFFF7225BFDA15DBA155D" box="[870,976,527,555]" degrees="11" direction="north" minutes="18" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="11.3">11°18ʹ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFAEFFF722E7FDA15A3B155D" box="[986,1105,527,555]" degrees="107" direction="east" minutes="06" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="107.1">107°06ʹ E</geoCoordinate>
;
<date id="FFAF102CFFAEFFF72561FDBE5B2B155D" box="[1116,1345,528,555]" pageId="20" pageNumber="21" value="2012-06-25" valueMax="2012-07-06" valueMin="2012-06-25">
<collectingDate id="EFEBE9C4FFAEFFF72561FDBE5B2B155D" box="[1116,1345,528,555]" pageId="20" pageNumber="21" value="2012-06-25" valueMax="2012-07-06" valueMin="2012-06-25">25 Jun.6 Jul. 2012</collectingDate>
</date>
; day time collecting;
<collectorName id="26E4533AFFAEFFF72042FD9A5C691538" box="[383,515,564,590]" pageId="20" pageNumber="21">J. Constant</collectorName>
and
<collectorName id="26E4533AFFAEFFF7237DFD9A5CD51538" box="[576,703,564,590]" pageId="20" pageNumber="21">J. Bresseel</collectorName>
leg.; I.G.: 32.161; “
<collectorName id="26E4533AFFAEFFF7228CFD9A5AEA1538" box="[945,1152,564,590]" pageId="20" pageNumber="21">Coll. I.R.Sc.N.B.</collectorName>
, Vietnam,
<collectingRegion id="49D5F80EFFAEFFF7243FFD9A5B1C1538" box="[1282,1398,564,590]" country="Vietnam" name="Dong Nai" pageId="20" pageNumber="21">Dong Nai</collectingRegion>
Biosphere Res.,
<geoCoordinate id="EE25502BFFAEFFF72043FDFB5F831504" box="[382,489,597,626]" degrees="11" direction="north" minutes="18" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="11.3">11°18ʹ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFAEFFF720CBFDFB5C071504" box="[502,621,597,626]" degrees="107" direction="east" minutes="06" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="107.1">107°06ʹ E</geoCoordinate>
,
<date id="FFAF102CFFAEFFF72347FDF95D211504" box="[634,843,599,626]" pageId="20" pageNumber="21" value="2012-06-25" valueMax="2012-07-06" valueMin="2012-06-25">
<collectingDate id="EFEBE9C4FFAEFFF72347FDF95D211504" box="[634,843,599,626]" pageId="20" pageNumber="21" value="2012-06-25" valueMax="2012-07-06" valueMin="2012-06-25">25.VI-6.VII.2012</collectingDate>
</date>
, day collecting,
<collectorName id="26E4533AFFAEFFF72531FDF65AA11507" box="[1036,1227,599,626]" pageId="20" pageNumber="21">Leg. J. Constant</collectorName>
&amp;
<collectorName id="26E4533AFFAEFFF725CCFDF65B181507" box="[1265,1394,599,626]" pageId="20" pageNumber="21">J. Bresseel</collectorName>
, I.G.: 32.161”;
<collectionCode id="ED00AE29FFAEFFF7205AFDD45FAB15E2" box="[359,449,634,660]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="20" pageNumber="21" type="Museum">RBINS</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFAEFFFB2180FD155B031466" lastPageId="24" lastPageNumber="25" pageId="20" pageNumber="21" type="description">
<paragraph id="8BAE36ECFFAEFFF72180FD155F2415A3" blockId="20.[189,334,699,725]" box="[189,334,699,725]" pageId="20" pageNumber="21">
<heading id="D0E68180FFAEFFF72180FD155F2415A3" bold="true" box="[189,334,699,725]" fontSize="11" level="3" pageId="20" pageNumber="21" reason="3">
<emphasis id="B965EAFEFFAEFFF72180FD155F2415A3" bold="true" box="[189,334,699,725]" pageId="20" pageNumber="21">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180FD445DD2145E" blockId="20.[189,1398,746,808]" pageId="20" pageNumber="21">
MEASUREMENTS AND RATIOS. LT:
<specimenCount id="9D17FD65FFAEFFF7237BFD455C361473" box="[582,604,747,773]" count="1" pageId="20" pageNumber="21" type="male"></specimenCount>
(n =1): 38.0 mm. LTg/BTg= 3.25; LH/BH=4.2; LF/BF=5.9; BH/ BPrH= 2.4; LH/LT= 0.39; wingspan (extrapolated):
<specimenCount id="9D17FD65FFAEFFF7221FFCA05D56145E" box="[802,828,782,808]" count="1" pageId="20" pageNumber="21" type="male"></specimenCount>
:
<quantity id="4CE99B09FFAEFFF72275FCA35DD9145E" box="[840,947,781,808]" metricMagnitude="-2" metricUnit="m" metricValue="4.52" pageId="20" pageNumber="21" unit="mm" value="45.2">45.2 mm</quantity>
.
</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180FCE15CED13F5" blockId="20.[189,1399,847,1155]" pageId="20" pageNumber="21">
HEAD (
<figureCitation id="132A2A69FFAEFFF72033FCE15FD3141F" box="[270,441,847,873]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="20" pageNumber="21">Fig. 12B, EF</figureCitation>
). Strongly elongate, representing 39% of total length, gently and evenly narrowing towards apex with apical ⅓ more or less parallel-sided in dorsal view; apex weakly widening, obliquely cut in lateral view and with incomplete, rim-shaped orange carina (missing in ventral portion); pale pinkish brown with irregular, moderately large, rather well-spaced, black pitting, more dense in basal half, especially on sides, and with very minute pits towards distal portion; some larger black pustules on side, along carina of vertex. Lateral carinae of frons weakly zigzagged, distinct towards apex, upcurved near apex and merging into a short dorsal carina reaching apex; lateral carinae of vertex distinct towards apex, merging with upcurved part of lateral carinae of frons; median carina of vertex obsolete. Labium yellowish brown, reaching mesocoxae.
</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180FB045B611391" blockId="20.[189,1399,1194,1256]" pageId="20" pageNumber="21">
THORAX (
<figureCitation id="132A2A69FFAEFFF7200DFB045F8A13B2" box="[304,480,1194,1220]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="20" pageNumber="21">Fig. 12B, EF</figureCitation>
). Pro- and mesonotum pale pinkish brown irregularly covered in small black pitting with pits much smaller than space between them (on average); median carina obsolete.
</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180FAA15AE512E5" blockId="20.[189,1398,1294,1427]" pageId="20" pageNumber="21">
TEGMINA (
<figureCitation id="132A2A69FFAEFFF7200BFAA05F8F125F" box="[310,485,1294,1321]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="20" pageNumber="21">Fig. 12A, CD</figureCitation>
). Strongly elongate and reticulate, pale (subhyaline) pinkish brown, moderately densely covered in minute to small black spots, mostly on veins, 3 slightly larger ones; costal and postclaval margins subparallel, weakly diverging from base towards apex and curved at level of apex of clavus; apical margin rounded. Veins yellowish in large basal portion. Clavus open.
</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180FA145B221281" blockId="20.[189,1398,1464,1527]" pageId="20" pageNumber="21">VENATION. ScP+R forked in basal ⅓, MP forked more basally in basal 1/4; CuA forked near apex of clavus; PCu and A1 fused near apex of clavus; PCu+A1 fused with postclaval margin at nodal line.</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180F9B15AB5112D" blockId="20.[189,1399,1566,1628]" pageId="20" pageNumber="21">
HIND WINGS (
<figureCitation id="132A2A69FFAEFFF72062F9B05F87114E" box="[351,493,1566,1592]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="20" pageNumber="21">Fig. 12A, C</figureCitation>
). Subhyaline, milky white with veins slightly darker, vein PCu dark brown in basal half; postclaval margin weakly emarginate at A1 vein, and infuscate along vein CuP.
</paragraph>
<paragraph id="8BAE36ECFFAEFFF72180F92D5FCB1070" blockId="20.[189,1399,1665,1798]" pageId="20" pageNumber="21">
VENATION. ScP+R and MP forked at distal ⅓; CuA forked slightly more basally then MP fork; PCu forked at basal ⅓; A1 forked close to base, A1
<subScript id="179534A9FFAEFFF723DEF9175C8611B1" attach="left" box="[739,748,1721,1735]" fontSize="6" pageId="20" pageNumber="21">2</subScript>
fused with A2 rather close to base and A1
<subScript id="179534A9FFAEFFF725D8F9175A8411B1" attach="right" box="[1253,1262,1721,1735]" fontSize="6" pageId="20" pageNumber="21">2</subScript>
+A2 forked slightly before reaching postclaval margin. Numerous crossveins in large portion along apical and postclaval margins.
</paragraph>
<paragraph id="8BAE36ECFFAEFFF42180F8805B1D161F" blockId="20.[189,1398,1838,2005]" lastBlockId="23.[189,1399,265,362]" lastPageId="23" lastPageNumber="24" pageId="20" pageNumber="21">
LEGS (
<figureCitation id="132A2A69FFAEFFF72037F8805FA8103E" box="[266,450,1838,1864]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="20" pageNumber="21">Fig. 12A, CD</figureCitation>
). Pale pinkish stramineous, pilose, rather short and robust. Pro- and mesocoxae with few small black markings; all femora with few small to minute black spots, sometimes absent, and conspicuous black spots aligned in a ring in distal ⅓, sometimes coalescent; pro- and mesotibiae with dark brown spots in basal ¼, and with apex brownish; pro- and mesotarsi with segments blackish brown apically; metatibiae weakly brown basally with all spines apically blackish brown, 45 lateral spines and 8 apical spines; first metatarsomere with 1113 apical spines ventrally; second metatarsomere with 9 apical spines ventrally; first and second metatarsomere with a dense pad of microsetae ventrally; third metatarsomere elongate with a subapical blackish brown ring. Metatibiotarsal formula: (45) 8/1113/9.
</paragraph>
<caption id="DF6E6664FFAFFFF62181F8295FCC1091" ID-DOI="http://doi.org/10.5281/zenodo.13837062" ID-Zenodo-Dep="13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="21" pageNumber="22" startId="21.[188,229,1927,1953]" targetBox="[215,1367,257,1885]" targetPageId="21">
<paragraph id="8BAE36ECFFAFFFF62181F8295FCC1091" blockId="21.[188,1397,1927,2023]" pageId="21" pageNumber="22">
<emphasis id="B965EAFEFFAFFFF62181F8295F7110D7" bold="true" box="[188,283,1927,1953]" pageId="21" pageNumber="22">Fig. 12.</emphasis>
<taxonomicName id="4C114D6FFFAFFFF62019F8265F8B10D7" authorityName="Constant &amp; Pham" authorityYear="2024" box="[292,481,1927,1953]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFAFFFF62019F8265F8B10D7" box="[292,481,1927,1953]" italics="true" pageId="21" pageNumber="22">Zanna kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFAFFFF620D5F8265C2C10D4" box="[488,582,1928,1954]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS).
<emphasis id="B965EAFEFFAFFFF62259F8295D1310D7" bold="true" box="[868,889,1927,1953]" pageId="21" pageNumber="22">A</emphasis>
. Habitus dorsal view.
<emphasis id="B965EAFEFFAFFFF625BBF8295AF010D7" bold="true" box="[1158,1178,1927,1953]" pageId="21" pageNumber="22">B</emphasis>
. Head and thorax, dorsal view.
<emphasis id="B965EAFEFFAFFFF62065F8045F0710B2" bold="true" box="[344,365,1962,1988]" pageId="21" pageNumber="22">C</emphasis>
. Habitus, ventral view.
<emphasis id="B965EAFEFFAFFFF623AFF8045CCD10B2" bold="true" box="[658,679,1962,1988]" pageId="21" pageNumber="22">D</emphasis>
. Habitus, lateral view.
<emphasis id="B965EAFEFFAFFFF622F9F8045DB210B2" bold="true" box="[964,984,1962,1988]" pageId="21" pageNumber="22">E</emphasis>
. Head, ventral view.
<emphasis id="B965EAFEFFAFFFF625E2F8045A9B10B2" bold="true" box="[1247,1265,1962,1988]" pageId="21" pageNumber="22">F</emphasis>
. Head and thorax, lateral view.
</paragraph>
</caption>
<caption id="DF6E6664FFACFFF52180F8285B1D1090" ID-DOI="http://doi.org/10.5281/zenodo.13837066" ID-Zenodo-Dep="13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="22" pageNumber="23" startId="22.[189,230,1926,1952]" targetBox="[218,1365,253,1887]" targetPageId="22">
<paragraph id="8BAE36ECFFACFFF52180F8285B1D1090" blockId="22.[186,1399,1926,2023]" pageId="22" pageNumber="23">
<emphasis id="B965EAFEFFACFFF52180F8285F7C10D6" bold="true" box="[189,278,1926,1952]" pageId="22" pageNumber="23">Fig. 13.</emphasis>
<emphasis id="B965EAFEFFACFFF52027F8285FA710D6" box="[282,461,1926,1952]" italics="true" pageId="22" pageNumber="23">Zannakusamae</emphasis>
<taxonomicNameLabel id="A2565785FFACFFF520E9F8295C4410D7" box="[468,558,1927,1953]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS), terminalia.
<emphasis id="B965EAFEFFACFFF52281F8285D9F10D6" bold="true" box="[956,1013,1926,1952]" pageId="22" pageNumber="23">AD</emphasis>
. Pygofer, anal tube and gonostyli.
<emphasis id="B965EAFEFFACFFF52187F8075EA510B5" bold="true" box="[186,207,1961,1987]" pageId="22" pageNumber="23">A</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFACFFF52089F8075FA210B5" bold="true" box="[436,456,1961,1987]" pageId="22" pageNumber="23">B</emphasis>
. Caudal view.
<emphasis id="B965EAFEFFACFFF52347F8075CE510B5" bold="true" box="[634,655,1961,1987]" pageId="22" pageNumber="23">C</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFACFFF52200F8075D3810B5" bold="true" box="[829,850,1961,1987]" pageId="22" pageNumber="23">D</emphasis>
. Ventral view.
<emphasis id="B965EAFEFFACFFF5253BF8075A5510B5" bold="true" box="[1030,1087,1961,1987]" pageId="22" pageNumber="23">EH</emphasis>
. Aedeagus.
<emphasis id="B965EAFEFFACFFF525EDF8075A8E10B5" bold="true" box="[1232,1252,1961,1987]" pageId="22" pageNumber="23">E</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFACFFF5203FF8635F7E1091" bold="true" box="[258,276,1997,2023]" pageId="22" pageNumber="23">F</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFACFFF52081F8625FB91090" bold="true" box="[444,467,1996,2022]" pageId="22" pageNumber="23">G</emphasis>
. Laterodorsal view.
<emphasis id="B965EAFEFFACFFF523FCF8635CB21091" bold="true" box="[705,728,1997,2023]" pageId="22" pageNumber="23">H</emphasis>
. Ventral view. Abbreviations: see Materials and methods.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFADFFF42180FE3F5A8F16B9" blockId="23.[189,1398,401,463]" pageId="23" pageNumber="24">
ABDOMEN (
<figureCitation id="132A2A69FFADFFF4207EFE3F5FB916DD" box="[323,467,401,427]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="23" pageNumber="24">Fig. 12A, C</figureCitation>
). Tergites smooth and shiny, yellowish brown; sternites (except pregenital one) yellowish brown with irregular small black dots more or less aligned into a transverse row.
</paragraph>
<caption id="DF6E6664FFADFFF42180F8305A6710AD" ID-DOI="http://doi.org/10.5281/zenodo.13837070" ID-Zenodo-Dep="13837070" httpUri="https://zenodo.org/record/13837070/files/figure.png" pageId="23" pageNumber="24" startId="23.[189,230,1950,1976]" targetBox="[187,1398,522,1886]" targetPageId="23">
<paragraph id="8BAE36ECFFADFFF42180F8305A6710AD" blockId="23.[189,1398,1950,2012]" pageId="23" pageNumber="24">
<emphasis id="B965EAFEFFADFFF42180F8305F7510CE" bold="true" box="[189,287,1950,1976]" pageId="23" pageNumber="24">Fig. 14.</emphasis>
<taxonomicName id="4C114D6FFFADFFF42011F8315F8110CE" authorityName="Constant &amp; Pham" authorityYear="2024" box="[300,491,1950,1976]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFADFFF42011F8315F8110CE" box="[300,491,1950,1976]" italics="true" pageId="23" pageNumber="24">Zanna kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFADFFF420C5F8305C3010CE" box="[504,602,1950,1976]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂, in nature, Dong Nai Biosphere Reserve, Vietnam, 3 Jul. 2012.
<emphasis id="B965EAFEFFADFFF42070F86F5F0810AD" bold="true" box="[333,354,1985,2011]" pageId="23" pageNumber="24">A</emphasis>
. Habitus lateral view.
<emphasis id="B965EAFEFFADFFF42357F86F5C1410AD" bold="true" box="[618,638,1985,2011]" pageId="23" pageNumber="24">B</emphasis>
. Habitus, dorsal view.
<emphasis id="B965EAFEFFADFFF422B6F86F5DCA10AD" bold="true" box="[907,928,1985,2011]" pageId="23" pageNumber="24">C</emphasis>
. Habitat.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFA2FFFB2180FEA75B031466" blockId="24.[189,1401,265,784]" pageId="24" pageNumber="25">
MALE TERMINALIA (
<figureCitation id="132A2A69FFA2FFFB2094FEA75C691655" box="[425,515,265,291]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">Fig. 13</figureCitation>
). Pygofer (
<emphasis id="B965EAFEFFA2FFFB23B0FEA45CC61655" box="[653,684,266,291]" italics="true" pageId="24" pageNumber="25">Py</emphasis>
<figureCitation id="132A2A69FFA2FFFB23EDFEA75D0B1655" box="[720,865,265,291]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">Fig. 13AD</figureCitation>
) about 1.51 times as high as long in lateral view, robust with posterior margin projecting posteriorly into a large posterior lobe in midheight; lobe angularly rounded; sides weakly excavate in dorsal portion (visible in dorsal aspect,
<figureCitation id="132A2A69FFA2FFFB2586FEE15B4D161F" box="[1211,1319,335,361]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">Fig. 13C</figureCitation>
). Anal tube (
<emphasis id="B965EAFEFFA2FFFB203CFEDD5F4816FB" box="[257,290,371,397]" italics="true" pageId="24" pageNumber="25">An</emphasis>
<figureCitation id="132A2A69FFA2FFFB207DFEDC5FD416FB" box="[320,446,370,397]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="24" pageNumber="25">Fig. 3AC</figureCitation>
) robust, 1.95 times as long as high and with ventral margin oblique in distal half and angularly projecting at a wide angle in lateral view; 1.26 times as long as wide in dorsal view, with lateral margins slightly diverging towards apex and apical margin bisinuate, weakly excavate in middle portion; suboval, more or less flattened dorsally, in caudal view; epiproct and paraproct strongly surpassing anal tube, angularly rounded posteriorly in dorsal aspect. Gonostyli (
<emphasis id="B965EAFEFFA2FFFB25A3FDAE5AD9156C" box="[1182,1203,512,538]" italics="true" pageId="24" pageNumber="25">G</emphasis>
<figureCitation id="132A2A69FFA2FFFB25E6FE515B04156F" box="[1243,1390,511,537]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">Fig. 13AD</figureCitation>
) elongate, 2.82 times as long as high; in lateral view, ventral margin broadly rounded and dorsal margin rather broadly rounded after lateral hook (
<emphasis id="B965EAFEFFA2FFFB238FFDE85CBD1516" box="[690,727,582,608]" italics="true" pageId="24" pageNumber="25">lhg</emphasis>
), apical margin rounded; lateral hook with basal portion projecting posterolaterad and dorsal hump in middle portion, and apex strongly curved lateroventrad; in ventral view, lateral protrusion at about midlength, rather strongly and angularly marked. Aedeagus (
<figureCitation id="132A2A69FFA2FFFB21FBFD015F3815BF" box="[198,338,687,713]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="24" pageNumber="25">Fig. 13EH</figureCitation>
) rather elongate with lateral processes (
<emphasis id="B965EAFEFFA2FFFB2213FD015D3915BF" box="[814,851,687,713]" italics="true" pageId="24" pageNumber="25">lpa</emphasis>
) forming multiple lobes; dorsal process (
<emphasis id="B965EAFEFFA2FFFB247CFD015B0715BF" box="[1345,1389,687,713]" italics="true" pageId="24" pageNumber="25">dpa</emphasis>
) strongly protruding dorsally in lateral view; dorsal endosomal process (
<emphasis id="B965EAFEFFA2FFFB22C5FD7C5A48159A" box="[1016,1058,722,748]" italics="true" pageId="24" pageNumber="25">dep</emphasis>
) in dorsal view subtriangular with lateral margins very weakly curved (nearly straight) and rather wide median slit in distal portion.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA2FFFB2180FC995D7C14D5" pageId="24" pageNumber="25" type="biology_ecology">
<paragraph id="8BAE36ECFFA2FFFB2180FC995F761427" blockId="24.[189,284,823,849]" box="[189,284,823,849]" pageId="24" pageNumber="25">
<heading id="D0E68180FFA2FFFB2180FC995F761427" bold="true" box="[189,284,823,849]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B965EAFEFFA2FFFB2180FC995F761427" bold="true" box="[189,284,823,849]" pageId="24" pageNumber="25">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA2FFFB2180FCC85D7C14D5" blockId="24.[189,1399,870,931]" pageId="24" pageNumber="25">
The
<typeStatus id="54AA884EFFA2FFFB21CEFCC85F3014F6" box="[243,346,870,896]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
(
<figureCitation id="132A2A69FFA2FFFB2057FCC85F9214F6" box="[362,504,870,896]" captionStart="Fig" captionStartId="23.[189,230,1950,1976]" captionTargetBox="[187,1398,522,1886]" captionTargetId="figure-106@23.[187,1401,522,1886]" captionTargetPageId="23" captionText="Fig. 14. Zanna kusamae sp. nov., holotype, ♂, in nature, Dong Nai Biosphere Reserve, Vietnam, 3 Jul. 2012. A. Habitus lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi="http://doi.org/10.5281/zenodo.13837070" httpUri="https://zenodo.org/record/13837070/files/figure.png" pageId="24" pageNumber="25">Fig. 14AB</figureCitation>
) was collected sitting on a leaf of a short plant along a gravel road near the clearing around the station in
<collectingRegion id="49D5F80EFFA2FFFB2327FC275CE714D5" box="[538,653,905,931]" country="Vietnam" name="Dong Nai" pageId="24" pageNumber="25">Dong Nai</collectingRegion>
(
<figureCitation id="132A2A69FFA2FFFB23A0FC275D6314D5" box="[669,777,905,931]" captionStart="Fig" captionStartId="23.[189,230,1950,1976]" captionTargetBox="[187,1398,522,1886]" captionTargetId="figure-106@23.[187,1401,522,1886]" captionTargetPageId="23" captionText="Fig. 14. Zanna kusamae sp. nov., holotype, ♂, in nature, Dong Nai Biosphere Reserve, Vietnam, 3 Jul. 2012. A. Habitus lateral view. B. Habitus, dorsal view. C. Habitat." figureDoi="http://doi.org/10.5281/zenodo.13837070" httpUri="https://zenodo.org/record/13837070/files/figure.png" pageId="24" pageNumber="25">Fig. 14C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA2FFFB2180FC645C131362" box="[189,633,970,1044]" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BAE36ECFFA2FFFB2180FC645F3C1492" blockId="24.[189,342,970,996]" box="[189,342,970,996]" pageId="24" pageNumber="25">
<heading id="D0E68180FFA2FFFB2180FC645F3C1492" bold="true" box="[189,342,970,996]" fontSize="11" level="3" pageId="24" pageNumber="25" reason="3">
<emphasis id="B965EAFEFFA2FFFB2180FC645F3C1492" bold="true" box="[189,342,970,996]" pageId="24" pageNumber="25">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA2FFFB2180FC575C131362" blockId="24.[189,633,1017,1044]" box="[189,633,1017,1044]" pageId="24" pageNumber="25">
<collectingCountry id="F306767CFFA2FFFB2180FC575F4F1365" box="[189,293,1017,1043]" name="Vietnam" pageId="24" pageNumber="25">Vietnam</collectingCountry>
:
<collectingRegion id="49D5F80EFFA2FFFB200DFC575C781365" box="[304,530,1017,1043]" country="Vietnam" name="Dong Nai" pageId="24" pageNumber="25">Dong Nai Province</collectingRegion>
(
<figureCitation id="132A2A69FFA2FFFB231FFC575C001362" box="[546,618,1017,1044]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="24" pageNumber="25">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,641 @@
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<mods:title id="9C3ECE85DBC6E6C2A8C519CF74A7010A">Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<mods:namePart id="9316F2E34590A4C141820D3DFC144267">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="C69500E0CB5B3E7BE2AF60E0CB126BB0">Royal Belgian Institute of Natural Sciences, O. D. Phylogeny and Taxonomy, Entomology, Vautier Street 29, B- 1000 Brussels, Belgium.</mods:affiliation>
<mods:nameIdentifier id="583E1FC026B97D9038F8A01863F262D9" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:namePart id="E867CB1A287B8A0170C5AAF714EDA969">Pham, Hong Thai</mods:namePart>
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<mods:affiliation id="FC278A2FC96DD3ECC8DCCEB97FED6902">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.</mods:affiliation>
<mods:nameIdentifier id="FC44D2BC3908B688C718D99C5981B32D" type="email">jerome.constant@naturalsciences.be</mods:nameIdentifier>
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<mods:title id="600B8B70FF1CCBECF8B41309B7E7B018">European Journal of Taxonomy</mods:title>
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<mods:part id="04939B98919EAFE4D26CC56D151BF919">
<mods:date id="3F8D182C8DE826C20234FF1F94FFF5CC">2024</mods:date>
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<mods:number id="90A8FF2954271B84AD189EDB1017C880">2024-09-25</mods:number>
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<mods:number id="F1F55441F7DFC71D6CF1B704EEAADDDC">958</mods:number>
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<mods:url id="1F8BB565159565D25BCB6810CB1B21B9">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2665/12349</mods:url>
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<mods:identifier id="1663D50D9C47F0FB4230B436057578C8" type="DOI">10.5852/ejt.2024.958.2665</mods:identifier>
<mods:identifier id="08273706BE142D7247C7AD1F58E4EB6A" type="ISSN">2118-9773</mods:identifier>
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<subSubSection id="C30B6567FFB0FFE9236CFB105AD6138D" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BAE36ECFFB0FFE9236CFB105D8813AE" blockId="10.[376,1212,1213,1314]" box="[593,994,1213,1240]" pageId="10" pageNumber="11">
<heading id="D0E68180FFB0FFE9236CFB105D8813AE" box="[593,994,1213,1240]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="4C114D6FFFB0FFE9236CFB105DEA13A1" authority="Constant" authorityName="Constant" authorityYear="2024" box="[593,896,1213,1240]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFB0FFE9236CFB105D7B13AE" bold="true" box="[593,785,1213,1240]" italics="true" pageId="10" pageNumber="11">Zanna chartieri</emphasis>
Constant
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<taxonomicNameLabel id="A2565785FFB0FFE922B5FB105D8813AE" box="[904,994,1214,1240]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C30B6567FFB0FFE9234EFAA95DAB1257" box="[627,961,1287,1314]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BAE36ECFFB0FFE9234EFAA95DAB1257" blockId="10.[376,1212,1213,1314]" box="[627,961,1287,1314]" pageId="10" pageNumber="11">
<figureCitation id="132A2A69FFB0FFE9234EFAA95C831254" box="[627,745,1287,1314]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStartId-0="9.[188,229,1994,2020]" captionStartId-1="12.[185,226,1930,1956]" captionStartId-2="13.[189,228,1925,1951]" captionStartId-3="14.[190,231,1950,1976]" captionStartId-4="15.[189,230,1896,1922]" captionStartId-5="16.[189,230,1894,1920]" captionStartId-6="18.[189,230,1926,1952]" captionTargetBox-0="[215,1372,718,1954]" captionTargetBox-1="[252,1334,322,1889]" captionTargetBox-2="[232,1368,257,1882]" captionTargetBox-3="[201,1399,285,1873]" captionTargetBox-4="[213,1370,268,1848]" captionTargetBox-5="[238,1351,274,1856]" captionTargetBox-6="[212,1369,538,1888]" captionTargetId-0="figure-180@9.[215,1372,718,1954]" captionTargetId-1="figure-47@12.[242,1334,317,1889]" captionTargetId-2="figure-62@13.[220,1377,238,1886]" captionTargetId-3="figure-25@14.[189,1399,275,1884]" captionTargetId-4="figure-62@15.[213,1370,268,1851]" captionTargetId-5="figure-84@16.[238,1351,274,1856]" captionTargetId-6="figure-156@18.[212,1369,538,1888]" captionTargetPageId-0="9" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="15" captionTargetPageId-5="16" captionTargetPageId-6="18" captionText-0="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." captionText-1="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." captionText-2="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." captionText-3="Fig. 8. Zanna chartieri Constant sp. nov., nymph. A. Habitus dorsal view. B. Habitus, ventral view. C. Habitus, lateral view." captionText-4="Fig. 9. Zanna chartieri Constant sp. nov., in nature in Tatai, Cambodia (photographs © G. Chartier). A. Adult on Desmodium sp., 20 Jan. 2017. B. Adult on unidentified plant, 17 Jan. 2017. C. Adult on Melastomasanguineum, lateral view, 17 Jan. 2017. D. Habitat, position of nymphs marked with red arrows, 20 Jan. 2017." captionText-5="Fig. 10. Zanna chartieri Constant sp. nov., nymphs in nature in Tatai, Cambodia, on Desmodium sp. (photographs © G. Chartier). AB. 4th instar nymphs. A. Laterodorsal view, 18 Jan. 2017. B. Lateral view, 20 Jan. 2017. CD. 5th instar nymphs, 17 Jan. 2017. C. Lateral view. D. Dorsal view. EF. Groups of nymphs. E. On vertical twig, 20 Jan. 2017. F. On creeping twig close to ground, 19 Jan. 2019." captionText-6="Fig. 11. Zannachartieri Constant sp. nov., emerging adult in nature in Tatai, Cambodia, onDesmodium sp. (photographs © G. Chartier). A. Emerging adult with 4th and 5th instar nymphs, 18 Jan. 2017. B. 18 Jan. 2017. CH. Time lapse of final moult of the same specimen, 18 Jan. 2017." figureDoi-0="http://doi.org/10.5281/zenodo.13837040" figureDoi-1="http://doi.org/10.5281/zenodo.13837042" figureDoi-2="http://doi.org/10.5281/zenodo.13837046" figureDoi-3="http://doi.org/10.5281/zenodo.13837048" figureDoi-4="http://doi.org/10.5281/zenodo.13837052" figureDoi-5="http://doi.org/10.5281/zenodo.13837056" figureDoi-6="http://doi.org/10.5281/zenodo.13837060" httpUri-0="https://zenodo.org/record/13837040/files/figure.png" httpUri-1="https://zenodo.org/record/13837042/files/figure.png" httpUri-2="https://zenodo.org/record/13837046/files/figure.png" httpUri-3="https://zenodo.org/record/13837048/files/figure.png" httpUri-4="https://zenodo.org/record/13837052/files/figure.png" httpUri-5="https://zenodo.org/record/13837056/files/figure.png" httpUri-6="https://zenodo.org/record/13837060/files/figure.png" pageId="10" pageNumber="11">Figs 511</figureCitation>
,
<figureCitation id="132A2A69FFB0FFE923C8FAA95D351257" box="[757,863,1287,1313]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="10" pageNumber="11">18B, F, J</figureCitation>
,
<figureCitation id="132A2A69FFB0FFE92251FAA95DAB1257" box="[876,961,1287,1313]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="10" pageNumber="11">19CD</figureCitation>
</paragraph>
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<subSubSection id="C30B6567FFB0FFE92180FAE65D5A1062" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BAE36ECFFB0FFE92180FAE65F5C1214" blockId="10.[189,310,1352,1378]" box="[189,310,1352,1378]" pageId="10" pageNumber="11">
<heading id="D0E68180FFB0FFE92180FAE65F5C1214" bold="true" box="[189,310,1352,1378]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B965EAFEFFB0FFE92180FAE65F5C1214" bold="true" box="[189,310,1352,1378]" pageId="10" pageNumber="11">Diagnosis</emphasis>
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<paragraph id="8BAE36ECFFB0FFE92180FAD95D5A1062" blockId="10.[188,1399,1399,1813]" pageId="10" pageNumber="11">
The species is closest to
<taxonomicName id="4C114D6FFFB0FFE920EDFAD65CF012E7" authorityName="Constant &amp; Pham" authorityYear="2024" box="[464,666,1399,1425]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFB0FFE920EDFAD65CF012E7" box="[464,666,1399,1425]" italics="true" pageId="10" pageNumber="11">Zanna bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB0FFE9239FFAD95C9112E7" box="[674,763,1399,1425]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C114D6FFFB0FFE92239FAD65DEC12E7" authorityName="Constant &amp; Pham" authorityYear="2024" box="[772,902,1399,1425]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFB0FFE92239FAD65DEC12E7" box="[772,902,1399,1425]" italics="true" pageId="10" pageNumber="11">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB0FFE922B0FAD95D8F12E7" box="[909,997,1399,1425]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C114D6FFFB0FFE92525FAD65B7812E7" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1048,1298,1399,1425]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFB0FFE92525FAD65ACC12E7" box="[1048,1190,1399,1425]" italics="true" pageId="10" pageNumber="11">Z.limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB0FFE92425FAD95B1812E7" box="[1304,1394,1399,1425]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can be separated by the following characters: basal portion of head (
<figureCitation id="132A2A69FFB0FFE92566FA345A9B12C2" box="[1115,1265,1434,1460]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="10" pageNumber="11">Fig. 6B, EF</figureCitation>
) covered in large, rather well-spaced, black pitting with background of dense, very small black pitting (dense, irregular, rather large, coarse, sometimes coalescent black pitting in
<taxonomicName id="4C114D6FFFB0FFE9226AFA4F5D9A128C" authorityName="Constant &amp; Pham" authorityYear="2024" box="[855,1008,1504,1530]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bidoupana">
<emphasis id="B965EAFEFFB0FFE9226AFA4F5D9A128C" box="[855,1008,1504,1530]" italics="true" pageId="10" pageNumber="11">Z. bidoupana</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFB0FFE92531FA4E5ACE128D" box="[1036,1188,1504,1531]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3B, EF</figureCitation>
; moderately large, rather well-spaced, black pitting without background of dense, very small black pitting in
<taxonomicName id="4C114D6FFFB0FFE925E1F9AA5B0A116B" authorityName="Constant &amp; Pham" authorityYear="2024" box="[1244,1376,1539,1565]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFB0FFE925E1F9AA5B0A116B" box="[1244,1376,1539,1565]" italics="true" pageId="10" pageNumber="11">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFB0FFE92180F9895F0C1137" box="[189,358,1575,1601]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="10" pageNumber="11">Fig. 12B, EF</figureCitation>
); anal tube of male (
<figureCitation id="132A2A69FFB0FFE92368F9895CDB1137" box="[597,689,1575,1601]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="10" pageNumber="11">Fig. 7A</figureCitation>
) rather flattened dorsoventrally, 2.14 times as long as high in lateral view and with ventral angle at midlength (rather high in lateral view, 1.601.65 times as long as high, with ventral angle in basal half in
<taxonomicName id="4C114D6FFFB0FFE92340F9C05D7F11F1" authorityName="Constant &amp; Pham" authorityYear="2024" box="[637,789,1645,1671]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bidoupana">
<emphasis id="B965EAFEFFB0FFE92340F9C05D7F11F1" box="[637,789,1645,1671]" italics="true" pageId="10" pageNumber="11">Z. bidoupana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFB0FFE92277F9C05DBD11F1" authorityName="Constant" authorityYear="2024" box="[842,983,1645,1671]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFB0FFE92277F9C05DBD11F1" box="[842,983,1645,1671]" italics="true" pageId="10" pageNumber="11">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFB0FFE922CEF9C35A3A11F1" box="[1011,1104,1645,1671]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Figs 3A</figureCitation>
,
<figureCitation id="132A2A69FFB0FFE92566F9C35AFB11F1" box="[1115,1169,1645,1671]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="10" pageNumber="11">16A</figureCitation>
); pygofer of male in lateral view (
<figureCitation id="132A2A69FFB0FFE9206DF93E5FC311DD" box="[336,425,1680,1707]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="10" pageNumber="11">Fig. 7A</figureCitation>
) with posterior margin moderately sinuate, forming a large, broadly rounded posterior lobe in dorsal half (posterior margin projecting into a large posterior lobe angularly rounded in midheight in
<taxonomicName id="4C114D6FFFB0FFE921E4F9765F311187" authorityName="Constant &amp; Pham" authorityYear="2024" box="[217,347,1751,1777]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFB0FFE921E4F9765F311187" box="[217,347,1751,1777]" italics="true" pageId="10" pageNumber="11">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFB0FFE92048F9795FB71187" box="[373,477,1751,1777]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="10" pageNumber="11">Fig. 13A</figureCitation>
; posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle in
<taxonomicName id="4C114D6FFFB0FFE92333F9555CF61062" authorityName="Constant" authorityYear="2024" box="[526,668,1786,1812]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFB0FFE92333F9555CF61062" box="[526,668,1786,1812]" italics="true" pageId="10" pageNumber="11">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFB0FFE92384F9545D491062" box="[697,803,1786,1812]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="10" pageNumber="11">Fig. 16A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFB0FFE92180F8925A3310BC" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BAE36ECFFB0FFE92180F8925F2E1020" blockId="10.[189,324,1852,1878]" box="[189,324,1852,1878]" pageId="10" pageNumber="11">
<heading id="D0E68180FFB0FFE92180F8925F2E1020" bold="true" box="[189,324,1852,1878]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B965EAFEFFB0FFE92180F8925F2E1020" bold="true" box="[189,324,1852,1878]" pageId="10" pageNumber="11">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB0FFE92180F8C45A3310BC" blockId="10.[189,1399,1898,1995]" pageId="10" pageNumber="11">
The species epithet is a patronym dedicated to Mr Gerard “Gee” Chartier (Tatai,
<collectingCountry id="F306767CFFB0FFE92573F8C45AA310F2" box="[1102,1225,1898,1924]" name="Cambodia" pageId="10" pageNumber="11">Cambodia</collectingCountry>
) who collected the
<typeStatus id="54AA884EFFB0FFE921D0F8205F4A10DE" box="[237,288,1934,1960]" pageId="10" pageNumber="11">type</typeStatus>
series and documented the biology of the species, in acknowledgment of his enthusiastic involvement in the progress of the knowledge of the Cambodian entomofauna.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFB1FFE82180FEA75A6115C5" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BAE36ECFFB1FFE82180FEA75F061654" blockId="11.[189,364,264,291]" box="[189,364,264,291]" pageId="11" pageNumber="12">
<heading id="D0E68180FFB1FFE82180FEA75F061654" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B965EAFEFFB1FFE82180FEA75F061654" bold="true" box="[189,364,264,291]" pageId="11" pageNumber="12">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB1FFE821D7FE995F361627" blockId="11.[189,1399,311,443]" box="[234,348,311,337]" pageId="11" pageNumber="12">
<emphasis id="B965EAFEFFB1FFE821D7FE995F361627" bold="true" box="[234,348,311,337]" pageId="11" pageNumber="12">
<typeStatus id="54AA884EFFB1FFE821D7FE995F361627" box="[234,348,311,337]" pageId="11" pageNumber="12" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180FEF45B1D16CD" blockId="11.[189,1399,311,443]" pageId="11" pageNumber="12">
<materialsCitation id="3B793CB1FFB1FFE82180FEF45B1D16CD" collectingDate="2017-01-18" collectingDateMax="2017-02-02" collectingDateMin="2017-01-18" collectionCode="RBINS" collectorName="G. Chartier &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc." country="Cambodia" latitude="11.586945" location="Tatai" longLatPrecision="21" longitude="103.09722" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Koh Kong" typeStatus="holotype">
<collectingCountry id="F306767CFFB1FFE82180FEF45F311602" box="[189,347,346,372]" name="Cambodia" pageId="11" pageNumber="12">CAMBODIA</collectingCountry>
<specimenCount id="9D17FD65FFB1FFE82040FEF45FDB1603" box="[381,433,346,373]" count="1" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D5F80EFFB1FFE820FFFEF45CD21602" box="[450,696,346,373]" country="Cambodia" name="Koh Kong" pageId="11" pageNumber="12">Koh Kong Province</collectingRegion>
,
<location id="8ECE6037FFB1FFE823F5FEF45D6C1602" LSID="urn:lsid:plazi:treatment:03B887FAFFB0FFF0236CFB105CFD13C9:8ECE6037FFB1FFE823F5FEF45D6C1602" box="[712,774,346,372]" country="Cambodia" latitude="11.586945" longLatPrecision="21" longitude="103.09722" name="Tatai" pageId="11" pageNumber="12" stateProvince="Koh Kong">Tatai</location>
;
<geoCoordinate id="EE25502BFFB1FFE8222BFEF75DC41603" box="[790,942,345,373]" degrees="11" direction="north" minutes="35" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="13" value="11.586945">11°35ʹ13ʺ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFB1FFE82283FEF75A091603" box="[958,1123,345,373]" degrees="103" direction="east" minutes="05" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="50" value="103.09722">103°05ʹ50ʺ E</geoCoordinate>
;
<date id="FFAF102CFFB1FFE82549FEF45B181602" box="[1140,1394,346,372]" pageId="11" pageNumber="12" value="2017-01-18" valueMax="2017-02-02" valueMin="2017-01-18">
<collectingDate id="EFEBE9C4FFB1FFE82549FEF45B181602" box="[1140,1394,346,372]" pageId="11" pageNumber="12" value="2017-01-18" valueMax="2017-02-02" valueMin="2017-01-18">18 Jan.2 Feb. 2017</collectingDate>
</date>
; day time collecting;
<collectorName id="26E4533AFFB1FFE82084FED05C2B16E1" box="[441,577,381,408]" pageId="11" pageNumber="12">G. Chartier</collectorName>
leg.; I.G.: 33.551; “
<collectorName id="26E4533AFFB1FFE82201FED35A6716EE" box="[828,1037,381,408]" pageId="11" pageNumber="12">Coll. I.R.Sc.N.B.</collectorName>
, Cambodia,
<collectingRegion id="49D5F80EFFB1FFE82594FED35B1816EE" box="[1193,1394,381,408]" country="Cambodia" name="Koh Kong" pageId="11" pageNumber="12">Koh Kong prov.</collectingRegion>
, Tatai,
<geoCoordinate id="EE25502BFFB1FFE82038FE315FFD16CD" box="[261,407,415,443]" degrees="11" direction="north" minutes="35" orientation="latitude" pageId="11" pageNumber="12" precision="15" seconds="13" value="11.586945">11°35ʹ13ʺ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFB1FFE8209FFE315C2A16CD" box="[418,576,415,443]" degrees="103" direction="east" minutes="05" orientation="longitude" pageId="11" pageNumber="12" precision="15" seconds="50" value="103.09722">103°05ʹ50ʺ E</geoCoordinate>
,
<date id="FFAF102CFFB1FFE82371FE0F5C9816CD" box="[588,754,417,443]" pageId="11" pageNumber="12" value="2017-01-18" valueMax="2017-02-02" valueMin="2017-01-18">
<collectingDate id="EFEBE9C4FFB1FFE82371FE0F5C9816CD" box="[588,754,417,443]" pageId="11" pageNumber="12" value="2017-01-18" valueMax="2017-02-02" valueMin="2017-01-18">18.I-2.II.2017</collectingDate>
</date>
, day collecting, leg.
<collectorName id="26E4533AFFB1FFE822DCFE0F5A0C16CD" box="[993,1126,417,443]" pageId="11" pageNumber="12">G. Chartier</collectorName>
, I.G.: 33.551”;
<collectionCode id="ED00AE29FFB1FFE82424FE0F5B1916CD" box="[1305,1395,417,443]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="12" type="Museum">RBINS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAE36ECFFB1FFE821D7FE4C5F0D168A" blockId="11.[189,1399,482,579]" box="[234,359,482,508]" pageId="11" pageNumber="12">
<emphasis id="B965EAFEFFB1FFE821D7FE4C5F0D168A" bold="true" box="[234,359,482,508]" pageId="11" pageNumber="12">
<typeStatus id="54AA884EFFB1FFE821D7FE4C5F0D168A" box="[234,359,482,508]" pageId="11" pageNumber="12" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180FDAB5C691535" blockId="11.[189,1399,482,579]" pageId="11" pageNumber="12">
<materialsCitation id="3B793CB1FFB1FFE82180FDAB5A5A1569" box="[189,1072,517,544]" collectingDate="2017-01-18" collectingDateMax="2017-02-02" collectingDateMin="2017-01-18" collectionCode="RBINS" collectorName="G. Chartier &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc." country="Cambodia" latitude="11.586945" location="Tatai" longLatPrecision="21" longitude="103.09722" pageId="11" pageNumber="12" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Koh Kong" typeStatus="paratype">
<collectingCountry id="F306767CFFB1FFE82180FDAB5F311569" box="[189,347,517,543]" name="Cambodia" pageId="11" pageNumber="12">CAMBODIA</collectingCountry>
<specimenCount id="9D17FD65FFB1FFE8204BFDAB5FD11556" box="[374,443,517,544]" count="3" pageId="11" pageNumber="12" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D17FD65FFB1FFE820F4FDAB5C671556" box="[457,525,517,544]" count="4" pageId="11" pageNumber="12" type="female">4 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED00AE29FFB1FFE822E7FDAB5A5A1569" box="[986,1072,517,543]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="12" type="Museum">RBINS</collectionCode>
</materialsCitation>
<materialsCitation id="3B793CB1FFB1FFE82571FDAB5F951535" collectingDate="2017-01-18" collectingDateMax="2017-02-02" collectingDateMin="2017-01-18" collectionCode="RBINS" collectorName="G. Chartier &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc." country="Cambodia" latitude="11.586945" location="Tatai" longLatPrecision="21" longitude="103.09722" pageId="11" pageNumber="12" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Koh Kong" typeStatus="paratype">
<specimenCount id="9D17FD65FFB1FFE82571FDAB5A171569" box="[1100,1149,517,543]" count="1" pageId="11" pageNumber="12" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D17FD65FFB1FFE825BEFDAB5AC41556" box="[1155,1198,517,544]" count="1" pageId="11" pageNumber="12" type="female">1 ♀</specimenCount>
; same collection data as for holotype; RUPP
</materialsCitation>
.
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180FDC45C5E15F2" blockId="11.[189,564,618,644]" box="[189,564,618,644]" pageId="11" pageNumber="12">
<heading id="D0E68180FFB1FFE82180FDC45C5E15F2" bold="true" box="[189,564,618,644]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B965EAFEFFB1FFE82180FDC45C5E15F2" bold="true" box="[189,564,618,644]" pageId="11" pageNumber="12">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180FD375A6115C5" blockId="11.[189,1035,664,691]" box="[189,1035,664,691]" pageId="11" pageNumber="12">
<materialsCitation id="3B793CB1FFB1FFE82180FD375A6D15C5" box="[189,1031,664,691]" collectingDate="2017-01-18" collectingDateMax="2017-02-02" collectingDateMin="2017-01-18" collectionCode="RBINS" collectorName="G. Chartier &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc." country="Cambodia" latitude="11.586945" location="Tatai" longLatPrecision="21" longitude="103.09722" pageId="11" pageNumber="12" specimenCount="1" stateProvince="Koh Kong">
<collectingCountry id="F306767CFFB1FFE82180FD375F3115C5" box="[189,347,665,691]" name="Cambodia" pageId="11" pageNumber="12">CAMBODIA</collectingCountry>
• 6 nymphs; same collection data as for holotype;
<collectionCode id="ED00AE29FFB1FFE82290FD375A6D15C5" box="[941,1031,665,691]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="11" pageNumber="12" type="Museum">RBINS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFB1FFF02180FD745DD516FA" lastPageId="19" lastPageNumber="20" pageId="11" pageNumber="12" type="description">
<paragraph id="8BAE36ECFFB1FFE82180FD745F241582" blockId="11.[189,334,730,756]" box="[189,334,730,756]" pageId="11" pageNumber="12">
<heading id="D0E68180FFB1FFE82180FD745F241582" bold="true" box="[189,334,730,756]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B965EAFEFFB1FFE82180FD745F241582" bold="true" box="[189,334,730,756]" pageId="11" pageNumber="12">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180FCA75D65141C" blockId="11.[189,1399,777,874]" pageId="11" pageNumber="12">
MEASUREMENTS AND RATIOS. LT:
<specimenCount id="9D17FD65FFB1FFE8236DFCA75C0C1455" box="[592,614,777,803]" count="1" pageId="11" pageNumber="12" type="male"></specimenCount>
(n= 4):
<quantity id="4CE99B09FFB1FFE823F2FCA75D561455" box="[719,828,777,803]" metricMagnitude="-2" metricUnit="m" metricValue="4.08" pageId="11" pageNumber="12" unit="mm" value="40.8">40.8 mm</quantity>
(38.441.8);
<specimenCount id="9D17FD65FFB1FFE822D8FCA75D911455" box="[997,1019,777,803]" count="1" pageId="11" pageNumber="12" type="female"></specimenCount>
(n =4): 41.8 (39.743.2). LTg/ BTg =3.8; LH/BH =4.8; LF/BF=6.0; BH/BPrH =2.2; LH/LT =0.41; wingspan (extrapolated):
<specimenCount id="9D17FD65FFB1FFE82180FCE15EBD141F" box="[189,215,847,873]" count="1" pageId="11" pageNumber="12" type="male"></specimenCount>
: 48.0 mm (45.249.3),
<specimenCount id="9D17FD65FFB1FFE820D7FCE15C68141F" box="[490,514,847,873]" count="1" pageId="11" pageNumber="12" type="female"></specimenCount>
:
<quantity id="4CE99B09FFB1FFE82332FCE15C12141F" box="[527,632,847,873]" metricMagnitude="-2" metricUnit="m" metricValue="4.93" pageId="11" pageNumber="12" unit="mm" value="49.3">49.3 mm</quantity>
(46.850.9).
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180FC3E5A5F13B2" blockId="11.[189,1399,912,1220]" pageId="11" pageNumber="12">
HEAD (
<figureCitation id="132A2A69FFB1FFE82032FC3E5FC714DC" box="[271,429,912,938]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="11" pageNumber="12">Fig. 6B, EF</figureCitation>
). Strongly elongate, representing 41% of total length, gently and evenly narrowing towards apex with apical ⅓ more or less parallel-sided in dorsal view; apex slightly widening, then obliquely cut in lateral view and with incomplete, rim-shaped orange carina (missing in ventral portion); rather pale greyish brown entirely covered in dense, irregular black pitting with smaller pits on all surface and some dispersed, moderately large, not or very rarely coalescent pits in basal 2/3; some larger black pustules on side, with always 25 along carina of vertex. Lateral carinae of frons moderate, weakly zigzagged, more prominent towards apex, upcurved near apex and merging into a short dorsal carina reaching apex; lateral carinae of vertex moderate, merging with upcurved part of lateral carinae of frons; median carina of vertex obsolete. Labium pale brown, reaching mesocoxae.
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180FB455C37123A" blockId="11.[189,1399,1259,1356]" pageId="11" pageNumber="12">
THORAX (
<figureCitation id="132A2A69FFB1FFE8200FFB455FB31273" box="[306,473,1259,1285]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="11" pageNumber="12">Fig. 6B, EF</figureCitation>
). Pro- and mesonotum rather pale greyish brown entirely covered in dense, irregular, moderate black pitting with pits smaller than space between them and not (or very rarely) coalescent; median carina obsolete.
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180FADD5F3D1148" blockId="11.[189,1399,1395,1598]" pageId="11" pageNumber="12">
TEGMINA (
<figureCitation id="132A2A69FFB1FFE82000FADD5F8712FB" box="[317,493,1395,1421]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="11" pageNumber="12">Fig. 6A, CD</figureCitation>
). Strongly elongate and reticulate, pale greyish brown, with more or less conspicuous, oblique brown band at basal ⅓, not extending on clavus; all surface irregularly covered in small to moderately large black spots, mostly on veins, larger ones slightly protruding; costal and postclaval margins subparallel, weakly diverging from base towards level of apex of clavus, then curved and weakly tapering towards apex; apical margin rounded. Main veins yellowish orange in basal portion. Clavus open.
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180F9CB5B2211D4" blockId="11.[189,1398,1635,1698]" pageId="11" pageNumber="12">VENATION. ScP+R forked in basal ⅓, MP forked more basally in basal 1/4; CuA forked near apex of clavus; PCu and A1 fused near apex of clavus; PCu+A1 fused with postclaval margin at nodal line.</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180F9645D751070" blockId="11.[189,1399,1737,1799]" pageId="11" pageNumber="12">
HIND WINGS (
<figureCitation id="132A2A69FFB1FFE8205CF9675F891195" box="[353,483,1737,1763]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="11" pageNumber="12">Fig. 6A, C</figureCitation>
). Subhyaline, milky white with veins slightly darker, vein PCu pale brown in basal half; postclaval margin emarginate at A1 vein.
</paragraph>
<paragraph id="8BAE36ECFFB1FFE82180F8805FCB10C7" blockId="11.[189,1399,1838,1969]" pageId="11" pageNumber="12">
VENATION. ScP+R and MP forked at midlength; CuA forked slightly more basally then MP fork; PCu forked at basal ⅓; A1 forked close to base, A1
<subScript id="179534A9FFB1FFE823EAF8CA5C8A1004" attach="left" box="[727,736,1892,1906]" fontSize="6" pageId="11" pageNumber="12">2</subScript>
fused with A
<quantity id="4CE99B09FFB1FFE82241F8FF5DC2101D" box="[892,936,1873,1899]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="11" pageNumber="12" unit="in" value="2.0">2 in</quantity>
large distal portion and A1
<subScript id="179534A9FFB1FFE825DBF8CA5A851004" attach="none" box="[1254,1263,1892,1906]" fontSize="6" pageId="11" pageNumber="12">2</subScript>
+A2 forked slightly before reaching postclaval margin. Numerous crossveins in large portion along apical and postclaval margins.
</paragraph>
<caption id="DF6E6664FFB6FFEF2184F8245FBF109C" ID-DOI="http://doi.org/10.5281/zenodo.13837042" ID-Zenodo-Dep="13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="12" pageNumber="13" startId="12.[185,226,1930,1956]" targetBox="[252,1334,322,1889]" targetPageId="12">
<paragraph id="8BAE36ECFFB6FFEF2184F8245FBF109C" blockId="12.[185,1395,1930,2026]" pageId="12" pageNumber="13">
<emphasis id="B965EAFEFFB6FFEF2184F8245F6210D2" bold="true" box="[185,264,1930,1956]" pageId="12" pageNumber="13">Fig. 6.</emphasis>
<taxonomicName id="4C114D6FFFB6FFEF202CF8255C5610D2" authority="Constant" authorityName="Constant" authorityYear="2024" box="[273,572,1930,1956]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFB6FFEF202CF8255FA710D2" box="[273,461,1930,1956]" italics="true" pageId="12" pageNumber="13">Zanna chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB6FFEF2378F8245CC910D2" box="[581,675,1930,1956]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS).
<emphasis id="B965EAFEFFB6FFEF2282F8245DBE10D2" bold="true" box="[959,980,1930,1956]" pageId="12" pageNumber="13">A</emphasis>
. Habitus dorsal view.
<emphasis id="B965EAFEFFB6FFEF25E2F8245A9910D2" bold="true" box="[1247,1267,1930,1956]" pageId="12" pageNumber="13">B</emphasis>
. Head and thorax, dorsal view.
<emphasis id="B965EAFEFFB6FFEF2097F8035FD510B1" bold="true" box="[426,447,1965,1991]" pageId="12" pageNumber="13">C</emphasis>
. Habitus, ventral view.
<emphasis id="B965EAFEFFB6FFEF23E7F8035C8510B1" bold="true" box="[730,751,1965,1991]" pageId="12" pageNumber="13">D</emphasis>
. Habitus, lateral view.
<emphasis id="B965EAFEFFB6FFEF253FF8035A7C10B1" bold="true" box="[1026,1046,1965,1991]" pageId="12" pageNumber="13">E</emphasis>
. Head, ventral view.
<emphasis id="B965EAFEFFB6FFEF242EF8035B4F10B1" bold="true" box="[1299,1317,1965,1991]" pageId="12" pageNumber="13">F</emphasis>
. Head and thorax, lateral view.
</paragraph>
</caption>
<caption id="DF6E6664FFB7FFEE2180F82B5B1D1090" ID-DOI="http://doi.org/10.5281/zenodo.13837046" ID-Zenodo-Dep="13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="13" pageNumber="14" startId="13.[189,228,1925,1951]" targetBox="[232,1368,257,1882]" targetPageId="13">
<paragraph id="8BAE36ECFFB7FFEE2180F82B5B1D1090" blockId="13.[189,1399,1925,2022]" pageId="13" pageNumber="14">
<emphasis id="B965EAFEFFB7FFEE2180F82B5F6310D6" bold="true" box="[189,265,1925,1952]" pageId="13" pageNumber="14">Fig. 7.</emphasis>
<taxonomicName id="4C114D6FFFB7FFEE202FF8285C5910D6" authority="Constant" authorityName="Constant" authorityYear="2024" box="[274,563,1925,1952]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFB7FFEE202FF8285FA210E9" box="[274,456,1925,1951]" italics="true" pageId="13" pageNumber="14">Zanna chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB7FFEE2301F8285CFD10D6" box="[572,663,1926,1952]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS), terminalia.
<emphasis id="B965EAFEFFB7FFEE2511F82B5A0910E9" bold="true" box="[1068,1123,1925,1951]" pageId="13" pageNumber="14">AD</emphasis>
. Pygofer, anal tube and gonostyli.
<emphasis id="B965EAFEFFB7FFEE2009F8065F2310B4" bold="true" box="[308,329,1960,1986]" pageId="13" pageNumber="14">A</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFB7FFEE231FF8075C5C10B5" bold="true" box="[546,566,1961,1987]" pageId="13" pageNumber="14">B</emphasis>
. Caudal view.
<emphasis id="B965EAFEFFB7FFEE23DCF8065C9C10B4" bold="true" box="[737,758,1960,1986]" pageId="13" pageNumber="14">C</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFB7FFEE22A0F8075DD810B5" bold="true" box="[925,946,1961,1987]" pageId="13" pageNumber="14">D</emphasis>
. Ventral view.
<emphasis id="B965EAFEFFB7FFEE2562F8075AFC10B5" bold="true" box="[1119,1174,1961,1987]" pageId="13" pageNumber="14">EH</emphasis>
. Aedeagus.
<emphasis id="B965EAFEFFB7FFEE241EF8075B5C10B5" bold="true" box="[1315,1334,1961,1987]" pageId="13" pageNumber="14">E</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFB7FFEE207AF8625F331090" bold="true" box="[327,345,1996,2022]" pageId="13" pageNumber="14">F</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFB7FFEE20C9F8655C611093" bold="true" box="[500,523,1995,2021]" pageId="13" pageNumber="14">G</emphasis>
. Laterodorsal view.
<emphasis id="B965EAFEFFB7FFEE23D4F8625D6A1090" bold="true" box="[745,768,1996,2022]" pageId="13" pageNumber="14">H</emphasis>
. Ventral view.Abbreviations: see Materials and methods.
</paragraph>
</caption>
<caption id="DF6E6664FFB4FFED2183F8305FB110AD" ID-DOI="http://doi.org/10.5281/zenodo.13837048" ID-Zenodo-Dep="13837048" httpUri="https://zenodo.org/record/13837048/files/figure.png" pageId="14" pageNumber="15" startId="14.[190,231,1950,1976]" targetBox="[201,1399,285,1873]" targetPageId="14">
<paragraph id="8BAE36ECFFB4FFED2183F8305FB110AD" blockId="14.[190,1400,1950,2012]" pageId="14" pageNumber="15">
<emphasis id="B965EAFEFFB4FFED2183F8305F6410CE" bold="true" box="[190,270,1950,1976]" pageId="14" pageNumber="15">Fig. 8.</emphasis>
<taxonomicName id="4C114D6FFFB4FFED2024F8315C2D10CE" authority="Constant" authorityName="Constant" authorityYear="2024" box="[281,583,1950,1976]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFB4FFED2024F8315FBC10CE" box="[281,470,1950,1976]" italics="true" pageId="14" pageNumber="15">Zanna chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB4FFED236FF8305CD810CE" box="[594,690,1950,1976]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
, nymph.
<emphasis id="B965EAFEFFB4FFED2219F8305D5310CE" bold="true" box="[804,825,1950,1976]" pageId="14" pageNumber="15">A</emphasis>
. Habitus dorsal view.
<emphasis id="B965EAFEFFB4FFED2576F8305A3510CE" bold="true" box="[1099,1119,1950,1976]" pageId="14" pageNumber="15">B</emphasis>
. Habitus, ventral view.
<emphasis id="B965EAFEFFB4FFED2183F86F5EB910AD" bold="true" box="[190,211,1985,2011]" pageId="14" pageNumber="15">C</emphasis>
. Habitus, lateral view.
</paragraph>
</caption>
<caption id="DF6E6664FFB5FFEC2180F8C65FD2109A" ID-DOI="http://doi.org/10.5281/zenodo.13837052" ID-Zenodo-Dep="13837052" httpUri="https://zenodo.org/record/13837052/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,230,1896,1922]" targetBox="[213,1370,268,1848]" targetPageId="15">
<paragraph id="8BAE36ECFFB5FFEC2180F8C65FD2109A" blockId="15.[189,1399,1895,2029]" pageId="15" pageNumber="16">
<emphasis id="B965EAFEFFB5FFEC2180F8C65F6710F5" bold="true" box="[189,269,1896,1923]" pageId="15" pageNumber="16">Fig. 9.</emphasis>
<taxonomicName id="4C114D6FFFB5FFEC202AF8C75C2E10F5" authority="Constant" authorityName="Constant" authorityYear="2024" box="[279,580,1896,1923]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFB5FFEC202AF8C75FBF10F4" box="[279,469,1896,1922]" italics="true" pageId="15" pageNumber="16">Zanna chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB5FFEC2373F8C75CC710F5" box="[590,685,1897,1923]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
, in nature in Tatai, Cambodia (photographs © G. Chartier).
<emphasis id="B965EAFEFFB5FFEC2180F8255EB810D3" bold="true" box="[189,210,1931,1957]" pageId="15" pageNumber="16">A</emphasis>
. Adult on
<taxonomicName id="4C114D6FFFB5FFEC2068F8225C6710D0" box="[341,525,1932,1958]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAFEFFB5FFEC2068F8225F8B10D0" box="[341,481,1932,1958]" italics="true" pageId="15" pageNumber="16">Desmodium</emphasis>
sp.
</taxonomicName>
, 20 Jan. 2017.
<emphasis id="B965EAFEFFB5FFEC23F4F8225CB010D0" bold="true" box="[713,730,1932,1958]" pageId="15" pageNumber="16">B</emphasis>
. Adult on unidentified plant, 17 Jan. 2017.
<emphasis id="B965EAFEFFB5FFEC25D5F8255A9710D3" bold="true" box="[1256,1277,1931,1957]" pageId="15" pageNumber="16">C</emphasis>
. Adult on
<emphasis id="B965EAFEFFB5FFEC2183F8015FB210BF" box="[190,472,1967,1993]" italics="true" pageId="15" pageNumber="16">Melastomasanguineum</emphasis>
, lateral view, 17 Jan. 2017.
<emphasis id="B965EAFEFFB5FFEC220DF8015D2F10BF" bold="true" box="[816,837,1967,1993]" pageId="15" pageNumber="16">D</emphasis>
. Habitat, position of nymphs marked with red arrows, 20 Jan. 2017.
</paragraph>
</caption>
<caption id="DF6E6664FFAAFFF32180F8C85B5A109C" ID-DOI="http://doi.org/10.5281/zenodo.13837056" ID-Zenodo-Dep="13837056" httpUri="https://zenodo.org/record/13837056/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,230,1894,1920]" targetBox="[238,1351,274,1856]" targetPageId="16">
<paragraph id="8BAE36ECFFAAFFF32180F8C85B5A109C" blockId="16.[188,1398,1894,2026]" pageId="16" pageNumber="17">
<emphasis id="B965EAFEFFAAFFF32180F8C85F7610F6" bold="true" box="[189,284,1894,1920]" pageId="16" pageNumber="17">Fig. 10.</emphasis>
<taxonomicName id="4C114D6FFFAAFFF3201BF8C95C3F10F6" authority="Constant" authorityName="Constant" authorityYear="2024" box="[294,597,1894,1920]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFAAFFF3201BF8C95F8810F6" box="[294,482,1894,1920]" italics="true" pageId="16" pageNumber="17">Zanna chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFAAFFF32360F8C85CD610F6" box="[605,700,1894,1920]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
, nymphs in nature in Tatai, Cambodia, on
<taxonomicName id="4C114D6FFFAAFFF325FFF8C85B1C10F6" box="[1218,1398,1894,1920]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAFEFFAAFFF325FFF8C85B2410F6" box="[1218,1358,1894,1920]" italics="true" pageId="16" pageNumber="17">Desmodium</emphasis>
sp.
</taxonomicName>
(photographs © G. Chartier).
<emphasis id="B965EAFEFFAAFFF3231FF8275C3010D5" bold="true" box="[546,602,1929,1955]" pageId="16" pageNumber="17">AB</emphasis>
. 4
<superScript id="7C649BA4FFAAFFF32344F8265CEC10E0" attach="left" box="[633,646,1928,1942]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
instar nymphs.
<emphasis id="B965EAFEFFAAFFF32276F8275D3410D5" bold="true" box="[843,862,1929,1955]" pageId="16" pageNumber="17">A</emphasis>
. Laterodorsal view, 18 Jan. 2017.
<emphasis id="B965EAFEFFAAFFF325C2F8275B7910D5" bold="true" box="[1279,1299,1929,1955]" pageId="16" pageNumber="17">B</emphasis>
. Lateral view, 20 Jan. 2017.
<emphasis id="B965EAFEFFAAFFF3209EF8025FB610B0" bold="true" box="[419,476,1964,1990]" pageId="16" pageNumber="17">CD</emphasis>
. 5
<superScript id="7C649BA4FFAAFFF320C4F8025C6C10CC" attach="left" box="[505,518,1964,1978]" fontSize="6" pageId="16" pageNumber="17">th</superScript>
instar nymphs, 17 Jan. 2017.
<emphasis id="B965EAFEFFAAFFF32262F8025D1E10B0" bold="true" box="[863,884,1964,1990]" pageId="16" pageNumber="17">C</emphasis>
. Lateral view.
<emphasis id="B965EAFEFFAAFFF3251CF8025A5C10B0" bold="true" box="[1057,1078,1964,1990]" pageId="16" pageNumber="17">D</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFAAFFF325E3F8025B7810B0" bold="true" box="[1246,1298,1964,1990]" pageId="16" pageNumber="17">EF</emphasis>
. Groups of nymphs.
<emphasis id="B965EAFEFFAAFFF32075F87E5F36109C" bold="true" box="[328,348,2000,2026]" pageId="16" pageNumber="17">E</emphasis>
. On vertical twig, 20 Jan. 2017.
<emphasis id="B965EAFEFFAAFFF323E1F87E5C84109C" bold="true" box="[732,750,2000,2026]" pageId="16" pageNumber="17">F</emphasis>
. On creeping twig close to ground, 19 Jan. 2019.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFABFFF22180FEA75A96156C" blockId="17.[189,1399,265,538]" pageId="17" pageNumber="18">
LEGS (
<figureCitation id="132A2A69FFABFFF22036FEA75FDC1655" box="[267,438,265,291]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="17" pageNumber="18">Fig. 6A, CD</figureCitation>
). Pale greyish stramineous, pilose, rather short and robust. Pro- and mesocoxae with some small black markings; all femora with few small to minute black spots and conspicuous black spots aligned in a ring in distal ⅓, sometimes coalescent; pro- and mesotibiae with some small black spots, regularly in basal ¼, and with apex brown; pro- and mesotarsi with segments darker apically; metatibiae weakly brown basally with all spines apically black, 56 lateral spines and 8 apical spines; first metatarsomere with 1115 apical spines ventrally; second metatarsomere with 910 apical spines ventrally; first and second metatarsomere with a dense pad of microsetae ventrally; third metatarsomere elongate with a subapical blackish brown ring. Metatibiotarsal formula: (56) 8/1115/910.
</paragraph>
<paragraph id="8BAE36ECFFABFFF22180FDEF5C9D1508" blockId="17.[189,1398,577,638]" pageId="17" pageNumber="18">
ABDOMEN (
<figureCitation id="132A2A69FFABFFF22079FDEF5FAD152D" box="[324,455,577,603]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="17" pageNumber="18">Fig. 6A, C</figureCitation>
). Tergites smooth and shiny, yellowish brown; sternites (except pregenital one) yellowish brown with irregular small black dots.
</paragraph>
<paragraph id="8BAE36ECFFABFFF22180FD085A0E13FF" blockId="17.[189,1399,677,1161]" pageId="17" pageNumber="18">
MALE TERMINALIA (
<figureCitation id="132A2A69FFABFFF2209FFD0B5F8115B6" box="[418,491,677,704]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
). Pygofer (
<emphasis id="B965EAFEFFABFFF2234DFD085CE515C9" box="[624,655,678,703]" italics="true" pageId="17" pageNumber="18">Py</emphasis>
<figureCitation id="132A2A69FFABFFF2238DFD0B5D4515B6" box="[688,815,677,704]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="17" pageNumber="18">Fig. 7AD</figureCitation>
) about 1.96 times as high as long in lateral view, robust with posterior margin sinuate, forming a large posterior lobe in dorsal half; lobe broadly rounded; sides rather strongly excavate in dorsal portion (visible in dorsal aspect,
<figureCitation id="132A2A69FFABFFF222D0FD425A2C1470" box="[1005,1094,748,774]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="17" pageNumber="18">Fig. 7C</figureCitation>
). Anal tube (
<emphasis id="B965EAFEFFABFFF225EBFD425A9C1470" box="[1238,1270,748,774]" italics="true" pageId="17" pageNumber="18">An</emphasis>
<figureCitation id="132A2A69FFABFFF2242CFD425EBB145F" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="17" pageNumber="18">Fig. 7A C</figureCitation>
) robust, relatively flattened, 2.14 times as long as high and with ventral margin oblique in distal half and weakly angular in midlength in lateral view; 1.24 times as long as wide, with sides diverging towards apex and with apical margin incurved in dorsal view; oval in caudal view; epiproct and paraproct rather strongly supassing anal tube, rounded posteriorly in dorsal aspect. Gonostyli (
<emphasis id="B965EAFEFFABFFF222EEFCD75D8214E5" box="[979,1000,889,915]" italics="true" pageId="17" pageNumber="18">G</emphasis>
<figureCitation id="132A2A69FFABFFF22539FCD75AEB14E5" box="[1028,1153,889,915]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="17" pageNumber="18">Fig. 7AD</figureCitation>
) elongate, 2.62 times as long as high; in lateral view, ventral margin broadly rounded and dorsal margin rather broadly rounded after lateral hook (
<emphasis id="B965EAFEFFABFFF220A3FC115FA914AF" box="[414,451,959,985]" italics="true" pageId="17" pageNumber="18">lhg</emphasis>
) but more strongly rounded at about midlength, apical margin rather narrowly rounded; lateral hook with basal portion projecting posterolaterad and apex strongly curved lateroventrad; in ventral view, lateral protrusion at about midlength, rather strongly marked and with posterior angle curved. Aedeagus (
<figureCitation id="132A2A69FFABFFF220A7FB875C7F1335" box="[410,533,1065,1091]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="17" pageNumber="18">Fig. 7EH</figureCitation>
) rather elongate with lateral processes (
<emphasis id="B965EAFEFFABFFF222DFFB875A6D1335" box="[994,1031,1065,1091]" italics="true" pageId="17" pageNumber="18">lpa</emphasis>
) forming multiple lobes; dorsal process (
<emphasis id="B965EAFEFFABFFF22018FBE25F3A1310" box="[293,336,1100,1126]" italics="true" pageId="17" pageNumber="18">dpa</emphasis>
) weakly protruding dorsally in lateral view; dorsal endosomal process (
<emphasis id="B965EAFEFFABFFF225ABFBE25AD51310" box="[1174,1215,1100,1126]" italics="true" pageId="17" pageNumber="18">dep</emphasis>
) in dorsal view subtriangular with lateral margins nearly straight and median slit in distal portion.
</paragraph>
<paragraph id="8BAE36ECFFABFFF221D7FB1F5CCC13BD" blockId="17.[189,1399,1200,1509]" box="[234,678,1200,1227]" pageId="17" pageNumber="18">
<emphasis id="B965EAFEFFABFFF221D7FB1F5F2D13BD" bold="true" box="[234,327,1201,1227]" pageId="17" pageNumber="18">Nymph</emphasis>
(fifth instar
<figureCitation id="132A2A69FFABFFF220D2FB1E5C5313BD" box="[495,569,1200,1227]" captionStart="Fig" captionStartId="14.[190,231,1950,1976]" captionTargetBox="[201,1399,285,1873]" captionTargetId="figure-25@14.[189,1399,275,1884]" captionTargetPageId="14" captionText="Fig. 8. Zanna chartieri Constant sp. nov., nymph. A. Habitus dorsal view. B. Habitus, ventral view. C. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837048" httpUri="https://zenodo.org/record/13837048/files/figure.png" pageId="17" pageNumber="18">Figs 8</figureCitation>
,
<figureCitation id="132A2A69FFABFFF2237BFB1F5CF513BD" box="[582,671,1201,1227]" captionStart="Fig" captionStartId="16.[189,230,1894,1920]" captionTargetBox="[238,1351,274,1856]" captionTargetId="figure-84@16.[238,1351,274,1856]" captionTargetPageId="16" captionText="Fig. 10. Zanna chartieri Constant sp. nov., nymphs in nature in Tatai, Cambodia, on Desmodium sp. (photographs © G. Chartier). AB. 4th instar nymphs. A. Laterodorsal view, 18 Jan. 2017. B. Lateral view, 20 Jan. 2017. CD. 5th instar nymphs, 17 Jan. 2017. C. Lateral view. D. Dorsal view. EF. Groups of nymphs. E. On vertical twig, 20 Jan. 2017. F. On creeping twig close to ground, 19 Jan. 2019." figureDoi="http://doi.org/10.5281/zenodo.13837056" httpUri="https://zenodo.org/record/13837056/files/figure.png" pageId="17" pageNumber="18">10AD</figureCitation>
)
</paragraph>
<paragraph id="8BAE36ECFFABFFF22180FB7A5AAD1293" blockId="17.[189,1399,1200,1509]" pageId="17" pageNumber="18">
HEAD (
<figureCitation id="132A2A69FFABFFF22030FB7A5FE41398" box="[269,398,1236,1262]" captionStart="Fig" captionStartId="14.[190,231,1950,1976]" captionTargetBox="[201,1399,285,1873]" captionTargetId="figure-25@14.[189,1399,275,1884]" captionTargetPageId="14" captionText="Fig. 8. Zanna chartieri Constant sp. nov., nymph. A. Habitus dorsal view. B. Habitus, ventral view. C. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837048" httpUri="https://zenodo.org/record/13837048/files/figure.png" pageId="17" pageNumber="18">Figs 8AC</figureCitation>
,
<figureCitation id="132A2A69FFABFFF220A5FB7A5F991398" box="[408,499,1236,1262]" captionStart="Fig" captionStartId="16.[189,230,1894,1920]" captionTargetBox="[238,1351,274,1856]" captionTargetId="figure-84@16.[238,1351,274,1856]" captionTargetPageId="16" captionText="Fig. 10. Zanna chartieri Constant sp. nov., nymphs in nature in Tatai, Cambodia, on Desmodium sp. (photographs © G. Chartier). AB. 4th instar nymphs. A. Laterodorsal view, 18 Jan. 2017. B. Lateral view, 20 Jan. 2017. CD. 5th instar nymphs, 17 Jan. 2017. C. Lateral view. D. Dorsal view. EF. Groups of nymphs. E. On vertical twig, 20 Jan. 2017. F. On creeping twig close to ground, 19 Jan. 2019." figureDoi="http://doi.org/10.5281/zenodo.13837056" httpUri="https://zenodo.org/record/13837056/files/figure.png" pageId="17" pageNumber="18">10AD</figureCitation>
). Pale grey, strongly elongate, representing nearly half of total length, flattened dorsally and with well-marked longitudinal carinae; evenly tapering towards apex; apex slightly dilated, obliquely truncate, and with incomplete, rim-shaped orange carina (missing in ventral portion); carinae with protruding black spots, black spots limited to distal portion on lateral carinae of vertex; some black spots around eyes and on clypeus; minute tubercles between lateral and sublateral carinae of frons, more densely grouped towards base and towards apex. Lateral carinae of frons upcurved near apex and merging into a short dorsal carina reaching apex; lateral carinae of vertex merging with upcurved part of lateral carinae of frons. Labium pale brown with apex dark brown, reaching mesocoxae.
</paragraph>
<paragraph id="8BAE36ECFFABFFF22180F9A25D5D106A" blockId="17.[188,1399,1547,1820]" pageId="17" pageNumber="18">
THORAX (
<figureCitation id="132A2A69FFABFFF2200EF9A55FD71150" box="[307,445,1547,1574]" captionStart="Fig" captionStartId="14.[190,231,1950,1976]" captionTargetBox="[201,1399,285,1873]" captionTargetId="figure-25@14.[189,1399,275,1884]" captionTargetPageId="14" captionText="Fig. 8. Zanna chartieri Constant sp. nov., nymph. A. Habitus dorsal view. B. Habitus, ventral view. C. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837048" httpUri="https://zenodo.org/record/13837048/files/figure.png" pageId="17" pageNumber="18">Fig. 8A, C</figureCitation>
). Pale grey with irregular, rather wide, laterodorsal black stripe extending on pronotum and basal portion of mesonotum. Pronotum with anterior margin projecting cephalad, largely covering base of head, and with numerous minute tubercles on lateral fields and lateroventral lobes; two subparallel, sinuate lateral carinae, ventral one not visible in dorsal aspect. Mesonotum about 1.75 times as long as pronotum, with black spot in middle of well-marked peridiscal carinae, group of minute tubercles in lateral fields and longitudinal brown marking on buds of tegmina; buds of tegmina reaching apex of buds of hind wings. Metanotum about as long as pronotum, with two black spots on dorsolateral carinae and group of minute tubercles in lateral fields.
</paragraph>
<paragraph id="8BAE36ECFFABFFF02180F8ED5DD516FA" blockId="17.[189,1399,1859,1991]" lastBlockId="19.[189,1399,265,397]" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
LEGS (
<figureCitation id="132A2A69FFABFFF22037F8ED5FE6102B" box="[266,396,1859,1885]" captionStart="Fig" captionStartId="14.[190,231,1950,1976]" captionTargetBox="[201,1399,285,1873]" captionTargetId="figure-25@14.[189,1399,275,1884]" captionTargetPageId="14" captionText="Fig. 8. Zanna chartieri Constant sp. nov., nymph. A. Habitus dorsal view. B. Habitus, ventral view. C. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837048" httpUri="https://zenodo.org/record/13837048/files/figure.png" pageId="17" pageNumber="18">Fig. 8AC</figureCitation>
). Pale grey, pilose, rather short and robust. Coxae with some small black markings; metacoxae with gear-shaped structures. Pro- and mesofemora with more or less coalescent black spots aligned in a ring in distal ⅓. Pro- and mesotibiae with 3 evenly spaced, more or less complete black rings, with first one at basal ¼ and last one at apex. Pro- and mesotarsi black with basal half of apical segment pale grey. Metafemora with irregular black ring at distal ⅓ and more or less coalescent black spots in basal portion. Metatibiae dorsally with narrow, irregular blackish brown stripe at basal ⅓; ventrally with basal dark brown marking and black spots in distal half; laterally with irregular black spots; 5 lateral and 8 apical spines tipped with blackish brown. Metatarsi pale grey with apical half of last segment black; first metatarsomere with 9 apical spines ventrally; second metatarsomere with 89 apical spines ventrally; first and second metatarsomere with a dense pad of microsetae ventrally. Metatibiotarsal formula: (5) 8/9/89. ABDOMEN (
<figureCitation id="132A2A69FFA9FFF0207CFEA75FAB1655" box="[321,449,265,291]" captionStart="Fig" captionStartId="14.[190,231,1950,1976]" captionTargetBox="[201,1399,285,1873]" captionTargetId="figure-25@14.[189,1399,275,1884]" captionTargetPageId="14" captionText="Fig. 8. Zanna chartieri Constant sp. nov., nymph. A. Habitus dorsal view. B. Habitus, ventral view. C. Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837048" httpUri="https://zenodo.org/record/13837048/files/figure.png" pageId="19" pageNumber="20">Fig. 8AC</figureCitation>
). Pale grey. Tergites short and wide, with some small black spots in middle portion and curved black line on sides forming a zigzagged pattern not reaching apex; longitudinal ridge in lateral portion, not reaching anterior and posterior margins; groups of minute tubercles in posterolateral portion of visible segments. Sternites with some small black dots.
</paragraph>
</subSubSection>
<caption id="DF6E6664FFA8FFF12180F8285A4D1090" ID-DOI="http://doi.org/10.5281/zenodo.13837060" ID-Zenodo-Dep="13837060" httpUri="https://zenodo.org/record/13837060/files/figure.png" pageId="18" pageNumber="19" startId="18.[189,230,1926,1952]" targetBox="[212,1369,538,1888]" targetPageId="18">
<paragraph id="8BAE36ECFFA8FFF12180F8285A4D1090" blockId="18.[186,1399,1926,2023]" pageId="18" pageNumber="19">
<emphasis id="B965EAFEFFA8FFF12180F8285F7F10D6" bold="true" box="[189,277,1926,1952]" pageId="18" pageNumber="19">Fig. 11.</emphasis>
<emphasis id="B965EAFEFFA8FFF12027F8285FA710D6" box="[282,461,1926,1952]" italics="true" pageId="18" pageNumber="19">Zannachartieri</emphasis>
Constant
<taxonomicNameLabel id="A2565785FFA8FFF1237CF8295CF110D7" box="[577,667,1927,1953]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
, emerging adult in nature in Tatai, Cambodia, on
<taxonomicName id="4C114D6FFFA8FFF125FFF8285B1C10D7" box="[1218,1398,1926,1953]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="18" pageNumber="19" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAFEFFA8FFF125FFF8285B2410D6" box="[1218,1358,1926,1952]" italics="true" pageId="18" pageNumber="19">Desmodium</emphasis>
sp.
</taxonomicName>
(photographs © G. Chartier).
<emphasis id="B965EAFEFFA8FFF12325F8075C4710B5" bold="true" box="[536,557,1961,1987]" pageId="18" pageNumber="19">A</emphasis>
. Emerging adult with 4
<superScript id="7C649BA4FFA8FFF1227FF8065D2510C0" attach="left" box="[834,847,1960,1974]" fontSize="6" pageId="18" pageNumber="19">th</superScript>
and 5
<superScript id="7C649BA4FFA8FFF122ABF8065DC910C0" attach="left" box="[918,931,1960,1974]" fontSize="6" pageId="18" pageNumber="19">th</superScript>
instar nymphs, 18 Jan. 2017.
<emphasis id="B965EAFEFFA8FFF1243CF8075B7F10B5" bold="true" box="[1281,1301,1961,1987]" pageId="18" pageNumber="19">B</emphasis>
. 18 Jan. 2017.
<emphasis id="B965EAFEFFA8FFF1203BF8625F2B1090" bold="true" box="[262,321,1996,2022]" pageId="18" pageNumber="19">CH</emphasis>
. Time lapse of final moult of the same specimen, 18 Jan. 2017.
</paragraph>
</caption>
<subSubSection id="C30B6567FFA9FFF02180FE1A5B1C1339" pageId="19" pageNumber="20" type="biology_ecology">
<paragraph id="8BAE36ECFFA9FFF02180FE1A5F7616B8" blockId="19.[189,284,436,462]" box="[189,284,436,462]" pageId="19" pageNumber="20">
<heading id="D0E68180FFA9FFF02180FE1A5F7616B8" bold="true" box="[189,284,436,462]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B965EAFEFFA9FFF02180FE1A5F7616B8" bold="true" box="[189,284,436,462]" pageId="19" pageNumber="20">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA9FFF02180FE4D5D3D142B" blockId="19.[189,1399,482,861]" pageId="19" pageNumber="20">
In 2017, between 17 January and 25 March, many specimens, nymphs and adults, were observed in Tatai,
<collectingRegion id="49D5F80EFFA9FFF02180FDA85FC01556" box="[189,426,518,544]" country="Cambodia" name="Koh Kong" pageId="19" pageNumber="20">Koh Kong Province</collectingRegion>
,
<collectingCountry id="F306767CFFA9FFF02088FDA85C441556" box="[437,558,518,544]" name="Cambodia" pageId="19" pageNumber="20">Cambodia</collectingCountry>
(G. Chartier, pers. com.,
<date id="FFAF102CFFA9FFF0226AFDA85A671556" box="[855,1037,518,544]" pageId="19" pageNumber="20" value="2017-01" valueMax="2017-02-31" valueMin="2017-01-00">Jan.Feb. 2017</date>
), in a clearing covered in grass and other herbaceous plants at Rainbow Lodge. They were present in high density (
<figureCitation id="132A2A69FFA9FFF025B0FD875A811535" box="[1165,1259,553,579]" captionStart="Fig" captionStartId="15.[189,230,1896,1922]" captionTargetBox="[213,1370,268,1848]" captionTargetId="figure-62@15.[213,1370,268,1851]" captionTargetPageId="15" captionText="Fig. 9. Zanna chartieri Constant sp. nov., in nature in Tatai, Cambodia (photographs © G. Chartier). A. Adult on Desmodium sp., 20 Jan. 2017. B. Adult on unidentified plant, 17 Jan. 2017. C. Adult on Melastomasanguineum, lateral view, 17 Jan. 2017. D. Habitat, position of nymphs marked with red arrows, 20 Jan. 2017." figureDoi="http://doi.org/10.5281/zenodo.13837052" httpUri="https://zenodo.org/record/13837052/files/figure.png" pageId="19" pageNumber="20">Fig. 9D</figureCitation>
), with up to 40 nymphs at different stages, four emerging adults and
<specimenCount id="9D17FD65FFA9FFF02264FDE35DBD1510" box="[857,983,588,615]" count="2" pageId="19" pageNumber="20" type="adult">two adults</specimenCount>
, counted in an area aproximatively 1×
<quantity id="4CE99B09FFA9FFF021D9FDC15F4015FC" box="[228,298,623,650]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="19" pageNumber="20" unit="m" value="1.5">1.5 m</quantity>
on
<date id="FFAF102CFFA9FFF02061FDC15C4D15FF" box="[348,551,623,650]" pageId="19" pageNumber="20" value="2017-01-18">18 January 2017</date>
. The very cryptic nymphs were all feeding on an unidentified species of creeping
<taxonomicName id="4C114D6FFFA9FFF02070FD3C5FD015DA" box="[333,442,658,684]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
in the genus
<taxonomicName id="4C114D6FFFA9FFF02363FD3C5D5215DB" authority="Desv." authorityName="Desv." box="[606,824,658,685]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis id="B965EAFEFFA9FFF02363FD3C5C8015DA" box="[606,746,658,684]" italics="true" pageId="19" pageNumber="20">Desmodium</emphasis>
Desv.
</taxonomicName>
, leaving some characteristic white wax deposits on the stems they were sitting on (
<figureCitation id="132A2A69FFA9FFF02362FD185CD015A6" box="[607,698,694,720]" captionStart="Fig" captionStartId="16.[189,230,1894,1920]" captionTargetBox="[238,1351,274,1856]" captionTargetId="figure-84@16.[238,1351,274,1856]" captionTargetPageId="16" captionText="Fig. 10. Zanna chartieri Constant sp. nov., nymphs in nature in Tatai, Cambodia, on Desmodium sp. (photographs © G. Chartier). AB. 4th instar nymphs. A. Laterodorsal view, 18 Jan. 2017. B. Lateral view, 20 Jan. 2017. CD. 5th instar nymphs, 17 Jan. 2017. C. Lateral view. D. Dorsal view. EF. Groups of nymphs. E. On vertical twig, 20 Jan. 2017. F. On creeping twig close to ground, 19 Jan. 2019." figureDoi="http://doi.org/10.5281/zenodo.13837056" httpUri="https://zenodo.org/record/13837056/files/figure.png" pageId="19" pageNumber="20">Fig. 10</figureCitation>
). The last instar nymphs seemed to move to more vertical stems before their final moult (
<figureCitation id="132A2A69FFA9FFF02316FD775CDA1585" box="[555,688,729,755]" captionStart="Fig" captionStartId="15.[189,230,1896,1922]" captionTargetBox="[213,1370,268,1848]" captionTargetId="figure-62@15.[213,1370,268,1851]" captionTargetPageId="15" captionText="Fig. 9. Zanna chartieri Constant sp. nov., in nature in Tatai, Cambodia (photographs © G. Chartier). A. Adult on Desmodium sp., 20 Jan. 2017. B. Adult on unidentified plant, 17 Jan. 2017. C. Adult on Melastomasanguineum, lateral view, 17 Jan. 2017. D. Habitat, position of nymphs marked with red arrows, 20 Jan. 2017." figureDoi="http://doi.org/10.5281/zenodo.13837052" httpUri="https://zenodo.org/record/13837052/files/figure.png" pageId="19" pageNumber="20">Figs 9AB</figureCitation>
,
<figureCitation id="132A2A69FFA9FFF02380FD775CB11585" box="[701,731,729,755]" captionStart="Fig" captionStartId="18.[189,230,1926,1952]" captionTargetBox="[212,1369,538,1888]" captionTargetId="figure-156@18.[212,1369,538,1888]" captionTargetPageId="18" captionText="Fig. 11. Zannachartieri Constant sp. nov., emerging adult in nature in Tatai, Cambodia, onDesmodium sp. (photographs © G. Chartier). A. Emerging adult with 4th and 5th instar nymphs, 18 Jan. 2017. B. 18 Jan. 2017. CH. Time lapse of final moult of the same specimen, 18 Jan. 2017." figureDoi="http://doi.org/10.5281/zenodo.13837060" httpUri="https://zenodo.org/record/13837060/files/figure.png" pageId="19" pageNumber="20">11</figureCitation>
), even often climbing on different plant species like, for example, a newly emerged adult on the
<taxonomicName id="4C114D6FFFA9FFF023BAFD525ADD1460" authority=", Melastomasanguineum Sims" authorityName="Melastomasanguineum Sims" box="[647,1207,764,790]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="family">
Melastomataceae,
<emphasis id="B965EAFEFFA9FFF0225DFD525A1C1460" box="[864,1142,764,790]" italics="true" pageId="19" pageNumber="20">Melastomasanguineum</emphasis>
Sims
</taxonomicName>
(
<figureCitation id="132A2A69FFA9FFF025FBFD525B751460" box="[1222,1311,764,790]" captionStart="Fig" captionStartId="15.[189,230,1896,1922]" captionTargetBox="[213,1370,268,1848]" captionTargetId="figure-62@15.[213,1370,268,1851]" captionTargetPageId="15" captionText="Fig. 9. Zanna chartieri Constant sp. nov., in nature in Tatai, Cambodia (photographs © G. Chartier). A. Adult on Desmodium sp., 20 Jan. 2017. B. Adult on unidentified plant, 17 Jan. 2017. C. Adult on Melastomasanguineum, lateral view, 17 Jan. 2017. D. Habitat, position of nymphs marked with red arrows, 20 Jan. 2017." figureDoi="http://doi.org/10.5281/zenodo.13837052" httpUri="https://zenodo.org/record/13837052/files/figure.png" pageId="19" pageNumber="20">Fig. 9C</figureCitation>
) but no nymph was observed feeding on this plant. The adults seemed to move to other plants which were not found, and they were never seen feeding on
<taxonomicName id="4C114D6FFFA9FFF023F9FCEC5D3A142A" box="[708,848,834,860]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="genus">
<emphasis id="B965EAFEFFA9FFF023F9FCEC5D3A142A" box="[708,848,834,860]" italics="true" pageId="19" pageNumber="20">Desmodium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAE36ECFFA9FFF02180FC2B5B1C1339" blockId="19.[189,1399,900,1103]" pageId="19" pageNumber="20">
More specimens were recorded (photographed) by G.C., always at the same place in Tatai: on
<date id="FFAF102CFFA9FFF02180FC095FCB14B7" box="[189,417,935,961]" pageId="19" pageNumber="20" value="2012-11-01">
1
<superScript id="7C649BA4FFA9FFF021F1FC095EBD14C3" attach="left" box="[204,215,935,949]" fontSize="6" pageId="19" pageNumber="20">st</superScript>
November 2012
</date>
, an adult in a bathroom at night, presumably attracted to light; on
<date id="FFAF102CFFA9FFF02589FC095F1D1492" pageId="19" pageNumber="20" value="2014-02-20" valueMax="2014-03-25" valueMin="2014-02-20">20 February and 25 March 2014</date>
,
<specimenCount id="9D17FD65FFA9FFF020BEFC655F981492" box="[387,498,970,997]" count="1" pageId="19" pageNumber="20" type="adult">one adult</specimenCount>
each time in a house, the latter at night time, presumably attracted to light;
<date id="FFAF102CFFA9FFF02182FC405FE0137E" box="[191,394,1006,1032]" pageId="19" pageNumber="20" value="2017-08-02" valueMax="2017-08-04" valueMin="2017-08-02">24 August 2017</date>
, several nymphs on
<taxonomicName id="4C114D6FFFA9FFF02345FC435D45137E" box="[632,815,1005,1032]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAFEFFA9FFF02345FC435D6E1371" box="[632,772,1005,1031]" italics="true" pageId="19" pageNumber="20">Desmodium</emphasis>
sp.
</taxonomicName>
;
<date id="FFAF102CFFA9FFF02201FC405A7A137E" box="[828,1040,1005,1032]" pageId="19" pageNumber="20" value="2018-03-07" valueMax="2018-03-24" valueMin="2018-03-07">724 March 2018</date>
, several adults and nymphs on
<taxonomicName id="4C114D6FFFA9FFF02180FBBF5F1E135D" box="[189,372,1041,1067]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAFEFFA9FFF02180FBBF5F23135D" box="[189,329,1041,1067]" italics="true" pageId="19" pageNumber="20">Desmodium</emphasis>
sp.
</taxonomicName>
;
<date id="FFAF102CFFA9FFF020BCFBBF5C78135D" box="[385,530,1041,1067]" pageId="19" pageNumber="20" value="2018-06-01">1 June 2018</date>
,
<specimenCount id="9D17FD65FFA9FFF02320FBBF5CE0135D" box="[541,650,1041,1067]" count="1" pageId="19" pageNumber="20" type="adult">one adult</specimenCount>
on
<taxonomicName id="4C114D6FFFA9FFF02388FBBF5D09135D" box="[693,867,1041,1067]" class="Magnoliopsida" family="Melastomataceae" genus="Melastoma" kingdom="Plantae" order="Myrtales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAFEFFA9FFF02388FBBF5D5D135D" box="[693,823,1041,1067]" italics="true" pageId="19" pageNumber="20">Melastoma</emphasis>
sp.
</taxonomicName>
;
<date id="FFAF102CFFA9FFF0224DFBBF5A5F135D" box="[880,1077,1041,1067]" pageId="19" pageNumber="20" value="2019-03-02" valueMax="2019-03-04" valueMin="2019-03-02">24 March 2019</date>
, several adults and nymphs on
<taxonomicName id="4C114D6FFFA9FFF021DDFB9A5FFC1339" box="[224,406,1076,1103]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAFEFFA9FFF021DDFB9A5F061338" box="[224,364,1076,1102]" italics="true" pageId="19" pageNumber="20">Desmodium</emphasis>
sp.
</taxonomicName>
;
<date id="FFAF102CFFA9FFF0209FFB9A5C141339" box="[418,638,1076,1103]" pageId="19" pageNumber="20" value="2019-12-27">27 December 2019</date>
<date id="FFAF102CFFA9FFF023B3FB9A5D341338" box="[654,862,1076,1103]" pageId="19" pageNumber="20" value="2020-02-25">25 February 2020</date>
, several adults and nymphs on
<taxonomicName id="4C114D6FFFA9FFF025FEFB9A5B1C1339" box="[1219,1398,1076,1103]" class="Magnoliopsida" family="Fabaceae" genus="Desmodium" kingdom="Plantae" order="Fabales" pageId="19" pageNumber="20" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B965EAFEFFA9FFF025FEFB9A5B251338" box="[1219,1359,1076,1102]" italics="true" pageId="19" pageNumber="20">Desmodium</emphasis>
sp.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFA9FFF02180FBDB5CFD13C9" box="[189,663,1141,1215]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BAE36ECFFA9FFF02180FBDB5F3C13F9" blockId="19.[189,342,1141,1167]" box="[189,342,1141,1167]" pageId="19" pageNumber="20">
<heading id="D0E68180FFA9FFF02180FBDB5F3C13F9" bold="true" box="[189,342,1141,1167]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="B965EAFEFFA9FFF02180FBDB5F3C13F9" bold="true" box="[189,342,1141,1167]" pageId="19" pageNumber="20">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFA9FFF02180FB0A5CFD13C9" blockId="19.[189,663,1188,1215]" box="[189,663,1188,1215]" pageId="19" pageNumber="20">
<collectingCountry id="F306767CFFA9FFF02180FB0A5F5013C8" box="[189,314,1188,1214]" name="Cambodia" pageId="19" pageNumber="20">Cambodia</collectingCountry>
:
<collectingRegion id="49D5F80EFFA9FFF02078FB0A5C4513C8" box="[325,559,1188,1214]" country="Cambodia" name="Koh Kong" pageId="19" pageNumber="20">Koh Kong Province</collectingRegion>
(
<figureCitation id="132A2A69FFA9FFF0237DFB0A5CE213C9" box="[576,648,1188,1215]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="19" pageNumber="20">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,529 @@
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<mods:title id="35F0EEE866E8804EAA3A42B5132645C0">Four new species of the lanternfly genus Zanna Kirkaldy, 1902 from Cambodia and Vietnam (Hemiptera: Fulgoromorpha: Fulgoridae)</mods:title>
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<mods:namePart id="BC69EFA40395439E113D46B9C64FCD41">Constant, Jérôme</mods:namePart>
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<mods:affiliation id="0AB27199D31DCAD7FFDDDE1B7B98F2E7">Mientrung Institute for Scientific Research, Vietnam National Museum of Nature, VAST, 321 Huynh Thuc Khang, Hue, Vietnam. &amp; Graduate School of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam.</mods:affiliation>
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<paragraph id="8BAE36ECFFBEFFE72340FB5E5DDD127C" blockId="4.[368,1219,1263,1361]" box="[637,951,1263,1290]" pageId="4" pageNumber="5">
<heading id="D0E68180FFBEFFE72340FB5E5DDD127C" box="[637,951,1263,1290]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
<taxonomicName id="4C114D6FFFBEFFE72340FB5E5D3F127C" authorityName="Constant &amp; Pham" authorityYear="2024" box="[637,853,1263,1290]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE72340FB5E5D3F127C" bold="true" box="[637,853,1263,1290]" italics="true" pageId="4" pageNumber="5">Zanna bidoupana</emphasis>
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<taxonomicNameLabel id="A2565785FFBEFFE72261FB5E5DDD127C" box="[860,951,1264,1290]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C30B6567FFBEFFE72345FA985DD61226" box="[632,956,1334,1361]" pageId="4" pageNumber="5" type="description">
<paragraph id="8BAE36ECFFBEFFE72345FA985DD61226" blockId="4.[368,1219,1263,1361]" box="[632,956,1334,1361]" pageId="4" pageNumber="5">
<figureCitation id="132A2A69FFBEFFE72345FA985C8A1227" box="[632,736,1334,1361]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="6.[189,230,1928,1954]" captionStartId-1="7.[190,230,1927,1953]" captionStartId-2="8.[189,230,1909,1935]" captionStartId-3="9.[188,229,1994,2020]" captionTargetBox-0="[217,1369,263,1889]" captionTargetBox-1="[216,1346,273,1889]" captionTargetBox-2="[257,1349,265,1872]" captionTargetBox-3="[215,1372,718,1954]" captionTargetId-0="figure-46@6.[212,1369,258,1890]" captionTargetId-1="figure-62@7.[212,1369,260,1892]" captionTargetId-2="figure-45@8.[257,1349,265,1872]" captionTargetId-3="figure-180@9.[215,1372,718,1954]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionText-0="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." captionText-1="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." captionText-2="Fig. 4. AB. Zanna bidoupana sp. nov. at the light trap, Vietnam, Bidoup-Nui Ba National Park, 23 Jul. 2014. A. Dorsal view. B. Lateral view. C. Habitat of Z.bidoupana sp. nov. around Hon Giao station, 23 Jul. 2014." captionText-3="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi-0="http://doi.org/10.5281/zenodo.13837032" figureDoi-1="http://doi.org/10.5281/zenodo.13837034" figureDoi-2="http://doi.org/10.5281/zenodo.13837036" figureDoi-3="http://doi.org/10.5281/zenodo.13837040" httpUri-0="https://zenodo.org/record/13837032/files/figure.png" httpUri-1="https://zenodo.org/record/13837034/files/figure.png" httpUri-2="https://zenodo.org/record/13837036/files/figure.png" httpUri-3="https://zenodo.org/record/13837040/files/figure.png" pageId="4" pageNumber="5">Figs 25</figureCitation>
,
<figureCitation id="132A2A69FFBEFFE723D1FA985D301227" box="[748,858,1334,1361]" captionStart="Fig" captionStartId="29.[191,234,1880,1906]" captionTargetBox="[185,1406,518,1841]" captionTargetId="figure-165@29.[185,1406,515,1845]" captionTargetPageId="29" captionText="Fig. 18. Zanna, new spp. from Cambodia and Vietnam, key characters. AD. Prothotax and basal half of head, lateral view. EH. Head and thorax, dorsal view. IL. Right tegmen. A, E, I. Z. bidoupana sp. nov. B, F, J. Z. chartieri Constant sp. nov. C, G, K. Z. kusamae sp. nov. D, H, L. Z. limbourgi Constant sp. nov., (not to scale)." figureDoi="http://doi.org/10.5281/zenodo.13837080" httpUri="https://zenodo.org/record/13837080/files/figure.png" pageId="4" pageNumber="5">18A, E, I</figureCitation>
,
<figureCitation id="132A2A69FFBEFFE7225AFA985DD61226" box="[871,956,1334,1360]" captionStart="Fig" captionStartId="30.[193,236,1892,1918]" captionTargetBox="[193,1402,242,1853]" captionTargetId="figure-68@30.[189,1410,242,1853]" captionTargetPageId="30" captionText="Fig. 19. Zanna, new spp. from Cambodia and Vietnam, key characters of pygofer, anal tube and gonostyli. A, C, E, G. Lateral view. B, D, F, H. Dorsal view. AB. Z. bidoupana sp. nov. CD. Z. chartieri Constant sp. nov. EF. Z. kusamae sp. nov. GH. Z. limbourgi Constant sp. nov. (scale bars =1 mm). Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837082" httpUri="https://zenodo.org/record/13837082/files/figure.png" pageId="4" pageNumber="5">19AB</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBEFFE72180FAD95D2A104E" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BAE36ECFFBEFFE72180FAD95F5C12E7" blockId="4.[189,1406,1399,1849]" box="[189,310,1399,1425]" pageId="4" pageNumber="5">
<heading id="D0E68180FFBEFFE72180FAD95F5C12E7" bold="true" box="[189,310,1399,1425]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B965EAFEFFBEFFE72180FAD95F5C12E7" bold="true" box="[189,310,1399,1425]" pageId="4" pageNumber="5">Diagnosis</emphasis>
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</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180FA355D2A104E" blockId="4.[189,1406,1399,1849]" pageId="4" pageNumber="5">
The species is closest to
<taxonomicName id="4C114D6FFFBEFFE720EFFA325C9312C3" authority="Constant" authorityName="Constant" authorityYear="2024" box="[466,761,1435,1461]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chartieri" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE720EFFA325CE012C3" box="[466,650,1435,1461]" italics="true" pageId="4" pageNumber="5">Zanna chartieri</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE723C3FA355D3312C3" box="[766,857,1435,1461]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C114D6FFFBEFFE7225CFA325D8F12C3" authorityName="Constant &amp; Pham" authorityYear="2024" box="[865,997,1435,1461]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE7225CFA325D8F12C3" box="[865,997,1435,1461]" italics="true" pageId="4" pageNumber="5">Z. kusamae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE722D1FA355A2F12C3" box="[1004,1093,1435,1461]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C114D6FFFBEFFE7254AFA325B1412C3" authority="Constant" authorityName="Constant" authorityYear="2024" box="[1143,1406,1435,1461]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="limbourgi" status="sp. nov.">
<emphasis id="B965EAFEFFBEFFE7254AFA325B6312C3" box="[1143,1289,1435,1461]" italics="true" pageId="4" pageNumber="5">Z.limbourgi</emphasis>
Constant
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBEFFE72180FA115F7312AF" box="[189,281,1471,1497]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, from which it can be separated by the following characters: basal portion of head (
<figureCitation id="132A2A69FFBEFFE725EAFA105B0512AE" box="[1239,1391,1470,1496]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="4" pageNumber="5">Fig. 2B, EF</figureCitation>
) covered in dense, irregular, rather large, coarse, sometimes coalescent black pitting with few smaller pits (moderately large, rather well-spaced, black pitting in
<taxonomicName id="4C114D6FFFBEFFE7224EF9AB5D931168" authorityName="Constant &amp; Pham" authorityYear="2024" box="[883,1017,1540,1566]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFBEFFE7224EF9AB5D931168" box="[883,1017,1540,1566]" italics="true" pageId="4" pageNumber="5">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE72525F9AA5AA91169" box="[1048,1219,1540,1567]" captionStart="Fig" captionStartId="21.[188,229,1927,1953]" captionTargetBox="[215,1367,257,1885]" captionTargetId="figure-46@21.[210,1367,254,1885]" captionTargetPageId="21" captionText="Fig. 12. Zanna kusamae sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837062" httpUri="https://zenodo.org/record/13837062/files/figure.png" pageId="4" pageNumber="5">Fig. 12B, EF</figureCitation>
; same but with background of dense, very small black pitting in
<taxonomicName id="4C114D6FFFBEFFE72226F9865DC91134" authorityName="Constant" authorityYear="2024" box="[795,931,1576,1602]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chartieri">
<emphasis id="B965EAFEFFBEFFE72226F9865DC91134" box="[795,931,1576,1602]" italics="true" pageId="4" pageNumber="5">Z. chartieri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFBEFFE722E2F9865A191134" authorityName="Constant" authorityYear="2024" box="[991,1139,1576,1602]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFBEFFE722E2F9865A191134" box="[991,1139,1576,1602]" italics="true" pageId="4" pageNumber="5">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE725ACF9865B5B1134" box="[1169,1329,1575,1602]" captionStart="Fig" captionStartId="12.[185,226,1930,1956]" captionTargetBox="[252,1334,322,1889]" captionTargetId="figure-47@12.[242,1334,317,1889]" captionTargetPageId="12" captionText="Fig. 6. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837042" httpUri="https://zenodo.org/record/13837042/files/figure.png" pageId="4" pageNumber="5">Figs 6B, EF</figureCitation>
,
<figureCitation id="132A2A69FFBEFFE72403F9895E991113" captionStart="Fig" captionStartId="25.[189,230,1925,1951]" captionTargetBox="[240,1354,270,1868]" captionTargetId="figure-46@25.[230,1354,265,1868]" captionTargetPageId="25" captionText="Fig. 15. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS). A. Habitus dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837073" httpUri="https://zenodo.org/record/13837073/files/figure.png" pageId="4" pageNumber="5">15B, EF</figureCitation>
); anal tube of male rather high in lateral view (
<figureCitation id="132A2A69FFBEFFE7220CF9E55DFB1113" box="[817,913,1611,1637]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="4" pageNumber="5">Fig. 3A</figureCitation>
), about 1.60 times as long as high, with ventral angle in basal half (rather flattened dorsoventrally, 1.952.14 times as long as high in lateral view, with ventral angle at midlength in
<taxonomicName id="4C114D6FFFBEFFE723A6F93C5D4B11DD" authorityName="Constant" authorityYear="2024" box="[667,801,1681,1707]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chartieri">
<emphasis id="B965EAFEFFBEFFE723A6F93C5D4B11DD" box="[667,801,1681,1707]" italics="true" pageId="4" pageNumber="5">Z. chartieri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D6FFFBEFFE72261F93C5D8811DD" authorityName="Constant &amp; Pham" authorityYear="2024" box="[860,994,1681,1707]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFBEFFE72261F93C5D8811DD" box="[860,994,1681,1707]" italics="true" pageId="4" pageNumber="5">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE7253DF93F5A0B11DD" box="[1024,1121,1681,1707]" captionStart="Fig" captionStartId="13.[189,228,1925,1951]" captionTargetBox="[232,1368,257,1882]" captionTargetId="figure-62@13.[220,1377,238,1886]" captionTargetPageId="13" captionText="Fig. 7. Zanna chartieri Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view.F. Dorsal view.G. Laterodorsal view.H. Ventral view.Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837046" httpUri="https://zenodo.org/record/13837046/files/figure.png" pageId="4" pageNumber="5">Figs 7A</figureCitation>
,
<figureCitation id="132A2A69FFBEFFE72553F93F5ACE11DD" box="[1134,1188,1681,1707]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="4" pageNumber="5">13A</figureCitation>
); pygofer of male in lateral view (
<figureCitation id="132A2A69FFBEFFE72047F91A5FBD11B9" box="[378,471,1716,1743]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="4" pageNumber="5">Fig. 3A</figureCitation>
) with posterior margin sinuate, forming large, broadly rounded posterior lobe in dorsal half (posterior margin projecting into large posterior lobe angularly rounded in midheight in
<taxonomicName id="4C114D6FFFBEFFE72182F9525F291063" authorityName="Constant &amp; Pham" authorityYear="2024" box="[191,323,1787,1813]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kusamae">
<emphasis id="B965EAFEFFBEFFE72182F9525F291063" box="[191,323,1787,1813]" italics="true" pageId="4" pageNumber="5">Z. kusamae</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE72061F9555FAD1063" box="[348,455,1787,1813]" captionStart="Fig" captionStartId="22.[189,230,1926,1952]" captionTargetBox="[218,1365,253,1887]" captionTargetId="figure-62@22.[214,1371,238,1891]" captionTargetPageId="22" captionText="Fig. 13. Zannakusamae sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tubeand gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Materials and methods." figureDoi="http://doi.org/10.5281/zenodo.13837066" httpUri="https://zenodo.org/record/13837066/files/figure.png" pageId="4" pageNumber="5">Fig. 13A</figureCitation>
; posterior margin projecting posteriad in dorsal ⅓, in large, apically rounded lobe forming nearly right angle in
<taxonomicName id="4C114D6FFFBEFFE7232AF8B15CC3104E" authorityName="Constant" authorityYear="2024" box="[535,681,1822,1848]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="limbourgi">
<emphasis id="B965EAFEFFBEFFE7232AF8B15CC3104E" box="[535,681,1822,1848]" italics="true" pageId="4" pageNumber="5">Z. limbourgi</emphasis>
</taxonomicName>
<figureCitation id="132A2A69FFBEFFE723FBF8B05D58104E" box="[710,818,1822,1848]" captionStart="Fig" captionStartId="26.[197,236,1958,1984]" captionTargetBox="[230,1373,285,1917]" captionTargetId="figure-62@26.[226,1383,268,1920]" captionTargetPageId="26" captionText="Fig. 16. Zanna limbourgi Constant sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus. E. Left" figureDoi="http://doi.org/10.5281/zenodo.13837076" httpUri="https://zenodo.org/record/13837076/files/figure.png" pageId="4" pageNumber="5">Fig. 16A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBEFFE72180F8CE5FC010BD" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BAE36ECFFBEFFE72180F8CE5F2E100C" blockId="4.[189,324,1888,1914]" box="[189,324,1888,1914]" pageId="4" pageNumber="5">
<heading id="D0E68180FFBEFFE72180F8CE5F2E100C" bold="true" box="[189,324,1888,1914]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="B965EAFEFFBEFFE72180F8CE5F2E100C" bold="true" box="[189,324,1888,1914]" pageId="4" pageNumber="5">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFBEFFE72180F8205FC010BD" blockId="4.[189,1398,1934,1995]" pageId="4" pageNumber="5">
The species epithet refers to the
<typeStatus id="54AA884EFFBEFFE72378F8215C1210DF" box="[581,632,1935,1961]" pageId="4" pageNumber="5">type</typeStatus>
location, Bidoup-Nui Ba National Park in
<collectingRegion id="49D5F80EFFBEFFE72540F8215B1810DE" box="[1149,1394,1934,1961]" country="Vietnam" name="Lam Dong" pageId="4" pageNumber="5">Lam Dong Province</collectingRegion>
, Highlands
<collectingCountry id="F306767CFFBEFFE72003F81F5FCF10BD" box="[318,421,1969,1995]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBFFFE62180FEA75F741460" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BAE36ECFFBFFFE62180FEA75F061654" blockId="5.[189,364,264,291]" box="[189,364,264,291]" pageId="5" pageNumber="6">
<heading id="D0E68180FFBFFFE62180FEA75F061654" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B965EAFEFFBFFFE62180FEA75F061654" bold="true" box="[189,364,264,291]" pageId="5" pageNumber="6">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFBFFFE621D7FE995F361627" blockId="5.[189,1399,311,478]" box="[234,348,311,337]" pageId="5" pageNumber="6">
<emphasis id="B965EAFEFFBFFFE621D7FE995F361627" bold="true" box="[234,348,311,337]" pageId="5" pageNumber="6">
<typeStatus id="54AA884EFFBFFFE621D7FE995F361627" box="[234,348,311,337]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FEF45A3916A8" blockId="5.[189,1399,311,478]" pageId="5" pageNumber="6">
<materialsCitation id="3B793CB1FFBFFFE62180FEF45A3916A8" collectingDate="2014-07-21" collectingDateMax="2014-07-25" collectingDateMin="2014-07-21" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc. &amp; Leg. J. Constant" country="Vietnam" latitude="12.186611" location="Hon Giao station" longLatPrecision="1" longitude="108.714806" municipality="Bidoup-Nui Ba National Park" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong" typeStatus="holotype">
<collectingCountry id="F306767CFFBFFFE62180FEF45F2E1602" box="[189,324,346,372]" name="Vietnam" pageId="5" pageNumber="6">VIETNAM</collectingCountry>
<specimenCount id="9D17FD65FFBFFFE62059FEF45FFC1603" box="[356,406,346,373]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D5F80EFFBFFFE6209BFEF55CF71602" box="[422,669,346,373]" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="6">Lam Dong Province</collectingRegion>
,
<collectingMunicipality id="6BCAAC96FFBFFFE62391FEF45A7D1602" box="[684,1047,346,373]" pageId="5" pageNumber="6">Bidoup-Nui Ba National Park</collectingMunicipality>
,
<location id="8ECE6037FFBFFFE62518FEF55A921602" LSID="urn:lsid:plazi:treatment:03B887FAFFBEFFE92340FB5E5CEF130E:8ECE6037FFBFFFE62518FEF55A921602" box="[1061,1272,346,373]" country="Vietnam" latitude="12.186611" longLatPrecision="1" longitude="108.714806" municipality="Bidoup-Nui Ba National Park" name="Hon Giao station" pageId="5" pageNumber="6" stateProvince="Lam Dong">Hon Giao station</location>
;
<geoCoordinate id="EE25502BFFBFFFE6243AFEF75E9F16EE" degrees="12" direction="north" minutes="11" orientation="latitude" pageId="5" pageNumber="6" precision="1" seconds="11. 8" value="12.186611">12°11ʹ11. 8ʺ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFBFFFE6203DFED25FDE16EE" box="[256,436,380,408]" degrees="108" direction="east" minutes="42" orientation="longitude" pageId="5" pageNumber="6" precision="1" seconds="53.3" value="108.714806">108°42ʹ53.3ʺ E</geoCoordinate>
;
<date id="FFAF102CFFBFFFE620FDFED35C1716EE" box="[448,637,381,408]" pageId="5" pageNumber="6" value="2014-07-21" valueMax="2014-07-25" valueMin="2014-07-21">
<collectingDate id="EFEBE9C4FFBFFFE620FDFED35C1716EE" box="[448,637,381,408]" pageId="5" pageNumber="6" value="2014-07-21" valueMax="2014-07-25" valueMin="2014-07-21">2125 Jul. 2014</collectingDate>
</date>
;
<collectorName id="26E4533AFFBFFFE623B5FED05D6016EE" box="[648,778,382,408]" pageId="5" pageNumber="6">J. Constant</collectorName>
and
<collectorName id="26E4533AFFBFFFE6227CFED05DD716E1" box="[833,957,381,408]" pageId="5" pageNumber="6">J. Bresseel</collectorName>
leg.; GTI project; I.G.: 32.779; “
<collectorName id="26E4533AFFBFFFE62406FED35F2D16CD" pageId="5" pageNumber="6">Coll. I.R.Sc.N.B.</collectorName>
, Vietnam,
<collectingRegion id="49D5F80EFFBFFFE620F0FE0F5CF216CD" box="[461,664,417,443]" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="6">Lam Dong prov.</collectingRegion>
, Bidoup-Nui Ba N.P.,
<geoCoordinate id="EE25502BFFBFFFE62293FE315A7416CD" box="[942,1054,415,443]" degrees="12" direction="north" minutes="26" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="12.433333">12°26ʹ N</geoCoordinate>
,
<geoCoordinate id="EE25502BFFBFFFE62513FE315AC216CD" box="[1070,1192,415,443]" degrees="108" direction="east" minutes="30" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="108.5">108°30ʹ E</geoCoordinate>
,
<date id="FFAF102CFFBFFFE62584FE0E5B1816CD" box="[1209,1394,416,443]" pageId="5" pageNumber="6" value="2014-07-21" valueMax="2014-07-25" valueMin="2014-07-21">
<collectingDate id="EFEBE9C4FFBFFFE62584FE0E5B1816CD" box="[1209,1394,416,443]" pageId="5" pageNumber="6" value="2014-07-21" valueMax="2014-07-25" valueMin="2014-07-21">21-25.VII.2014</collectingDate>
</date>
,
<collectingMethod id="52504EFBFFBFFFE62180FE6A5F5F16A8" box="[189,309,452,478]" pageId="5" pageNumber="6">Light trap</collectingMethod>
,
<collectorName id="26E4533AFFBFFFE6207DFE6A5F9516A8" box="[320,511,452,478]" pageId="5" pageNumber="6">Leg. J. Constant</collectorName>
&amp;
<collectorName id="26E4533AFFBFFFE62319FE6A5CCE16A8" box="[548,676,452,478]" pageId="5" pageNumber="6">J. Bresseel</collectorName>
, GTI Project, I.G.: 32.779”;
<collectionCode id="ED00AE29FFBFFFE622C8FE6A5A2516A8" box="[1013,1103,452,478]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="5" pageNumber="6" type="Museum">RBINS</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BAE36ECFFBFFFE621D7FDAB5C4E1569" blockId="5.[189,1399,517,790]" box="[234,548,517,544]" pageId="5" pageNumber="6">
<emphasis id="B965EAFEFFBFFFE621D7FDAB5F0D1569" bold="true" box="[234,359,517,543]" pageId="5" pageNumber="6">
<typeStatus id="54AA884EFFBFFFE621D7FDAB5F0D1569" box="[234,359,517,543]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D17FD65FFBFFFE62045FDAB5FA61556" box="[376,460,517,544]" count="12" pageId="5" pageNumber="6" type="male">12 ♂♂</specimenCount>
,
<specimenCount id="9D17FD65FFBFFFE620E5FDAB5C721569" box="[472,536,517,543]" count="2" pageId="5" pageNumber="6" type="female">2 ♀♀</specimenCount>
)
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FD875F741460" blockId="5.[189,1399,517,790]" pageId="5" pageNumber="6">
<materialsCitation id="3B793CB1FFBFFFE62180FD875A7D1535" box="[189,1047,552,579]" collectingDate="2014-07-21" collectingDateMax="2014-07-25" collectingDateMin="2014-07-21" collectionCode="RBINS" collectorName="J. Constant &amp; J. Bresseel &amp; Coll. I. R. Sc. N. B. &amp; Coll. I. R. Sc. &amp; Leg. J. Constant" country="Vietnam" latitude="12.186611" location="Hon Giao station" longLatPrecision="1" longitude="108.714806" municipality="Bidoup-Nui Ba National Park" pageId="5" pageNumber="6" specimenCount="10" specimenCount-female="2" specimenCount-male="8" stateProvince="Lam Dong">
<collectingCountry id="F306767CFFBFFFE62180FD875F2E1535" box="[189,324,553,579]" name="Vietnam" pageId="5" pageNumber="6">VIETNAM</collectingCountry>
<specimenCount id="9D17FD65FFBFFFE62058FD875FC01535" box="[357,426,553,579]" count="8" pageId="5" pageNumber="6" type="male">8 ♂♂</specimenCount>
,
<specimenCount id="9D17FD65FFBFFFE6208AFD875F901535" box="[439,506,553,579]" count="2" pageId="5" pageNumber="6" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED00AE29FFBFFFE622FCFD875A7D1535" box="[961,1047,553,579]" country="Belgium" httpUri="http://biocol.org/urn:lsid:biocol.org:col:35271" lsid="urn:lsid:biocol.org:col:35271" name="Royal Belgian Institute of Natural Sciences" pageId="5" pageNumber="6" type="Museum">RBINS</collectionCode>
</materialsCitation>
<materialsCitation id="3B793CB1FFBFFFE6250CFD875C5A15FF" collectingDate="2013-06-04" collectionCode="VNMN" collectorName="H. T. Pham" country="Vietnam" location="Bidoup-Nui Ba National Park" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong">
<specimenCount id="9D17FD65FFBFFFE6250CFD875A0A1535" box="[1073,1120,553,579]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49D5F80EFFBFFFE62550FD875A811535" box="[1133,1259,553,579]" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="6">Lam Dong</collectingRegion>
[Province],
<location id="8ECE6037FFBFFFE62180FDE25C491510" LSID="urn:lsid:plazi:treatment:03B887FAFFBEFFE92340FB5E5CEF130E:8ECE6037FFBFFFE62180FDE25C491510" box="[189,547,588,614]" country="Vietnam" name="Bidoup-Nui Ba National Park" pageId="5" pageNumber="6" stateProvince="Lam Dong">Bidoup-Nui Ba National Park</location>
;
<date id="FFAF102CFFBFFFE6230DFDE25CD41510" box="[560,702,588,614]" pageId="5" pageNumber="6" value="2013-06-04">
<collectingDate id="EFEBE9C4FFBFFFE6230DFDE25CD41510" box="[560,702,588,614]" pageId="5" pageNumber="6" value="2013-06-04">4 Jun. 2013</collectingDate>
</date>
; [
<collectingMethod id="52504EFBFFBFFFE623EEFDE25D221510" box="[723,840,588,614]" pageId="5" pageNumber="6">light trap</collectingMethod>
]; [
<collectorName id="26E4533AFFBFFFE62259FDE25D8C1510" box="[868,998,588,614]" pageId="5" pageNumber="6">H.T. Pham</collectorName>
leg.]; “VQG Bidup-Nui Ba,
<collectingRegion id="49D5F80EFFBFFFE6247CFDE25F6B15FF" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="6">Lam Dong</collectingRegion>
,
<date id="FFAF102CFFBFFFE6202CFDC15F1615FF" box="[273,380,623,649]" pageId="5" pageNumber="6" value="2013-06-04">
<collectingDate id="EFEBE9C4FFBFFFE6202CFDC15F1615FF" box="[273,380,623,649]" pageId="5" pageNumber="6" value="2013-06-04">4/6/2013</collectingDate>
</date>
, VD”;
<collectionCode id="ED00AE29FFBFFFE620EBFDC15C5A15FF" box="[470,560,623,649]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
</materialsCitation>
<materialsCitation id="3B793CB1FFBFFFE6236CFDC15FA415A6" collectingDate="2016-10-13" collectionCode="VNMN" collectorName="T. T. Men" country="Vietnam" location="Bidoup National Park" pageId="5" pageNumber="6" specimenCode="E 000009331" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong">
<specimenCount id="9D17FD65FFBFFFE6236CFDC15CEE15FF" box="[593,644,623,649]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
; [
<collectingRegion id="49D5F80EFFBFFFE623A1FDC15DF115FF" box="[668,923,623,649]" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="6">Lam Dong Province</collectingRegion>
],
<location id="8ECE6037FFBFFFE6228FFDC15AD415FF" LSID="urn:lsid:plazi:treatment:03B887FAFFBEFFE92340FB5E5CEF130E:8ECE6037FFBFFFE6228FFDC15AD415FF" box="[946,1214,623,649]" country="Vietnam" name="Bidoup National Park" pageId="5" pageNumber="6" stateProvince="Lam Dong">Bidoup National Park</location>
;
<date id="FFAF102CFFBFFFE625F3FDC15B1815FF" box="[1230,1394,623,649]" pageId="5" pageNumber="6" value="2016-10-13">
<collectingDate id="EFEBE9C4FFBFFFE625F3FDC15B1815FF" box="[1230,1394,623,649]" pageId="5" pageNumber="6" value="2016-10-13">13 Oct. 2016</collectingDate>
</date>
;
<collectorName id="26E4533AFFBFFFE62180FD3D5F4315DB" box="[189,297,659,685]" pageId="5" pageNumber="6">T.T. Men</collectorName>
leg.;
<collectionCode id="ED00AE29FFBFFFE62050FD3D5FAD15DB" box="[365,455,659,685]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
_
<specimenCode id="DBB79E97FFBFFFE620EBFD3D5C1D15DA" box="[470,631,658,685]" collectionCode="VNMN" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">E 000009331</specimenCode>
; “Bidoup NP, LD,
<date id="FFAF102CFFBFFFE62265FD3C5D8B15DA" box="[856,993,658,684]" pageId="5" pageNumber="6" value="2016-10-13">
<collectingDate id="EFEBE9C4FFBFFFE62265FD3C5D8B15DA" box="[856,993,658,684]" pageId="5" pageNumber="6" value="2016-10-13">13/10/2016</collectingDate>
</date>
, light,
<collectorName id="26E4533AFFBFFFE6250EFD3D5AF515DB" box="[1075,1183,659,685]" pageId="5" pageNumber="6">T.T. Men</collectorName>
leg.”, “
<collectionCode id="ED00AE29FFBFFFE625C4FD3C5B3F15DA" box="[1273,1365,658,684]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
_E 000 009 331”;
<collectionCode id="ED00AE29FFBFFFE62049FD185FA415A6" box="[372,462,694,720]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
</materialsCitation>
<materialsCitation id="3B793CB1FFBFFFE620D0FD1B5B1D15A6" box="[493,1399,693,720]" collectingDate="2016-10-13" collectionCode="VNMN" collectorName="T. T. Men" country="Vietnam" location="Bidoup National Park" pageId="5" pageNumber="6" specimenCode="E 000009330" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong">
<specimenCount id="9D17FD65FFBFFFE620D0FD1B5C7515A6" box="[493,543,693,720]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectionCode id="ED00AE29FFBFFFE6253FFD185A3615A6" box="[1026,1116,694,720]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
_
<specimenCode id="DBB79E97FFBFFFE62556FD185B6515B9" box="[1131,1295,693,720]" collectionCode="VNMN" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">E 000009330</specimenCode>
;
<collectionCode id="ED00AE29FFBFFFE62420FD185B1D15A6" box="[1309,1399,694,720]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
</materialsCitation>
<materialsCitation id="3B793CB1FFBFFFE621F3FD775F711460" collectingDate="2016-10-13" collectionCode="VNMN" collectorName="T. T. Men &amp; N. V. Dat" country="Vietnam" location="Bidoup National Park" pageId="5" pageNumber="6" specimenCode="E 000009332" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong">
<specimenCount id="9D17FD65FFBFFFE621F3FD775E971585" box="[206,253,729,755]" count="1" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
; same collection data as for preceding;
<collectingMethod id="52504EFBFFBFFFE623F4FD765D7E1585" box="[713,788,728,755]" pageId="5" pageNumber="6">by net</collectingMethod>
;
<collectorName id="26E4533AFFBFFFE62222FD775DE01585" box="[799,906,729,755]" pageId="5" pageNumber="6">T.T. Men</collectorName>
and
<collectorName id="26E4533AFFBFFFE622FFFD775A421585" box="[962,1064,729,755]" pageId="5" pageNumber="6">N.V. Dat</collectorName>
leg.;
<collectionCode id="ED00AE29FFBFFFE62554FD775AA91585" box="[1129,1219,729,755]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
_
<specimenCode id="DBB79E97FFBFFFE625EFFD775B181585" box="[1234,1394,729,755]" collectionCode="VNMN" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">E 000009332</specimenCode>
;
<collectionCode id="ED00AE29FFBFFFE62180FD525F711460" box="[189,283,764,790]" country="Viet Nam" httpUri="http://grbio.org/cool/65ma-2bzq" name="Vietnam National Museum of Nature" pageId="5" pageNumber="6">VNMN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFBFFFE92180FC935B191402" lastPageId="10" lastPageNumber="11" pageId="5" pageNumber="6" type="description">
<paragraph id="8BAE36ECFFBFFFE62180FC935F241421" blockId="5.[189,334,829,855]" box="[189,334,829,855]" pageId="5" pageNumber="6">
<heading id="D0E68180FFBFFFE62180FC935F241421" bold="true" box="[189,334,829,855]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B965EAFEFFBFFFE62180FC935F241421" bold="true" box="[189,334,829,855]" pageId="5" pageNumber="6">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FCC25CAB14BB" blockId="5.[189,1399,876,973]" pageId="5" pageNumber="6">
MEASUREMENTS AND RATIOS. LT:
<specimenCount id="9D17FD65FFBFFFE62369FCC25C0014F0" box="[596,618,876,902]" count="1" pageId="5" pageNumber="6" type="male"></specimenCount>
(n =10): 41.0 mm (38.243.1),
<specimenCount id="9D17FD65FFBFFFE622C7FCC25A7A14F0" box="[1018,1040,876,902]" count="1" pageId="5" pageNumber="6" type="female"></specimenCount>
(n=2): 41.942.0 mm. LTg/ BTg =3.48; LH/BH=4.0; LF/BF =5.3; BH/BPrH = 2.2; LH/LT =0.35; wingspan (extrapolated):
<specimenCount id="9D17FD65FFBFFFE62180FC1D5EBD14BB" box="[189,215,947,973]" count="1" pageId="5" pageNumber="6" type="male"></specimenCount>
:
<quantity id="4CE99B09FFBFFFE621DFFC1C5F2114BB" box="[226,331,946,973]" metricMagnitude="-2" metricUnit="m" metricValue="5.279999999999999" pageId="5" pageNumber="6" unit="mm" value="52.8">52.8 mm</quantity>
(49.255.5),
<specimenCount id="9D17FD65FFBFFFE620D7FC1D5C6814BB" box="[490,514,947,973]" count="1" pageId="5" pageNumber="6" type="female"></specimenCount>
: 54.0
<quantity id="4CE99B09FFBFFFE6236EFC1C5CD714BB" box="[595,701,946,973]" metricMagnitude="-2" metricUnit="m" metricValue="5.41" pageId="5" pageNumber="6" unit="mm" value="54.1">54.1 mm</quantity>
.
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FC5A5F2D125E" blockId="5.[189,1399,1011,1320]" pageId="5" pageNumber="6">
HEAD (
<figureCitation id="132A2A69FFBFFFE62032FC5D5FC71378" box="[271,429,1011,1038]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="5" pageNumber="6">Fig. 2B, EF</figureCitation>
). Strongly elongate, representing 35% of total length, gently and evenly narrowing towards apex; apex obliquely cut in lateral view and with incomplete, rim-shaped orange carina (missing in ventral portion); rather pale pinkish brown entirely covered in dense, irregular black pitting with smaller pits towards distal portion and majority of rather large, sometimes coalescent pits in basal portion. Lateral carinae of frons weakly zigzagged, more prominent towards apex, upcurved near apex and merging into a short dorsal carina reaching apex; lateral carinae of vertex merging with upcurved part of lateral carinae of frons; weakly marked, weakly zigzagged carina in middle portion of vertex, and some cross-carinae between latter and lateral carinae of vertex. Labium pale brown, reaching mesocoxae.
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FAE15FB912D9" blockId="5.[189,1398,1358,1455]" pageId="5" pageNumber="6">
THORAX (
<figureCitation id="132A2A69FFBFFFE6200FFAE05FB3121F" box="[306,473,1358,1385]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="5" pageNumber="6">Fig. 2B, EF</figureCitation>
). Pro- and mesonotum rather pale pinkish brown entirely covered in dense, irregular, strong black pitting with pits often larger than space between them and sometimes coalescent; median carina obsolete.
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180FA785C5511D7" blockId="5.[189,1399,1494,1697]" pageId="5" pageNumber="6">
TEGMINA (
<figureCitation id="132A2A69FFBFFFE62006FA785F821286" box="[315,488,1494,1520]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="5" pageNumber="6">Fig. 2A, CD</figureCitation>
). Strongly elongate and reticulate, pale pinkish brown, progressively paler, subhyaline towards apex, basal portion densely covered with minute pink spots; all surface irregularly covered in small to rather large black spots, mostly on veins, larger ones slightly protruding, large middle portion with very dense small spots; costal and postclaval margins subparallel, weakly diverging from base towards apex and curved at level of apex of clavus; apical margin rounded. Veins orange in large basal portion. Clavus open.
</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180F9665B221073" blockId="5.[189,1398,1734,1798]" pageId="5" pageNumber="6">VENATION. ScP+R forked in basal ⅓, MP forked more basally in basal 1/4; CuA forked near apex of clavus; PCu and A1 fused near apex of clavus; PCu+A1 fused with postclaval margin at nodal line.</paragraph>
<paragraph id="8BAE36ECFFBFFFE62180F8835D11101C" blockId="5.[189,1399,1836,1898]" pageId="5" pageNumber="6">
HIND WINGS (
<figureCitation id="132A2A69FFBFFFE6205DF8825F881031" box="[352,482,1836,1863]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="5" pageNumber="6">Fig. 2A, C</figureCitation>
). Subhyaline, milky white with veins slightly darker, vein PCu dark brown in basal half; postclaval margin weakly emarginate at A1 vein.
</paragraph>
<caption id="DF6E6664FFBCFFE52180F8265FCC109E" ID-DOI="http://doi.org/10.5281/zenodo.13837032" ID-Zenodo-Dep="13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,230,1928,1954]" targetBox="[217,1369,263,1889]" targetPageId="6">
<paragraph id="8BAE36ECFFBCFFE52180F8265FCC109E" blockId="6.[189,1398,1928,2024]" pageId="6" pageNumber="7">
<emphasis id="B965EAFEFFBCFFE52180F8265F6610D4" bold="true" box="[189,268,1928,1954]" pageId="6" pageNumber="7">Fig. 2.</emphasis>
<taxonomicName id="4C114D6FFFBCFFE52029F8275F8F10D4" authorityName="Constant &amp; Pham" authorityYear="2024" box="[276,485,1928,1954]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFBCFFE52029F8275F8F10D4" box="[276,485,1928,1954]" italics="true" pageId="6" pageNumber="7">Zanna bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFBCFFE520D1F8265C2010D4" box="[492,586,1928,1954]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS).
<emphasis id="B965EAFEFFBCFFE52259F8265D1310D4" bold="true" box="[868,889,1928,1954]" pageId="6" pageNumber="7">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B965EAFEFFBCFFE525B4F8265AF710D4" bold="true" box="[1161,1181,1928,1954]" pageId="6" pageNumber="7">B</emphasis>
. Head and thorax, dorsal view.
<emphasis id="B965EAFEFFBCFFE52064F8055F0410B3" bold="true" box="[345,366,1963,1989]" pageId="6" pageNumber="7">C</emphasis>
. Habitus, ventral view.
<emphasis id="B965EAFEFFBCFFE523AEF8055CC210B3" bold="true" box="[659,680,1963,1989]" pageId="6" pageNumber="7">D</emphasis>
. Habitus, lateral view.
<emphasis id="B965EAFEFFBCFFE522F9F8055DB210B3" bold="true" box="[964,984,1963,1989]" pageId="6" pageNumber="7">E</emphasis>
. Head, ventral view.
<emphasis id="B965EAFEFFBCFFE525E2F8055A9B10B3" bold="true" box="[1247,1265,1963,1989]" pageId="6" pageNumber="7">F</emphasis>
. Head and thorax, lateral view.
</paragraph>
</caption>
<caption id="DF6E6664FFBDFFE42183F8295B7E109E" ID-DOI="http://doi.org/10.5281/zenodo.13837034" ID-Zenodo-Dep="13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="7" pageNumber="8" startId="7.[190,230,1927,1953]" targetBox="[216,1346,273,1889]" targetPageId="7">
<paragraph id="8BAE36ECFFBDFFE42183F8295B7E109E" blockId="7.[188,1400,1927,2024]" pageId="7" pageNumber="8">
<emphasis id="B965EAFEFFBDFFE42183F8295F6D10D7" bold="true" box="[190,263,1927,1953]" pageId="7" pageNumber="8">Fig. 3.</emphasis>
<emphasis id="B965EAFEFFBDFFE42030F8295FBD10D7" box="[269,471,1927,1953]" italics="true" pageId="7" pageNumber="8">Zannabidoupana</emphasis>
<taxonomicNameLabel id="A2565785FFBDFFE420E1F8265C5F10D4" box="[476,565,1928,1954]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♂ (RBINS), terminalia.
<emphasis id="B965EAFEFFBDFFE42282F8295D9210D7" bold="true" box="[959,1016,1927,1953]" pageId="7" pageNumber="8">AD</emphasis>
. Pygofer, anal tube and gonostyli.
<emphasis id="B965EAFEFFBDFFE42181F8045EBB10B2" bold="true" box="[188,209,1962,1988]" pageId="7" pageNumber="8">A</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFBDFFE42098F8055FD310B3" bold="true" box="[421,441,1963,1989]" pageId="7" pageNumber="8">B</emphasis>
. Caudal view.
<emphasis id="B965EAFEFFBDFFE42362F8045C1E10B2" bold="true" box="[607,628,1962,1988]" pageId="7" pageNumber="8">C</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFBDFFE4222AF8055D4610B3" bold="true" box="[791,812,1963,1989]" pageId="7" pageNumber="8">D</emphasis>
. Ventral view.
<emphasis id="B965EAFEFFBDFFE422EEF8055A6610B3" bold="true" box="[979,1036,1963,1989]" pageId="7" pageNumber="8">EH</emphasis>
. Aedeagus.
<emphasis id="B965EAFEFFBDFFE425AEF8055ACD10B3" bold="true" box="[1171,1191,1963,1989]" pageId="7" pageNumber="8">E</emphasis>
. Left lateral view.
<emphasis id="B965EAFEFFBDFFE42180F8605EA5109E" bold="true" box="[189,207,1998,2024]" pageId="7" pageNumber="8">F</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFBDFFE42048F8635FE61091" bold="true" box="[373,396,1997,2023]" pageId="7" pageNumber="8">G</emphasis>
. Laterodorsal view.
<emphasis id="B965EAFEFFBDFFE42348F8605CE6109E" bold="true" box="[629,652,1998,2024]" pageId="7" pageNumber="8">H</emphasis>
. Ventral view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF6E6664FFB2FFEB2180F8DB5FDE10A0" ID-DOI="http://doi.org/10.5281/zenodo.13837036" ID-Zenodo-Dep="13837036" httpUri="https://zenodo.org/record/13837036/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,230,1909,1935]" targetBox="[257,1349,265,1872]" targetPageId="8">
<paragraph id="8BAE36ECFFB2FFEB2180F8DB5FDE10A0" blockId="8.[189,1398,1909,2006]" pageId="8" pageNumber="9">
<emphasis id="B965EAFEFFB2FFEB2180F8DB5F3F10F9" bold="true" box="[189,341,1909,1936]" pageId="8" pageNumber="9">Fig. 4. AB</emphasis>
.
<taxonomicName id="4C114D6FFFB2FFEB2055F8D95C5710E6" authorityName="Constant &amp; Pham" authorityYear="2024" box="[360,573,1910,1936]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFB2FFEB2055F8D95C5710E6" box="[360,573,1910,1936]" italics="true" pageId="8" pageNumber="9">Zanna bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB2FFEB2377F8D85CC010E6" box="[586,682,1910,1936]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
at the light trap, Vietnam, Bidoup-Nui Ba National Park, 23 Jul. 2014.
<emphasis id="B965EAFEFFB2FFEB205FF8375F1D10C5" bold="true" box="[354,375,1945,1971]" pageId="8" pageNumber="9">A</emphasis>
. Dorsal view.
<emphasis id="B965EAFEFFB2FFEB2315F8375C5610C5" bold="true" box="[552,572,1945,1971]" pageId="8" pageNumber="9">B</emphasis>
. Lateral view.
<emphasis id="B965EAFEFFB2FFEB23CCF8375D6C10C5" bold="true" box="[753,774,1945,1971]" pageId="8" pageNumber="9">C</emphasis>
. Habitat of
<taxonomicName id="4C114D6FFFB2FFEB22A6F8345A5D10C5" authorityName="Constant &amp; Pham" authorityYear="2024" box="[923,1079,1945,1971]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bidoupana" status="sp. nov.">
<emphasis id="B965EAFEFFB2FFEB22A6F8345A5D10C5" box="[923,1079,1945,1971]" italics="true" pageId="8" pageNumber="9">Z.bidoupana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2565785FFB2FFEB257CF8375AF710C5" box="[1089,1181,1945,1971]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
around Hon Giao station, 23 Jul. 2014.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFB3FFEA2180FEA75B3F161F" blockId="9.[187,1399,265,362]" pageId="9" pageNumber="10">
VENATION. ScP+R and MP forked at midlength; CuA forked slightly more basally then MP fork; PCu forked at basal ⅓; A1 forked close to base, A1
<subScript id="179534A9FFB3FFEA2344FE915CE8163B" attach="left" box="[633,642,319,333]" fontSize="6" pageId="9" pageNumber="10">2</subScript>
fused with A
<quantity id="4CE99B09FFB3FFEA2226FE825D2C1630" box="[795,838,300,326]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="9" pageNumber="10" unit="in" value="2.0">2 in</quantity>
distal portion and A1
<subScript id="179534A9FFB3FFEA2501FE915A2E163B" attach="none" box="[1084,1092,319,333]" fontSize="6" pageId="9" pageNumber="10">2</subScript>
+A2 forked slightly before reaching postclaval margin. Numerous crossveins in large portion along apical and postclaval margins.
</paragraph>
<paragraph id="8BAE36ECFFB3FFEA2180FE3F5B6F1508" blockId="9.[189,1399,401,639]" pageId="9" pageNumber="10">
LEGS (
<figureCitation id="132A2A69FFB3FFEA203AFE3F5FC316DD" box="[263,425,401,427]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="9" pageNumber="10">Fig. 2A, CD</figureCitation>
). Pale pinkish brown, pilose, rather short and robust. Pro- and mesocoxae with some black markings; all femora with some small black spots and conspicuous ring in distal ⅓, formed from large, more or less coalescent, black spots; pro- and mesotibiae with some small black spots and with apex black; pro- and mesotarsi blackish brown; metatibiae brown basally with all spines apically black, 56 lateral spines and 8 apical spines; first metatarsomere with 11 apical spines ventrally; second metatarsomere with 9 apical spines ventrally; first and second metatarsomere with a dense pad of microsetae ventrally; third metatarsomere elongate with a blackish brown ring. Metatibiotarsal formula: (56) 8/11/9.
</paragraph>
<caption id="DF6E6664FFB3FFEA2181F8645A6D1093" ID-DOI="http://doi.org/10.5281/zenodo.13837040" ID-Zenodo-Dep="13837040" box="[188,1031,1994,2021]" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="9" pageNumber="10" startId="9.[188,229,1994,2020]" targetBox="[215,1372,718,1954]" targetPageId="9">
<paragraph id="8BAE36ECFFB3FFEA2181F8645A6D1093" blockId="9.[188,1031,1994,2021]" box="[188,1031,1994,2021]" pageId="9" pageNumber="10">
<emphasis id="B965EAFEFFB3FFEA2181F8645F601093" bold="true" box="[188,266,1994,2021]" pageId="9" pageNumber="10">Fig. 5.</emphasis>
<taxonomicName id="4C114D6FFFB3FFEA202FF8625F371093" authorityName="Kirkaldy" authorityYear="1902" box="[274,349,1996,2021]" class="Insecta" family="Fulgoridae" genus="Zanna" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="B965EAFEFFB3FFEA202FF8625F371093" box="[274,349,1996,2021]" italics="true" pageId="9" pageNumber="10">Zanna</emphasis>
</taxonomicName>
, new spp. from Cambodia and Vietnam, distribution map.
</paragraph>
</caption>
<paragraph id="8BAE36ECFFB0FFE92180FEA75C9116FA" blockId="10.[189,1398,265,396]" pageId="10" pageNumber="11">
ABDOMEN (
<figureCitation id="132A2A69FFB0FFE9207AFEA75FBB1655" box="[327,465,265,291]" captionStart="Fig" captionStartId="6.[189,230,1928,1954]" captionTargetBox="[217,1369,263,1889]" captionTargetId="figure-46@6.[212,1369,258,1890]" captionTargetPageId="6" captionText="Fig. 2. Zanna bidoupana sp. nov., holotype, ♂ (RBINS). A. Habitus, dorsal view. B. Head and thorax, dorsal view. C. Habitus, ventral view. D. Habitus, lateral view. E. Head, ventral view. F. Head and thorax, lateral view." figureDoi="http://doi.org/10.5281/zenodo.13837032" httpUri="https://zenodo.org/record/13837032/files/figure.png" pageId="10" pageNumber="11">Fig. 2A, C</figureCitation>
). Tergites smooth and shiny, blackish brown bordered with yellowish along posterior margin; sternites pale pinkish brown with transverse irregular row of strong black dots and rounded, brown marking along middle portion of posterior margin, except pregenital sternite with large subtriangular blackish brown marking in middle.
</paragraph>
<paragraph id="8BAE36ECFFB0FFE92180FE1A5B191402" blockId="10.[189,1399,436,885]" pageId="10" pageNumber="11">
MALE TERMINALIA (
<figureCitation id="132A2A69FFB0FFE92099FE1A5F8616B8" box="[420,492,436,462]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3</figureCitation>
). Pygofer (
<emphasis id="B965EAFEFFB0FFE9234EFE1B5CF816B8" box="[627,658,437,462]" italics="true" pageId="10" pageNumber="11">Py</emphasis>
<figureCitation id="132A2A69FFB0FFE9238CFE1A5D5B16B8" box="[689,817,436,462]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3AD</figureCitation>
) about 1.77 times as high as long in lateral view, robust with posterior margin sinuate, forming large, broadly rounded posterior lobe in dorsal half; sides excavate in dorsal portion (visible in dorsal aspect,
<figureCitation id="132A2A69FFB0FFE9225FFE545DD41563" box="[866,958,506,533]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3C</figureCitation>
). Anal tube (
<emphasis id="B965EAFEFFB0FFE92565FE555A131563" box="[1112,1145,507,533]" italics="true" pageId="10" pageNumber="11">An</emphasis>
<figureCitation id="132A2A69FFB0FFE925AAFE545B7F1563" box="[1175,1301,506,533]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3AC</figureCitation>
) robust, 1.60 times as long as high and with ventral margin oblique in large posterior portion and angularly projecting in lateral view; 1.2 times as long as wide in dorsal view and with sides nearly straight, weakly diverging towards apex; more or less round in caudal view; epiproct and paraproct slightly supassing anal tube, rounded posteriorly in dorsal aspect. Gonostyli (
<emphasis id="B965EAFEFFB0FFE92253FD265DE915D4" box="[878,899,648,674]" italics="true" pageId="10" pageNumber="11">G</emphasis>
<figureCitation id="132A2A69FFB0FFE9229DFD295A4A15D7" box="[928,1056,647,673]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3AD</figureCitation>
) elongate, 2.46 times as long as high; in lateral view, ventral margin broadly rounded and dorsal margin rather broadly rounded after lateral hook (
<emphasis id="B965EAFEFFB0FFE92067FD605F15159E" box="[346,383,718,744]" italics="true" pageId="10" pageNumber="11">lhg</emphasis>
), apical margin rounded; lateral hook with basal portion projecting posterolaterad and apex strongly curved ventrad; in ventral view, lateral protrusion at about midlength, rather strongly and angularly marked. Aedeagus (
<figureCitation id="132A2A69FFB0FFE92318FCBA5CCF1458" box="[549,677,788,814]" captionStart="Fig" captionStartId="7.[190,230,1927,1953]" captionTargetBox="[216,1346,273,1889]" captionTargetId="figure-62@7.[212,1369,260,1892]" captionTargetPageId="7" captionText="Fig. 3. Zannabidoupana sp. nov., holotype, ♂ (RBINS), terminalia. AD. Pygofer, anal tube and gonostyli. A. Left lateral view. B. Caudal view. C. Dorsal view. D. Ventral view. EH. Aedeagus.E. Left lateral view. F. Dorsal view. G. Laterodorsal view. H. Ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.13837034" httpUri="https://zenodo.org/record/13837034/files/figure.png" pageId="10" pageNumber="11">Fig. 3EH</figureCitation>
) rather robust with lateral processes (
<emphasis id="B965EAFEFFB0FFE9254FFCBA5AFD1458" box="[1138,1175,788,814]" italics="true" pageId="10" pageNumber="11">lpa</emphasis>
) forming multiple lobes; dorsal process (
<emphasis id="B965EAFEFFB0FFE920F7FC995F9C1427" box="[458,502,823,849]" italics="true" pageId="10" pageNumber="11">dpa</emphasis>
) moderately protruding dorsally in lateral view; dorsal endosomal process (
<emphasis id="B965EAFEFFB0FFE921FAFCF45E9B1402" box="[199,241,858,884]" italics="true" pageId="10" pageNumber="11">dep</emphasis>
) in dorsal view subtriangular with lateral margins weakly curved and median slit in distal portion.
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFB0FFE92180FC325B20137E" pageId="10" pageNumber="11" type="biology_ecology">
<paragraph id="8BAE36ECFFB0FFE92180FC325F7614C0" blockId="10.[189,284,924,950]" box="[189,284,924,950]" pageId="10" pageNumber="11">
<heading id="D0E68180FFB0FFE92180FC325F7614C0" bold="true" box="[189,284,924,950]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B965EAFEFFB0FFE92180FC325F7614C0" bold="true" box="[189,284,924,950]" pageId="10" pageNumber="11">Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB0FFE92180FC645B20137E" blockId="10.[189,1399,970,1032]" pageId="10" pageNumber="11">
The specimens were collected at a mercury vapour light trap at Hon Giao station in July (
<figureCitation id="132A2A69FFB0FFE925DAFC645B001492" box="[1255,1386,970,996]" captionStart="Fig" captionStartId="8.[189,230,1909,1935]" captionTargetBox="[257,1349,265,1872]" captionTargetId="figure-45@8.[257,1349,265,1872]" captionTargetPageId="8" captionText="Fig. 4. AB. Zanna bidoupana sp. nov. at the light trap, Vietnam, Bidoup-Nui Ba National Park, 23 Jul. 2014. A. Dorsal view. B. Lateral view. C. Habitat of Z.bidoupana sp. nov. around Hon Giao station, 23 Jul. 2014." figureDoi="http://doi.org/10.5281/zenodo.13837036" httpUri="https://zenodo.org/record/13837036/files/figure.png" pageId="10" pageNumber="11">Fig. 4AB</figureCitation>
). The habitat around was a clearing covered in grass, herbaceaous weeds and small bushes (
<figureCitation id="132A2A69FFB0FFE925DDFC435B57137E" box="[1248,1341,1005,1032]" captionStart="Fig" captionStartId="8.[189,230,1909,1935]" captionTargetBox="[257,1349,265,1872]" captionTargetId="figure-45@8.[257,1349,265,1872]" captionTargetPageId="8" captionText="Fig. 4. AB. Zanna bidoupana sp. nov. at the light trap, Vietnam, Bidoup-Nui Ba National Park, 23 Jul. 2014. A. Dorsal view. B. Lateral view. C. Habitat of Z.bidoupana sp. nov. around Hon Giao station, 23 Jul. 2014." figureDoi="http://doi.org/10.5281/zenodo.13837036" httpUri="https://zenodo.org/record/13837036/files/figure.png" pageId="10" pageNumber="11">Fig. 4C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B6567FFB0FFE92180FB815CEF130E" box="[189,645,1071,1144]" pageId="10" pageNumber="11" type="distribution">
<paragraph id="8BAE36ECFFB0FFE92180FB815F3C133F" blockId="10.[189,342,1071,1097]" box="[189,342,1071,1097]" pageId="10" pageNumber="11">
<heading id="D0E68180FFB0FFE92180FB815F3C133F" bold="true" box="[189,342,1071,1097]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B965EAFEFFB0FFE92180FB815F3C133F" bold="true" box="[189,342,1071,1097]" pageId="10" pageNumber="11">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BAE36ECFFB0FFE92180FBF35CEF130E" blockId="10.[189,645,1117,1144]" box="[189,645,1117,1144]" pageId="10" pageNumber="11">
<collectingCountry id="F306767CFFB0FFE92180FBF35F4F1301" box="[189,293,1117,1143]" name="Vietnam" pageId="10" pageNumber="11">Vietnam</collectingCountry>
:
<collectingRegion id="49D5F80EFFB0FFE9200DFBF05C771301" box="[304,541,1117,1144]" country="Vietnam" name="Lam Dong" pageId="10" pageNumber="11">Lam Dong Province</collectingRegion>
(
<figureCitation id="132A2A69FFB0FFE92313FBF35C1C130E" box="[558,630,1117,1144]" captionStart="Fig" captionStartId="9.[188,229,1994,2020]" captionTargetBox="[215,1372,718,1954]" captionTargetId="figure-180@9.[215,1372,718,1954]" captionTargetPageId="9" captionText="Fig. 5. Zanna, new spp. from Cambodia and Vietnam, distribution map." figureDoi="http://doi.org/10.5281/zenodo.13837040" httpUri="https://zenodo.org/record/13837040/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,352 @@
<document id="13BB7177FCBB98CA182E7B34DA133126" ID-DOI="10.5252/zoosystema2024v46a23" ID-ISSN="1638-9387" ID-Zenodo-Dep="13837101" ID-ZooBank="9195A8AB-6FA8-4828-B8D1-DDEC94E99A6D" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727253036749" checkinUser="plazi" docAuthor="Pati, Sameer K., Peter, Santanu Mitra &amp; L., Peter K." docDate="2024" docId="03F38788FFC4FFB0FC21F9204DBD2FDB" docLanguage="en" docName="zoosystema.2024.46.23.pdf" docOrigin="Zoosystema 46 (23)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.9:Zoosystema.2020-.journal_article.type1" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article.type1" docStyleVersion="9" docTitle="Capitamon meitei Pati, Peter &amp; L., 2024, n. sp." docType="treatment" docUuid="109C2D01-A922-4EE1-ABCE-710D7E91F2F2" docUuidSource="ZooBank" docVersion="2" lastPageNumber="606" masterDocId="FFCAFFF0FFD6FFA6FFCAFF9E4F402A36" masterDocTitle="Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae)" masterLastPageNumber="615" masterPageNumber="589" pageNumber="605" updateTime="1727253493743" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="103B8524C60A991D681A1155DEF9B126">Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae)</mods:title>
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<mods:namePart id="C05B7D43EF9425837FD0C3B6A2CE0C80">Pati, Sameer K.</mods:namePart>
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<mods:namePart id="16135F59EA73B1F0AB6AF15226F48BE9">Peter, Santanu Mitra</mods:namePart>
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<mods:namePart id="C9A3C028EADECE0C9959E13A213CDC80">L., Peter K.</mods:namePart>
<mods:affiliation id="D3F8FFC2073BF288859FCF2EE36CCE39">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377 (Republic of Singapore)</mods:affiliation>
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<mods:title id="ACE103A62A1A4A1C23217630AF84786C">Zoosystema</mods:title>
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<mods:part id="441AC30A1763691CC6449F1C4B39E2F1">
<mods:date id="542A48D6EDBEFA3437AEB5F3EEB0F9CD">2024</mods:date>
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<mods:number id="AB9303F12D4CE391D08C3FA254DF911D">2024-09-24</mods:number>
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<mods:number id="7CFBBE465E971B30DA6598A78AC69C47">46</mods:number>
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<mods:number id="A7E5EC005AA4DD08D39EEB03CB448691">23</mods:number>
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<mods:url id="38039082DDABFF019232FAC975048AAB">https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/zoosystema2024v46a23.pdf</mods:url>
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<mods:identifier id="20C6793D500286342737813CFF0FF8C2" type="DOI">10.5252/zoosystema2024v46a23</mods:identifier>
<mods:identifier id="7B58D9FDBADB0C6BBBA563204DAA2F9B" type="ISSN">1638-9387</mods:identifier>
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<taxonomicName id="4C5A4D1DFFC4FFB4FC21F9204BEF2CEE" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" box="[1003,1199,1726,1752]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="605" phylum="Arthropoda" rank="species" species="meitei" status="sp. nov.">
<emphasis id="B92EEA8CFFC4FFB4FC21F9204BEF2CEE" bold="true" box="[1003,1199,1726,1752]" italics="true" pageId="18" pageNumber="605">Capitamon meitei</emphasis>
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<taxonomicNameLabel id="A21D57F7FFC4FFB4FB7CF9204BB12CEE" box="[1206,1265,1726,1752]" pageId="18" pageNumber="605" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="C3406515FFC4FFB4FBDEF9404AD52D1A" pageId="18" pageNumber="605" type="description">
<paragraph id="8BE5369EFFC4FFB4FBDEF9404B882CCE" blockId="18.[1003,1265,1726,1784]" box="[1044,1224,1757,1784]" pageId="18" pageNumber="605">
(
<figureCitation id="13612A1BFFC4FFB4FBD6F9434B2B2CCE" box="[1052,1131,1757,1784]" captionStart="FIG" captionStartId="20.[132,143,1985,2003]" captionTargetBox="[132,1359,215,1943]" captionTargetId="figure-79@20.[698,1267,759,1307]" captionTargetPageId="20" captionText="FIG. 12. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bars: A, C, 20 mm; B, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13837134" httpUri="https://zenodo.org/record/13837134/files/figure.png" pageId="18" pageNumber="605">Figs 12</figureCitation>
;
<figureCitation id="13612A1BFFC4FFB4FBB3F9404BD52CCE" box="[1145,1173,1758,1784]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="18" pageNumber="605">13</figureCitation>
;
<figureCitation id="13612A1BFFC4FFB4FB69F9404BFF2CCE" box="[1187,1215,1758,1784]" captionStart="FIG" captionStartId="22.[131,142,1309,1327]" captionTargetBox="[146,1443,214,1267]" captionTargetId="figure-506@22.[132,1455,204,1274]" captionTargetPageId="22" captionText="FIG. 14. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, left G1 ultimate article (dorsal view); B, left G2; C, left G1 ultimate article (ventral view); D, left G1 (dorsal view); E, left G1 (ventral view). Scale bar: A, C, 1 mm; B, D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837138" httpUri="https://zenodo.org/record/13837138/files/figure.png" pageId="18" pageNumber="605">14</figureCitation>
)
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urn:lsid:zoobank.org:act:
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. —
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<typeStatus id="54E1883CFFC4FFB4FC3BF8D04B0F2D50" box="[1009,1103,1870,1894]" pageId="18" pageNumber="605" type="holotype">Holotype</typeStatus>
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<collectingCountry id="F34D760EFFC4FFB4FB97F8CE4BD22D5E" box="[1117,1170,1872,1896]" name="India" pageId="18" pageNumber="605">India</collectingCountry>
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);
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:
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:
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leg.;
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C.2405.
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<typeStatus id="54E1883CFFC4FFB4FCE7F8004CC72D80" box="[813,903,1950,1974]" pageId="18" pageNumber="605" type="paratype">Paratypes</typeStatus>
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.
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<collectingCountry id="F34D760EFFC4FFB4FC5BF8014C832D81" box="[913,963,1951,1975]" name="India" pageId="18" pageNumber="605">India</collectingCountry>
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(39.9
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<quantity id="4CA29B7BFFC4FFB4FBFBF8004BC82D80" box="[1073,1160,1950,1974]" metricMagnitude="-2" metricUnit="m" metricValue="3.04" pageId="18" pageNumber="605" unit="mm" value="30.4">30.4 mm</quantity>
),
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(36.0
<emphasis id="B92EEA8CFFC4FFB4FB2EF8004BB22D83" box="[1252,1266,1950,1973]" italics="true" pageId="18" pageNumber="605">×</emphasis>
<quantity id="4CA29B7BFFC4FFB4FB3EF8004A0B2D80" box="[1268,1355,1950,1974]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="18" pageNumber="605" unit="mm" value="27.4">27.4 mm</quantity>
);
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:
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:
<collectingMunicipality id="6B81ACE4FFC4FFB4FBE2F8264BE62DE6" box="[1064,1190,1976,2000]" pageId="18" pageNumber="605">Leimakhong</collectingMunicipality>
;
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,
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; alt.
<quantity id="4CA29B7BFFC4FFB4FC98F84D4CD42DDC" box="[850,916,2003,2027]" metricMagnitude="2" metricUnit="m" metricValue="8.84" pageId="18" pageNumber="605" unit="m" value="884.0">
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;
<collectorName id="26AF5348FFC4FFB4FBD8F84D4BFE2DDD" box="[1042,1214,2003,2027]" pageId="18" pageNumber="605">Thangpurba Devi</collectorName>
leg.;
<collectionCode id="ED4BAE5BFFC4FFB4FB39F84D4A182DDD" box="[1267,1368,2003,2027]" pageId="18" pageNumber="605">ZSI-WRC</collectionCode>
C.2406.
<smallCapsWord id="8D03A042FFC5FFB5FF4EFA174FAF2FA9" baselines="1434,1435" box="[132,239,1417,1440]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="19" pageNumber="606">DIAGNOSIS</smallCapsWord>
. —
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ovate, broader than long (
<collectionCode id="ED4BAE5BFFC5FFB5FDBEFA174DDE2F97" box="[628,670,1417,1441]" pageId="19" pageNumber="606">CW</collectionCode>
/CL = 1.3- 1.4), relatively low (
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/CW = 0.4); epigastric cristae well-developed, rugose; postorbital cristae well-developed, relatively sharp, oblique in dorsal view; external orbital angle triangular, with short outer margin,
<emphasis id="B92EEA8CFFC5FFB5FF12FA6A4FA82C3D" box="[216,232,1524,1547]" italics="true" pageId="19" pageNumber="606">c.</emphasis>
2
<emphasis id="B92EEA8CFFC5FFB5FEC9FA6A4E512C3D" box="[259,273,1524,1547]" italics="true" pageId="19" pageNumber="606">×</emphasis>
inner margin; epibranchial tooth low, blunt, positioned above level of postorbital cristae; cervical grooves continuous, reaching to level of postorbital cristae; epistome posterior margin with well-developed, triangular median tooth, outer parts sloping downwards laterally (
<figureCitation id="13612A1BFFC5FFB5FE9DF9C04E8D2C40" box="[343,461,1630,1654]" captionStart="FIG" captionStartId="20.[132,143,1985,2003]" captionTargetBox="[132,1359,215,1943]" captionTargetId="figure-79@20.[698,1267,759,1307]" captionTargetPageId="20" captionText="FIG. 12. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bars: A, C, 20 mm; B, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13837134" httpUri="https://zenodo.org/record/13837134/files/figure.png" pageId="19" pageNumber="606">Fig. 12A, B</figureCitation>
).
<collectorName id="26AF5348FFC5FFB5FE2AF9C04D562C40" box="[480,534,1630,1654]" pageId="19" pageNumber="606">Third</collectorName>
maxilliped exopod with short flagellum, shorter than width of merus (
<figureCitation id="13612A1BFFC5FFB5FDF4F9E74DD42CA7" box="[574,660,1657,1681]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">Fig. 13B</figureCitation>
).
<collectorName id="26AF5348FFC5FFB5FD6FF9E74C462CA7" box="[677,774,1657,1681]" pageId="19" pageNumber="606">Chelipeds</collectorName>
rugose, unequal in adult males, inner distal tooth on carpus relatively broad (
<figureCitation id="13612A1BFFC5FFB5FECBF9304E322CF0" box="[257,370,1710,1734]" captionStart="FIG" captionStartId="20.[132,143,1985,2003]" captionTargetBox="[132,1359,215,1943]" captionTargetId="figure-79@20.[698,1267,759,1307]" captionTargetPageId="20" captionText="FIG. 12. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bars: A, C, 20 mm; B, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13837134" httpUri="https://zenodo.org/record/13837134/files/figure.png" pageId="19" pageNumber="606">Figs 12A, C</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FEB0F9304EE82CF0" box="[378,424,1710,1734]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">13C</figureCitation>
).
<collectorName id="26AF5348FFC5FFB5FE73F9304EAA2CF0" box="[441,490,1710,1734]" pageId="19" pageNumber="606">Male</collectorName>
s3/s4 indiscernible (
<figureCitation id="13612A1BFFC5FFB5FD61F9304C422CF0" box="[683,770,1710,1734]" captionStart="FIG" captionStartId="20.[132,143,1985,2003]" captionTargetBox="[132,1359,215,1943]" captionTargetId="figure-79@20.[698,1267,759,1307]" captionTargetPageId="20" captionText="FIG. 12. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bars: A, C, 20 mm; B, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13837134" httpUri="https://zenodo.org/record/13837134/files/figure.png" pageId="19" pageNumber="606">Figs 12C</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FF4EF9574FF42CD7" box="[132,180,1737,1761]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">13D</figureCitation>
).
<collectorName id="26AF5348FFC5FFB5FF02F9564FBA2CD6" box="[200,250,1736,1760]" pageId="19" pageNumber="606">Male</collectorName>
pleonal somite 6 relatively broader (proximal width
<emphasis id="B92EEA8CFFC5FFB5FF4CF97A4FCD2CCD" box="[134,141,1764,1787]" italics="true" pageId="19" pageNumber="606">c</emphasis>
. 2.1-2.2
<emphasis id="B92EEA8CFFC5FFB5FF21F97A4FB92CCD" box="[235,249,1764,1787]" italics="true" pageId="19" pageNumber="606">×</emphasis>
medial length), equal in length to pleonal somite 5, with convex lateral margins (
<figureCitation id="13612A1BFFC5FFB5FE51F9604EB32D20" box="[411,499,1790,1814]" captionStart="FIG" captionStartId="20.[132,143,1985,2003]" captionTargetBox="[132,1359,215,1943]" captionTargetId="figure-79@20.[698,1267,759,1307]" captionTargetPageId="20" captionText="FIG. 12. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bars: A, C, 20 mm; B, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13837134" httpUri="https://zenodo.org/record/13837134/files/figure.png" pageId="19" pageNumber="606">Figs 12C</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FE34F9604D692D20" box="[510,553,1790,1814]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">13E</figureCitation>
).
<collectorName id="26AF5348FFC5FFB5FDF0F9604D2B2D20" box="[570,619,1790,1814]" pageId="19" pageNumber="606">Male</collectorName>
telson relatively broader (proximal width
<emphasis id="B92EEA8CFFC5FFB5FE4AF8874ECF2D06" box="[384,399,1817,1840]" italics="true" pageId="19" pageNumber="606">c.</emphasis>
1.3-1.4
<emphasis id="B92EEA8CFFC5FFB5FE2EF8874EB22D06" box="[484,498,1817,1840]" italics="true" pageId="19" pageNumber="606">×</emphasis>
medial length), with gently to strongly concave lateral margins (
<figureCitation id="13612A1BFFC5FFB5FE20F8AD4D042D7D" box="[490,580,1843,1867]" captionStart="FIG" captionStartId="20.[132,143,1985,2003]" captionTargetBox="[132,1359,215,1943]" captionTargetId="figure-79@20.[698,1267,759,1307]" captionTargetPageId="20" captionText="FIG. 12. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, overall dorsal view; B, frontal view of cephalothorax; C, overall ventral view. Scale bars: A, C, 20 mm; B, 10 mm." figureDoi="http://doi.org/10.5281/zenodo.13837134" httpUri="https://zenodo.org/record/13837134/files/figure.png" pageId="19" pageNumber="606">Figs 12C</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FD9AF8AD4D3C2D7D" box="[592,636,1843,1867]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">13E</figureCitation>
). G1 ultimate article relatively less strongly bent at angle of about 75° from longitudinal axis, with sinuous outer margin, dorsal flap relatively high, narrowly triangular (
<figureCitation id="13612A1BFFC5FFB5FE81F81D4EDF2DAC" box="[331,415,1923,1947]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">Figs 13F</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FE62F81D4E932DAD" box="[424,467,1923,1947]" captionStart="FIG" captionStartId="22.[131,142,1309,1327]" captionTargetBox="[146,1443,214,1267]" captionTargetId="figure-506@22.[132,1455,204,1274]" captionTargetPageId="22" captionText="FIG. 14. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, left G1 ultimate article (dorsal view); B, left G2; C, left G1 ultimate article (ventral view); D, left G1 (dorsal view); E, left G1 (ventral view). Scale bar: A, C, 1 mm; B, D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837138" httpUri="https://zenodo.org/record/13837138/files/figure.png" pageId="19" pageNumber="606">14A</figureCitation>
, C-E). G2
<emphasis id="B92EEA8CFFC5FFB5FDF0F81A4D0A2DAD" box="[570,586,1924,1947]" italics="true" pageId="19" pageNumber="606">c.</emphasis>
1.2
<emphasis id="B92EEA8CFFC5FFB5FDBEF81A4DC22DAD" box="[628,642,1924,1947]" italics="true" pageId="19" pageNumber="606">×</emphasis>
length of G1; ultimate article long,
<emphasis id="B92EEA8CFFC5FFB5FE80F8004E192D83" box="[330,345,1950,1973]" italics="true" pageId="19" pageNumber="606">c.</emphasis>
0.5
<emphasis id="B92EEA8CFFC5FFB5FE48F8004ED02D83" box="[386,400,1950,1973]" italics="true" pageId="19" pageNumber="606">×</emphasis>
length of penultimate article (
<figureCitation id="13612A1BFFC5FFB5FD61F8004C432D80" box="[683,771,1950,1974]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">Figs 13G</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FF4EF8264F8F2DE6" box="[132,207,1976,2000]" captionStart="FIG" captionStartId="22.[131,142,1309,1327]" captionTargetBox="[146,1443,214,1267]" captionTargetId="figure-506@22.[132,1455,204,1274]" captionTargetPageId="22" captionText="FIG. 14. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, left G1 ultimate article (dorsal view); B, left G2; C, left G1 ultimate article (ventral view); D, left G1 (dorsal view); E, left G1 (ventral view). Scale bar: A, C, 1 mm; B, D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837138" httpUri="https://zenodo.org/record/13837138/files/figure.png" pageId="19" pageNumber="606">14B, D</figureCitation>
).
<collectorName id="26AF5348FFC5FFB5FF2AF8264E612DE6" box="[224,289,1976,2000]" pageId="19" pageNumber="606">Vulvae</collectorName>
laterally completely covered by protruding sternal cover, invisible in ventral view (
<figureCitation id="13612A1BFFC5FFB5FE72F84D4D472DDD" box="[440,519,2003,2027]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">Fig. 13I</figureCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF256616FFC5FFB5FF4EFA834BEC2F73" ID-DOI="http://doi.org/10.5281/zenodo.13837132" ID-Zenodo-Dep="13837132" httpUri="https://zenodo.org/record/13837132/files/figure.png" pageId="19" pageNumber="606" startId="19.[132,143,1309,1327]" targetBox="[140,1449,214,1265]" targetPageId="19" targetType="figure">
<paragraph id="8BE5369EFFC5FFB5FF4EFA834BEC2F73" blockId="19.[132,1457,1308,1349]" pageId="19" pageNumber="606">
<smallCapsWord id="8D03A042FFC5FFB5FF4EFA834FDD2F1B" baselines="1322,1322" box="[132,157,1309,1327]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="19" pageNumber="606">FIG</smallCapsWord>
. 11. —
<taxonomicName id="4C5A4D1DFFC5FFB5FF2AFA824D682F19" authority="(Alcock, 1909)" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" baseAuthorityName="Alcock" baseAuthorityYear="1909" box="[224,552,1308,1327]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="606" phylum="Arthropoda" rank="species" species="manipurense" status="comb. nov.">
<emphasis id="B92EEA8CFFC5FFB5FF2AFA824EEC2F19" box="[224,428,1308,1327]" italics="true" pageId="19" pageNumber="606">Capitamon manipurense</emphasis>
(
<bibRefCitation id="EFCB4B6FFFC5FFB5FE7CFA834D642F19" author="ALCOCK A." box="[438,548,1309,1327]" pageId="19" pageNumber="606" pagination="243 - 252" refId="ref17242" refString="ALCOCK A. 1909. - Diagnoses of new species and varieties of freshwater crabs. Nos. 1 - 3. Records of the Indian Museum 3 (3): 243 - 252. https: // doi. org / 10.26515 / rzsi / v 3 / i 3 / 1909 / 163275" type="journal article" year="1909">Alcock, 1909</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A21D57F7FFC5FFB5FDE4FA834D3D2F19" box="[558,637,1309,1327]" pageId="19" pageNumber="606" rank="species">n. comb.</taxonomicNameLabel>
, lectotype ♂ (39.9 × 30.0 mm) (ZSIK C.6923/3):
<emphasis id="B92EEA8CFFC5FFB5FBDAFA834B5D2F19" bold="true" box="[1040,1053,1309,1327]" pageId="19" pageNumber="606">A</emphasis>
, left G1 ultimate article (dorsal view);
<emphasis id="B92EEA8CFFC5FFB5FA90FA834A272F19" bold="true" box="[1370,1383,1309,1327]" pageId="19" pageNumber="606">B</emphasis>
, left G2;
<emphasis id="B92EEA8CFFC5FFB5FF4EFAAD4FD22F73" bold="true" box="[132,146,1331,1349]" pageId="19" pageNumber="606">C</emphasis>
, left G1 ultimate article (ventral view);
<emphasis id="B92EEA8CFFC5FFB5FE05FAAD4E9D2F73" bold="true" box="[463,477,1331,1349]" pageId="19" pageNumber="606">D</emphasis>
, left G1 (dorsal view);
<emphasis id="B92EEA8CFFC5FFB5FD5CFAAD4DE22F73" bold="true" box="[662,674,1331,1349]" pageId="19" pageNumber="606">E</emphasis>
, left G1 (ventral view). Scale bar: A, C, 0.5 mm; B, D, E, 1 mm.
</paragraph>
</caption>
<subSubSection id="C3406515FFC5FFB5FCE7FA174AE62C16" pageId="19" pageNumber="606" type="etymology">
<paragraph id="8BE5369EFFC5FFB5FCE7FA174CDB2FE0" blockId="19.[813,1457,1417,1494]" pageId="19" pageNumber="606">
<smallCapsWord id="8D03A042FFC5FFB5FCE7FA174CE82FA9" baselines="1434,1435" box="[813,936,1417,1440]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="19" pageNumber="606">ETYMOLOGY</smallCapsWord>
. — The species name,
<taxonomicName id="4C5A4D1DFFC5FFB5FB5CFA144B8E2F97" box="[1174,1230,1418,1441]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="606" phylum="Arthropoda" rank="species" species="meitei">
<emphasis id="B92EEA8CFFC5FFB5FB5CFA144B8E2F97" box="[1174,1230,1418,1441]" italics="true" pageId="19" pageNumber="606">meitei</emphasis>
</taxonomicName>
, means Manipuri, the people/language of
<collectingRegion id="499EF87CFFC5FFB5FC27FA3D4B062F8D" box="[1005,1094,1443,1467]" country="India" name="Manipur" pageId="19" pageNumber="606">Manipur</collectingRegion>
. The name is used as a Latin noun in apposition.
</paragraph>
<paragraph id="8BE5369EFFC5FFB5FCE7FA734AE62C16" blockId="19.[812,1456,1517,1568]" pageId="19" pageNumber="606">
<smallCapsWord id="8D03A042FFC5FFB5FCE7FA734C842C32" baselines="1535,1536" box="[813,964,1517,1541]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Geographical" pageId="19" pageNumber="606">GEOGRAPHICAL</smallCapsWord>
<smallCapsWord id="8D03A042FFC5FFB5FC00FA6D4B122C32" baselines="1536" box="[970,1106,1523,1540]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="distribution" pageId="19" pageNumber="606">DISTRIBUTION</smallCapsWord>
. —
<taxonomicName id="4C5A4D1DFFC5FFB5FBB7FA704A652C33" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" box="[1149,1317,1518,1541]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="606" phylum="Arthropoda" rank="species" species="meitei" status="n. gen., n. sp.">
<emphasis id="B92EEA8CFFC5FFB5FBB7FA704A652C33" box="[1149,1317,1518,1541]" italics="true" pageId="19" pageNumber="606">Capitamon meitei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57F7FFC5FFB5FAE6FA704AF02C33" box="[1324,1456,1518,1542]" pageId="19" pageNumber="606" rank="species">n. gen., n. sp.</taxonomicNameLabel>
is known only from
<collectingRegion id="499EF87CFFC5FFB5FC39F9964B3C2C29" box="[1011,1148,1544,1568]" country="India" name="Manipur" pageId="19" pageNumber="606">Manipur state</collectingRegion>
of northeastern
<collectingCountry id="F34D760EFFC5FFB5FAD4F9964A112C16" box="[1310,1361,1544,1568]" name="India" pageId="19" pageNumber="606">India</collectingCountry>
(
<figureCitation id="13612A1BFFC5FFB5FAAAF9964AD82C16" box="[1376,1432,1544,1568]" captionStart="FIG" captionStartId="4.[146,157,1767,1785]" captionTargetBox="[191,1399,241,1671]" captionTargetId="figure-108@4.[735,1284,766,1327]" captionTargetPageId="4" captionText="FIG. 1. — Map showing Myanmar, India, northeastern India, and distribution of Indochinamon beieri (Pretzmann, 1966) and the species of Capitamon n. gen." figureDoi="http://doi.org/10.5281/zenodo.13837103" httpUri="https://zenodo.org/record/13837103/files/figure.png" pageId="19" pageNumber="606">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3406515FFC5FFB0FCE7F9CD4DBD2FDB" lastPageId="22" lastPageNumber="609" pageId="19" pageNumber="606" type="discussion">
<paragraph id="8BE5369EFFC5FFB5FCE7F9CD4CD12C5D" blockId="19.[811,1457,1619,2028]" box="[813,913,1619,1645]" pageId="19" pageNumber="606">
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</paragraph>
<paragraph id="8BE5369EFFC5FFB0FCE7F9EC4DBD2FDB" blockId="19.[811,1457,1619,2028]" lastBlockId="22.[130,775,1427,1517]" lastPageId="22" lastPageNumber="609" pageId="19" pageNumber="606">
Among the nominal species of the genus,
<taxonomicName id="4C5A4D1DFFC5FFB5FB3DF9ED4A162CBA" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" box="[1271,1366,1651,1677]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="606" phylum="Arthropoda" rank="species" species="meitei" status="n. gen., n. sp.">
<emphasis id="B92EEA8CFFC5FFB5FB3DF9ED4A162CBA" box="[1271,1366,1651,1677]" italics="true" pageId="19" pageNumber="606">C. meitei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57F7FFC5FFB5FA94F9ED4C272C9B" pageId="19" pageNumber="606" rank="species">n. gen., n. sp.</taxonomicNameLabel>
most resembles
<taxonomicName id="4C5A4D1DFFC5FFB5FBDAF90D4BEE2C9A" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" box="[1040,1198,1683,1709]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="606" phylum="Arthropoda" rank="species" species="mizoramense" status="n. gen., n. sp.">
<emphasis id="B92EEA8CFFC5FFB5FBDAF90D4BEE2C9A" box="[1040,1198,1683,1709]" italics="true" pageId="19" pageNumber="606">C. mizoramense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57F7FFC5FFB5FB7EF90D4A012C9B" box="[1204,1345,1683,1709]" pageId="19" pageNumber="606" rank="species">n. gen., n. sp.</taxonomicNameLabel>
mainly by the G1 ultimate article, which has a sinuous outer margin, and the relatively high and narrowly triangular dorsal flap (
<figureCitation id="13612A1BFFC5FFB5FA9BF94C4AEB2CDA" box="[1361,1451,1746,1772]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">Figs 13F</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FCE7F96C4C1B2D3A" box="[813,859,1778,1804]" captionStart="FIG" captionStartId="22.[131,142,1309,1327]" captionTargetBox="[146,1443,214,1267]" captionTargetId="figure-506@22.[132,1455,204,1274]" captionTargetPageId="22" captionText="FIG. 14. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, left G1 ultimate article (dorsal view); B, left G2; C, left G1 ultimate article (ventral view); D, left G1 (dorsal view); E, left G1 (ventral view). Scale bar: A, C, 1 mm; B, D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837138" httpUri="https://zenodo.org/record/13837138/files/figure.png" pageId="19" pageNumber="606">14A</figureCitation>
, C-E; 16F; 17A, C-E).
<taxonomicName id="4C5A4D1DFFC5FFB5FB98F96C4A442D3A" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" box="[1106,1284,1778,1804]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="606" phylum="Arthropoda" rank="species" species="meitei" status="n. gen., n. sp.">
<emphasis id="B92EEA8CFFC5FFB5FB98F96C4A442D3A" box="[1106,1284,1778,1804]" italics="true" pageId="19" pageNumber="606">Capitamon meitei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57F7FFC5FFB5FAC1F96C4AD92D3A" box="[1291,1433,1778,1804]" pageId="19" pageNumber="606" rank="species">n. gen., n. sp.</taxonomicNameLabel>
is nevertheless differentiated from
<taxonomicName id="4C5A4D1DFFC5FFB5FBB0F88C4A592D1D" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" box="[1146,1305,1810,1836]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="606" phylum="Arthropoda" rank="species" species="mizoramense" status="n. gen., n. sp.">
<emphasis id="B92EEA8CFFC5FFB5FBB0F88C4A592D1D" box="[1146,1305,1810,1836]" italics="true" pageId="19" pageNumber="606">C. mizoramense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57F7FFC5FFB5FAEAF88C4AF02D1A" box="[1312,1456,1810,1836]" pageId="19" pageNumber="606" rank="species">n. gen., n. sp.</taxonomicNameLabel>
by the relatively less strongly bent G1 ultimate article, which is bent at an angle of about 75° from longitudinal axis of the G1 (
<figureCitation id="13612A1BFFC5FFB5FCFFF8EF4CCE2DBA" box="[821,910,1905,1932]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="19" pageNumber="606">Figs 13F</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FC52F8EF4CA42DBD" box="[920,996,1905,1932]" captionStart="FIG" captionStartId="22.[131,142,1309,1327]" captionTargetBox="[146,1443,214,1267]" captionTargetId="figure-506@22.[132,1455,204,1274]" captionTargetPageId="22" captionText="FIG. 14. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405): A, left G1 ultimate article (dorsal view); B, left G2; C, left G1 ultimate article (ventral view); D, left G1 (dorsal view); E, left G1 (ventral view). Scale bar: A, C, 1 mm; B, D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837138" httpUri="https://zenodo.org/record/13837138/files/figure.png" pageId="19" pageNumber="606">14D, E</figureCitation>
) (vs G1 ultimate article more strongly bent at an angle of about 110° from longitudinal axis of the G1;
<figureCitation id="13612A1BFFC5FFB5FA9BF80F4AEB2D9A" box="[1361,1451,1937,1964]" captionStart="FIG" captionStartId="24.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-138@24.[698,1267,782,1352]" captionTargetPageId="24" captionText="FIG. 16. — Capitamon mizoramense n. gen., n. sp., holotype ♂ (46.3 × 35.7 mm) (ZSIK C.6122/2A) (A-G), paratype ♀ (51.3 × 37.6 mm) (ZSIK C.6122/2B) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837142" httpUri="https://zenodo.org/record/13837142/files/figure.png" pageId="19" pageNumber="606">Figs 16F</figureCitation>
;
<figureCitation id="13612A1BFFC5FFB5FCE7F82F4C3B2DFD" box="[813,891,1969,1996]" captionStart="FIG" captionStartId="25.[132,143,1367,1385]" captionTargetBox="[145,1444,215,1325]" captionTargetId="figure-507@25.[132,1455,205,1333]" captionTargetPageId="25" captionText="FIG. 17. — Capitamon mizoramense n. gen., n. sp., holotype ♂ (46.3 × 35.7 mm) (ZSIK C.6122/2A): A, left G1 ultimate article (dorsal view); B, left G2; C, left G1 ultimate article (ventral view); D, left G1 (dorsal view); E, left G1 (ventral view). Scale bar: A, C, 1 mm; B, D, E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837144" httpUri="https://zenodo.org/record/13837144/files/figure.png" pageId="19" pageNumber="606">17D, E</figureCitation>
). In addition, the vulvae are completely covered laterally by the protruding sternal cover and invisible in the ventral view in
<taxonomicName id="4C5A4D1DFFC0FFB0FF55FA0D4FB82F9B" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" box="[159,248,1427,1453]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="609" phylum="Arthropoda" rank="species" species="meitei" status="n. gen., n. sp.">
<emphasis id="B92EEA8CFFC0FFB0FF55FA0D4FB82F9B" box="[159,248,1427,1453]" italics="true" pageId="22" pageNumber="609">C. meitei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57F7FFC0FFB0FF34FA0A4EC62F98" box="[254,390,1428,1454]" pageId="22" pageNumber="609" rank="species">n. gen., n. sp.</taxonomicNameLabel>
(
<figureCitation id="13612A1BFFC0FFB0FE5EFA0D4EA42F9B" box="[404,484,1427,1453]" captionStart="FIG" captionStartId="21.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-132@21.[698,1267,782,1352]" captionTargetPageId="21" captionText="FIG. 13. — Capitamon meitei n. gen., n. sp., holotype ♂ (51.7 × 38.8 mm) (ZSI-WRC C.2405) (A-G), paratype ♀ (36.0 × 27.4 mm) (ZSI-WRC C.2406) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837136" httpUri="https://zenodo.org/record/13837136/files/figure.png" pageId="22" pageNumber="609">Fig. 13I</figureCitation>
), whereas the vulvae are partially covered laterally by the protruding sternal cover and visible in the ventral view in
<taxonomicName id="4C5A4D1DFFC0FFB0FEA8FA4D4EBF2FDB" authorityName="Pati &amp; Peter &amp; L." authorityYear="2024" box="[354,511,1491,1517]" class="Malacostraca" family="Potamidae" genus="Capitamon" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="609" phylum="Arthropoda" rank="species" species="mizoramense" status="n. gen., n. sp.">
<emphasis id="B92EEA8CFFC0FFB0FEA8FA4D4EBF2FDB" box="[354,511,1491,1517]" italics="true" pageId="22" pageNumber="609">C. mizoramense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57F7FFC0FFB0FDCFFA4D4DD02FDB" box="[517,656,1491,1517]" pageId="22" pageNumber="609" rank="species">n. gen., n. sp.</taxonomicNameLabel>
(
<figureCitation id="13612A1BFFC0FFB0FD57FA4D4DB02FDB" box="[669,752,1491,1517]" captionStart="FIG" captionStartId="24.[132,143,1962,1980]" captionTargetBox="[132,1455,215,1920]" captionTargetId="figure-138@24.[698,1267,782,1352]" captionTargetPageId="24" captionText="FIG. 16. — Capitamon mizoramense n. gen., n. sp., holotype ♂ (46.3 × 35.7 mm) (ZSIK C.6122/2A) (A-G), paratype ♀ (51.3 × 37.6 mm) (ZSIK C.6122/2B) (H, I): A, mouthparts; B, right third maxilliped; C, chela (outer view); D, s1-s8 showing right G1 in situ; E, pleon and telson; F, left G1 (dorsal view); G, left G2; H, pleonal somites 3-6 and telson; I, s1-s8 showing vulvae. Scale bars: A, B, D, E, H, I, 5 mm; C, 10 mm; F, G, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13837142" httpUri="https://zenodo.org/record/13837142/files/figure.png" pageId="22" pageNumber="609">Fig. 16I</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>

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@ -0,0 +1,85 @@
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<mods:title id="D35207803A593F9E0FBB4C2349CDE03D">Identity of the freshwater crab Indochinamon beieri (Pretzmann, 1966), with the description of a new genus and four new species from northeastern India (Decapoda, Brachyura, Potamidae)</mods:title>
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<heading id="D0AD81F2FFD3FFA3FF3FFE294DD62BE7" box="[245,662,438,465]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="592" reason="2">
Genus
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<emphasis id="B92EEA8CFFD3FFA3FE88FE284E992BE6" box="[322,473,438,464]" italics="true" pageId="5" pageNumber="592">Indochinamon</emphasis>
<bibRefCitation id="EFCB4B6FFFD3FFA3FE2AFE294DD62BE7" author="YEO D. C. J. &amp; NG P. K. L." box="[480,662,439,465]" pageId="5" pageNumber="592" pagination="273 - 308" refId="ref18959" refString="YEO D. C. J. &amp; NG P. K. L. 2007. - On the genus &quot; Potamon &quot; and allies in Indochina (Crustacea: Decapoda: Brachyura: Potamidae). Raffles Bulletin of Zoology 16 (Suppl.): 273 - 308." type="journal article" year="2007">Yeo &amp; Ng, 2007</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3406515FFD3FFA3FF4EFE6F4ED02815" pageId="5" pageNumber="592" type="materials_examined">
<paragraph id="8BE5369EFFD3FFA3FF4EFE6F4ED02815" blockId="5.[132,775,497,547]" pageId="5" pageNumber="592">
<smallCapsWord id="8D03A042FFD3FFA3FF4EFE6F4FF62831" baselines="514,515" box="[132,182,497,520]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="5" pageNumber="592">
<typeStatus id="54E1883CFFD3FFA3FF4EFE6F4FF62831" box="[132,182,497,520]" pageId="5" pageNumber="592">TYPE</typeStatus>
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<smallCapsWord id="8D03A042FFD3FFA3FF74FE684E452831" baselines="515" box="[190,261,502,519]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="species" pageId="5" pageNumber="592">SPECIES</smallCapsWord>
. —
<taxonomicName id="4C5A4D1DFFD3FFA3FEFCFE6C4DD0283F" authority="Yeo &amp; Ng, 1998" authorityName="Yeo &amp; Ng" authorityYear="1998" box="[310,656,497,521]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="592" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="B92EEA8CFFD3FFA3FEFCFE6C4EA0283F" box="[310,480,497,521]" italics="true" pageId="5" pageNumber="592">Potamon villosum</emphasis>
<bibRefCitation id="EFCB4B6FFFD3FFA3FE2CFE6F4DD0283F" author="YEO D. C. J. &amp; NG P. K. L." box="[486,656,497,521]" pageId="5" pageNumber="592" pagination="627 - 650" refId="ref18843" refString="YEO D. C. J. &amp; NG P. K. L. 1998. - Freshwater crabs of the Potamon tannanti species group (Crustacea: Decapoda: Brachyura: Potamidae) from northern Indochina. Raffles Bulletin of Zoology 46 (2): 627 - 650." type="journal article" year="1998">Yeo &amp; Ng, 1998</bibRefCitation>
</taxonomicName>
, by original designation; gender neuter.
</paragraph>
</subSubSection>
</treatment>
</document>